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<document id="8A2CD406E479B19B439DEA361CDCC892" ID-DOI="10.11646/zootaxa.5570.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734630" ID-ZooBank="8EF2BC5F-5939-440C-A053-A8924F07A42C" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1737736316380" checkinUser="plazi" docAuthor="Zheng, Xinyi, Li, Yuang, Tan, Zhixiang &amp; Wu, San’An" docDate="2025" docId="03CD8437232BFF83FF5C5405FA29FA08" docLanguage="en" docName="zootaxa.5570.1.5.pdf" docOrigin="Zootaxa 5570 (1)" docSource="https://doi.org/10.11646/zootaxa.5570.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Puto Signoret" docType="key" docVersion="4" lastPageNumber="134" masterDocId="FFF4FC4F2325FF8CFFCB537EFFB5FF8F" masterDocTitle="A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu" masterLastPageNumber="137" masterPageNumber="119" pageNumber="133" updateTime="1738059700552" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="7A8A3A36EA277457BE9E79960EBF0C44">A new species of the genus Puto Signoret (Hemiptera: Coccomorpha: Putoidae) from China and redescription of Puto huangshanensis Wu</mods:title>
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<mods:namePart id="16F2CCF975A103EA7A2CB7645448939A">Zheng, Xinyi</mods:namePart>
<mods:nameIdentifier id="A8DE8B54A494FF6C1981DE4597D7655C" type="ORCID">0000-0002-7396-7488</mods:nameIdentifier>
<mods:affiliation id="CA2820A98CF369ABA29275046DE05A56">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:namePart id="00C2144747458911DB05E189D1878C2B">Li, Yuang</mods:namePart>
<mods:nameIdentifier id="980F5EE7B6DBD8122508E722EC6A9522" type="ORCID">0009-0004-8851-6796</mods:nameIdentifier>
<mods:affiliation id="810E2795A6872808C5027B2344232DF8">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
<mods:name id="B00AECF2A91766D9E0E1A5C03F574854" type="personal">
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<mods:namePart id="A4F879F754B8C69D1374978FD8247FA6">Tan, Zhixiang</mods:namePart>
<mods:nameIdentifier id="863E54A9637D4DA72C3B0B2C33E8F41A" type="ORCID">0000-0002-9754-8341</mods:nameIdentifier>
<mods:affiliation id="83FE22B6A362EBDBD562389A0C07F9FB">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025, China</mods:affiliation>
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<mods:namePart id="A072B8B4D1D2EA84C156290DF527DE70">Wu, San’An</mods:namePart>
<mods:nameIdentifier id="ADE6813A9BC8A89BC4AB257ABF9F177F" type="ORCID">0000-0002-9671-9401</mods:nameIdentifier>
<mods:affiliation id="279597B913F514451AAB055BBD4A8B16">The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
<mods:nameIdentifier id="C1DE74D23FDDA4D0656B5162F3156CD3" type="email">sananwu@bjfu.edu.cn</mods:nameIdentifier>
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<mods:title id="5294DBEC5790D5BE9A82B3E939E5506D">Zootaxa</mods:title>
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<mods:date id="BC2E25C4C333DBABAB3559B517DE4285">2025</mods:date>
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<mods:number id="E0A82033701E889361058152FD5759B4">2025-01-16</mods:number>
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<mods:number id="2556E72B049C7DD9B10879026AA40E9A">5570</mods:number>
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<mods:number id="CE16EB7404E4E380D224ECF885AD9003">1</mods:number>
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<mods:url id="614D6B61B38337DB04042BC33EC67493">https://doi.org/10.11646/zootaxa.5570.1.5</mods:url>
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<mods:identifier id="B74A604A4ED5F08F4CC7D9AF41A7FB36" type="DOI">10.11646/zootaxa.5570.1.5</mods:identifier>
<mods:identifier id="66D94422C5DF75F5C287A0B2BFEE112E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CD8437232BFF83FF5C5405FA29FA08" LSID="urn:lsid:plazi:treatment:03CD8437232BFF83FF5C5405FA29FA08" httpUri="http://treatment.plazi.org/id/03CD8437232BFF83FF5C5405FA29FA08" lastPageId="15" lastPageNumber="134" pageId="14" pageNumber="133">
<subSubSection id="C37E66AA232BFF82FF5C5405FB43F81B" box="[151,1270,1914,1941]" pageId="14" pageNumber="133" type="nomenclature">
<paragraph id="8BDB3521232BFF82FF5C5405FB43F81B" blockId="14.[151,1270,1914,1941]" box="[151,1270,1914,1941]" pageId="14" pageNumber="133">
<heading id="D093824D232BFF82FF5C5405FB43F81B" bold="true" box="[151,1270,1914,1941]" fontSize="11" level="1" pageId="14" pageNumber="133" reason="1">
<emphasis id="B910E933232BFF82FF5C5405FB43F81B" bold="true" box="[151,1270,1914,1941]" pageId="14" pageNumber="133">
Key to adult females of
<taxonomicName id="4C644EA2232BFF82FE745405FE4DF81A" ID-CoL="75LQ" ID-ENA="249564" authority="Signoret" authorityName="Signoret" box="[447,504,1915,1941]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis id="B910E933232BFF82FE745405FE4DF81A" bold="true" box="[447,504,1915,1941]" italics="true" pageId="14" pageNumber="133">Puto</emphasis>
</taxonomicName>
from the Old World (updated from Danzig &amp; Gavrilov 2014)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E66AA232BFF83FF5C54BDFB96FF5A" lastPageId="15" lastPageNumber="134" pageId="14" pageNumber="133" type="discussion">
<paragraph id="8BDB3521232BFF83FF5C54BDFB96FF5A" blockId="14.[151,1437,1986,2013]" lastBlockId="15.[151,1437,151,213]" lastPageId="15" lastPageNumber="134" pageId="14" pageNumber="133">
<emphasis id="B910E933232BFF82FF5C54BDFF50F852" bold="true" box="[151,229,1987,2013]" pageId="14" pageNumber="133">Notes:</emphasis>
The key does not include
<taxonomicName id="4C644EA2232BFF82FDD854BDFCC2F852" authority="(Signoret 1875)" baseAuthorityName="Signoret" baseAuthorityYear="1875" box="[531,887,1986,2013]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="brunnitarsis">
<emphasis id="B910E933232BFF82FDD854BDFD0EF852" box="[531,699,1987,2013]" italics="true" pageId="14" pageNumber="133">P. brunnitarsis</emphasis>
(
<bibRefCitation id="EFF548D0232BFF82FD0154BDFCDAF852" author="Signoret, V." box="[714,879,1986,2013]" pageId="14" pageNumber="133" pagination="305 - 352" refId="ref19865" refString="Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 15 e partie. Annales de la Societe entomologique de France (serie 5), 5, 305 - 352." type="journal article" year="1875">Signoret 1875</bibRefCitation>
)
</taxonomicName>
because the available description is inadequate. The data of
<taxonomicName id="4C644EA2232AFF83FEEE53E9FD92FF3E" authority="(Cockerell)" baseAuthorityName="Cockerell" baseAuthorityYear="1895" box="[293,551,151,177]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="barberi">
<emphasis id="B910E933232AFF83FEEE53E9FE2CFF3E" box="[293,409,151,177]" italics="true" pageId="15" pageNumber="134">P. barberi</emphasis>
(Cockerell)
</taxonomicName>
,
<taxonomicName id="4C644EA2232AFF83FDFF53E9FD60FF3F" authorityName="Marotta &amp; Tranfaglia" authorityYear="1993" box="[564,725,151,176]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="marsicanus">
<emphasis id="B910E933232AFF83FDFF53E9FD60FF3F" box="[564,725,151,176]" italics="true" pageId="15" pageNumber="134">P. marsicanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C644EA2232AFF83FCDB53E9FC09FF3E" authorityName="Williams" authorityYear="2004" box="[784,956,151,177]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis id="B910E933232AFF83FCDB53E9FC09FF3E" box="[784,956,151,177]" italics="true" pageId="15" pageNumber="134">P. thailandicus</emphasis>
</taxonomicName>
are derived from
<bibRefCitation id="EFF548D0232AFF83FB4553E9FEF1FF5A" author="Williams, D. J. &amp; Granara de Willink, M. C." pageId="15" pageNumber="134" refId="ref20099" refString="Williams, D. J. &amp; Granara de Willink, M. C. (1992) Mealybugs of Central and South America. CAB International, Wallingford, 635 pp." type="book" year="1992">Williams &amp; Granara de Willink (1992)</bibRefCitation>
,
<bibRefCitation id="EFF548D0232AFF83FE9B53C5FD2DFF5A" author="Marotta, S. &amp; Tranfaglia, A." box="[336,664,187,213]" pageId="15" pageNumber="134" pagination="171 - 205" refId="ref19748" refString="Marotta, S. &amp; Tranfaglia, A. (1993) Le specie del genere Puto Signoret (Homoptera Coccoidea Pseudococcidae) dell'Europa centrale e del bacino del Mediterraneo. Bollettino del Laboratorio di Entomologia Agraria ' Filippo Silvestri', 48, 171 - 205." type="journal article" year="1993">Marotta &amp; Tranfaglia (1993)</bibRefCitation>
and
<bibRefCitation id="EFF548D0232AFF83FD1B53C5FC3EFF5A" author="Williams, D. J." box="[720,907,187,213]" pageId="15" pageNumber="134" refId="ref20067" refString="Williams, D. J. (2004) Mealybugs of Southern Asia. The Natural History Museum, London. Southdene SDN. BHD, Kuala Lumpur, 896 pp." type="book" year="2004">Williams (2004)</bibRefCitation>
respectively.
</paragraph>
</subSubSection>
<subSubSection id="C37E66AA232AFF83FF5C5387FA29FA08" pageId="15" pageNumber="134" type="key">
<keyStep id="30902D84232AFF83FF5C5387FA29FEA3" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5387FA29FE81" blockId="15.[151,1437,249,1415]" box="[151,1436,249,271]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5387FA29FE81" box="[151,1436,249,271]" pageId="15" pageNumber="134">1(0) Claw with denticle well developed........................................................................ 2</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5269FA29FEA3" blockId="15.[151,1437,249,1415]" box="[151,1436,278,300]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5269FA29FEA3" box="[151,1436,278,300]" pageId="15" pageNumber="134">
- Claw without denticle...............................................................
<taxonomicName id="4C644EA2232AFF83FB615269FA29FEA3" ID-CoL="4QL8M" authority="(Borchsenius)" baseAuthorityName="Borchsenius" baseAuthorityYear="1948" box="[1194,1436,278,300]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B910E933232AFF83FB615269FAA5FEA4" box="[1194,1296,278,300]" italics="true" pageId="15" pageNumber="134">P. borealis</emphasis>
(Borchsenius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C524AFA29FEE9" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C524AFA29FEC6" blockId="15.[151,1437,249,1415]" box="[151,1436,308,330]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C524AFA29FEC6" box="[151,1436,308,330]" pageId="15" pageNumber="134">2(1) Dorsal tubular ducts present............................................................................. 9</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C522FFA29FEE9" blockId="15.[151,1437,249,1415]" box="[151,1436,337,359]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C522FFA29FEE9" box="[151,1436,337,359]" pageId="15" pageNumber="134">- Dorsal tubular ducts absent............................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5210FA29FE2D" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5210FA29FE0C" blockId="15.[151,1437,249,1415]" box="[151,1436,366,388]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5210FA29FE0C" box="[151,1436,366,388]" pageId="15" pageNumber="134">3(2) Ventral tubular ducts of 2 or 3 sizes....................................................................... 4</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C52F2FA29FE2D" blockId="15.[151,1437,249,1415]" box="[151,1436,396,418]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C52F2FA29FE2D" box="[151,1436,396,418]" pageId="15" pageNumber="134">- Ventral tubular ducts all same size........................................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C52D7FA29FE53" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C52D7FA29FE30" blockId="15.[151,1437,249,1415]" box="[151,1436,425,447]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C52D7FA29FE30" box="[151,1436,425,447]" pageId="15" pageNumber="134">4(3) Ventral tubular ducts of 2 sizes........................................................................... 5</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C52B9FA29FE53" blockId="15.[151,1437,249,1415]" box="[151,1436,454,476]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C52B9FA29FE53" box="[151,1436,454,476]" pageId="15" pageNumber="134">
- Ventral tubular ducts of 3 sizes............................................................
<taxonomicName id="4C644EA2232AFF83FB1652B9FA29FE53" ID-CoL="78P6W" authority="Danzig" authorityName="Danzig" authorityYear="1978" box="[1245,1436,454,476]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B910E933232AFF83FB1652B9FAE4FE54" box="[1245,1361,454,476]" italics="true" pageId="15" pageNumber="134">P. orientalis</emphasis>
Danzig
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C529DFA29FDDD" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C529DFA28FD98" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C529DFA28FD98" pageId="15" pageNumber="134">
5(4) Large ducts present along margins of thorax and posterior abdominal sternites, absent from median areas of posterior abdominal sternites...........................................................................
<taxonomicName id="4C644EA2232AFF83FB7F517CFA28FD98" ID-CoL="78P76" authority="(Signoret)" baseAuthorityName="Signoret" baseAuthorityYear="1875" box="[1204,1437,513,535]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="B910E933232AFF83FB7F517CFA80FD98" box="[1204,1333,514,535]" italics="true" pageId="15" pageNumber="134">P. antennatus</emphasis>
(Signoret)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5161FA29FDDD" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5161FA29FDDD" pageId="15" pageNumber="134">
- Large ducts absent from margins of thorax and posterior abdominal sternites, but present on median areas of posterior abdominal sternites.............................................................................
<taxonomicName id="4C644EA2232AFF83FB025142FA29FDDD" ID-CoL="4QL8L" authority="(Cockerell)" baseAuthorityName="Cockerell" baseAuthorityYear="1895" box="[1225,1436,572,594]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="barberi">
<emphasis id="B910E933232AFF83FB025142FA9CFDDE" box="[1225,1321,572,593]" italics="true" pageId="15" pageNumber="134">P. barberi</emphasis>
(Cockerell)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5127FA29FD03" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5127FA29FDE0" blockId="15.[151,1437,249,1415]" box="[151,1436,601,623]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5127FA29FDE0" box="[151,1436,601,623]" pageId="15" pageNumber="134">
6(3) Ventral tubular ducts scattered throughout posterior part of body, but absent from anterior part of body...
<taxonomicName id="4C644EA2232AFF83FB235124FA29FDE0" ID-CoL="4QL9M" authority="Takahashi" authorityName="Takahashi" authorityYear="1941" box="[1256,1436,601,623]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="konoi">
<emphasis id="B910E933232AFF83FB235124FA80FDE1" box="[1256,1333,601,623]" italics="true" pageId="15" pageNumber="134">P. konoi</emphasis>
Takahashi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5109FA29FD03" blockId="15.[151,1437,249,1415]" box="[151,1436,630,652]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5109FA29FD03" box="[151,1436,630,652]" pageId="15" pageNumber="134">- Ventral tubular ducts present in frontal group only, or (rarely) totally absent....................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C51EAFA29FD49" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C51EAFA29FD25" blockId="15.[151,1437,249,1415]" box="[151,1436,660,682]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C51EAFA29FD25" box="[151,1436,660,682]" pageId="15" pageNumber="134">
7(6) Multilocular disc-pores numerous, scattered on all medial zone of venter..................
<taxonomicName id="4C644EA2232AFF83FBB651EAFAEEFD25" ID-CoL="6WP5J" authority="(Leonardi)" authorityName="" baseAuthorityName="Leonardi" baseAuthorityYear="1907" box="[1149,1371,660,682]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="B910E933232AFF83FBB651EAFB59FD26" box="[1149,1260,660,681]" italics="true" pageId="15" pageNumber="134">P. superbus</emphasis>
(Leonardi)
</taxonomicName>
, partly
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C51CFFA29FD49" blockId="15.[151,1437,249,1415]" box="[151,1436,689,711]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C51CFFA29FD49" box="[151,1436,689,711]" pageId="15" pageNumber="134">- Multilocular disc-pores few, present on cephalothoracic sternites only or totally absent.............................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C51B0FA29FC8E" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C51B0FA29FD6B" blockId="15.[151,1437,249,1415]" box="[151,1436,718,740]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C51B0FA29FD6B" box="[151,1436,718,740]" pageId="15" pageNumber="134">
8(7) Multilocular disc-pores present on cephalothoracic sternites...............................
<taxonomicName id="4C644EA2232AFF83FB5251B1FA29FD6B" ID-CoL="78P7C" authority="(Vayssiere)" baseAuthorityName="Vayssiere" baseAuthorityYear="1923" box="[1177,1436,718,740]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="peyerimhoffi">
<emphasis id="B910E933232AFF83FB5251B1FA9FFD6C" box="[1177,1322,718,740]" italics="true" pageId="15" pageNumber="134">P. peyerimhoffi</emphasis>
(Vayssiere)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5192FA29FC8E" blockId="15.[151,1437,249,1415]" box="[151,1436,748,770]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5192FA29FC8E" box="[151,1436,748,770]" pageId="15" pageNumber="134">
- Multilocular disc-pores absent..........................................................
<taxonomicName id="4C644EA2232AFF83FB0B5192FA29FC8E" ID-CoL="78P79" authority="Ben-Dov" authorityName="Ben-Dov" authorityYear="2001" box="[1216,1436,748,769]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis id="B910E933232AFF83FB0B5192FA8AFC8E" box="[1216,1343,748,769]" italics="true" pageId="15" pageNumber="134">P. israelensis</emphasis>
Ben-Dov
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5077FA29FCB4" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5077FA29FC90" blockId="15.[151,1437,249,1415]" box="[151,1436,777,799]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5077FA29FC90" box="[151,1436,777,799]" pageId="15" pageNumber="134">
9(2) Dorsal tubular ducts of 2 sizes; smaller ducts present along margins of abdominal tergites..........
<taxonomicName id="4C644EA2232AFF83FB7A5074FA29FC90" ID-CoL="4QL8Z" authority="Hadzibejli" authorityName="Hadzibejli" authorityYear="1956" box="[1201,1436,777,799]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="caucasicus">
<emphasis id="B910E933232AFF83FB7A5074FA87FC90" box="[1201,1330,778,799]" italics="true" pageId="15" pageNumber="134">P. caucasicus</emphasis>
Hadzibejli
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5059FA29FCB4" blockId="15.[151,1437,249,1415]" box="[151,1436,806,828]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5059FA29FCB4" box="[151,1436,806,828]" pageId="15" pageNumber="134">- Dorsal tubular ducts all same size, large.................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C503AFA29FCF9" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C503AFA29FCD6" blockId="15.[151,1437,249,1415]" box="[151,1436,836,858]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C503AFA29FCD6" box="[151,1436,836,858]" pageId="15" pageNumber="134">10(9) Dorsal tubular ducts wide; diameter of their openings more than 2 times larger than diameter of a trilocular pore......... 11</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C501FFA29FCF9" blockId="15.[151,1437,249,1415]" box="[151,1436,865,887]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C501FFA29FCF9" box="[151,1436,865,887]" pageId="15" pageNumber="134">- Dorsal tubular ducts narrow; diameter of their openings about the same size as diameter of a trilocular pore............. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5000FA29FC41" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5000FA29FC3D" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5000FA29FC3D" pageId="15" pageNumber="134">
11(10) Dorsal tubular ducts located only in cerarii. Ventral tubular ducts all same size, present only in frontal group................................................................................................
<emphasis id="B910E933232AFF83FB0E50E2FA9FFC3E" box="[1221,1322,924,945]" italics="true" pageId="15" pageNumber="134">subericola</emphasis>
(Vayssiere)
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C50C7FA29FC41" blockId="15.[151,1437,249,1415]" box="[151,1436,953,975]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C50C7FA29FC41" box="[151,1436,953,975]" pageId="15" pageNumber="134">- Dorsal tubular ducts located between cerarii. Ventral tubular ducts of 2 or 3 sizes, numerous on all abdominal sternites.... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C50A8FA29FB85" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C50A8FA28FC63" blockId="15.[151,1437,249,1415]" box="[151,1437,982,1004]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C50A8FA28FC63" box="[151,1437,982,1004]" pageId="15" pageNumber="134">
12(11) Quinquelocular disc-pores present.......................................................
<taxonomicName id="4C644EA2232AFF83FB0F50A9FA28FC63" authority="Wu" authorityName="Wu" box="[1220,1437,982,1004]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="huangshanensis">
<emphasis id="B910E933232AFF83FB0F50A9FAC1FC64" box="[1220,1396,982,1004]" italics="true" pageId="15" pageNumber="134">P. huangshanensis</emphasis>
Wu
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C508AFA29FB85" blockId="15.[151,1437,249,1415]" box="[151,1436,1012,1034]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C508AFA29FB85" box="[151,1436,1012,1034]" pageId="15" pageNumber="134">
- Quinquelocular disc-pores absent...........................................................
<taxonomicName id="4C644EA2232AFF83FB2C508AFA29FB85" ID-CoL="6WP5N" authority="Danzig" authorityName="Danzig" authorityYear="1978" box="[1255,1436,1012,1034]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="tubulifer">
<emphasis id="B910E933232AFF83FB2C508AFAE4FB86" box="[1255,1361,1012,1033]" italics="true" pageId="15" pageNumber="134">P. tubulifer</emphasis>
Danzig
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C576FFA29FBCB" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C576FFA29FBA9" blockId="15.[151,1437,249,1415]" box="[151,1436,1041,1063]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C576FFA29FBA9" box="[151,1436,1041,1063]" pageId="15" pageNumber="134">13(10) Dorsal tubular ducts present along body margin only........................................................ 14</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5751FA29FBCB" blockId="15.[151,1437,249,1415]" box="[151,1436,1070,1092]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5751FA29FBCB" box="[151,1436,1070,1092]" pageId="15" pageNumber="134">- Dorsal tubular ducts present in medial zones of tergites as well as along margins.................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5732FA29FB36" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5732FA29FBF0" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5732FA29FBF0" pageId="15" pageNumber="134">
14(13) Dorsum with additional cerarii, forming 6 longitudinal rows. Ventral margins without small oral collar tubular ducts....................................................................................
<taxonomicName id="4C644EA2232AFF83FBB65714FAEEFBF0" ID-CoL="6WP5J" authority="(Leonardi)" authorityName="" baseAuthorityName="Leonardi" baseAuthorityYear="1907" box="[1149,1371,1129,1151]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="B910E933232AFF83FBB65714FB59FBF1" box="[1149,1260,1129,1151]" italics="true" pageId="15" pageNumber="134">P. superbus</emphasis>
(Leonardi)
</taxonomicName>
, partly
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C57F9FA29FB36" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C57F9FA29FB36" pageId="15" pageNumber="134">
- Dorsum without additional cerarii. Ventral margins with groups of small oral collar tubular ducts..................................................................................................
<taxonomicName id="4C644EA2232AFF83FBB857DAFAE9FB36" authority="Zheng &amp; Wu" authorityName="Zheng &amp; Wu" authorityYear="2025" box="[1139,1372,1188,1210]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="sinensis" status="sp. n.">
<emphasis id="B910E933232AFF83FBB857DAFB62FB36" box="[1139,1239,1188,1209]" italics="true" pageId="15" pageNumber="134">P. sinensis</emphasis>
Zheng &amp; Wu
</taxonomicName>
,
<emphasis id="B910E933232AFF83FAAD57DAFA29FB36" bold="true" box="[1382,1436,1188,1209]" pageId="15" pageNumber="134">
<taxonomicNameLabel id="A2235448232AFF83FAAD57DAFA29FB36" box="[1382,1436,1188,1209]" pageId="15" pageNumber="134" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C57BFFA29FB7C" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C57BFFA29FB58" blockId="15.[151,1437,249,1415]" box="[151,1436,1217,1239]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C57BFFA29FB58" box="[151,1436,1217,1239]" pageId="15" pageNumber="134">15(13) Ventral tubular ducts present on thoracic and abdominal sternites as well as in frontal group......................... 16</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C57A1FA29FB7C" blockId="15.[151,1437,249,1415]" box="[151,1436,1246,1268]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C57A1FA29FB7C" box="[151,1436,1246,1268]" pageId="15" pageNumber="134">- Ventral tubular ducts absent from thoracic and abdominal sternites, present only in frontal group..................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C5785FA29FAC3" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C5785FA29FAA0" blockId="15.[151,1437,249,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5785FA29FAA0" pageId="15" pageNumber="134">
16(15) Dorsum with additional cerarii forming 2 longitudinal rows, plus several additional cerarii on abdominal segment VI.....................................................................................
<taxonomicName id="4C644EA2232AFF83FB5B5664FA29FAA0" ID-CoL="4QL9V" authority="(Borchsenius)" baseAuthorityName="Borchsenius" baseAuthorityYear="1948" box="[1168,1436,1305,1327]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="megriensis">
<emphasis id="B910E933232AFF83FB5B5664FAA5FAA0" box="[1168,1296,1306,1327]" italics="true" pageId="15" pageNumber="134">P. megriensis</emphasis>
(Borchsenius)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C5649FA29FAC3" blockId="15.[151,1437,249,1415]" box="[151,1436,1334,1356]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C5649FA29FAC3" box="[151,1436,1334,1356]" pageId="15" pageNumber="134">
- Dorsum without additional cerarii......................................................
<taxonomicName id="4C644EA2232AFF83FB7A5649FA29FAC3" authority="Williams" authorityName="Williams" authorityYear="2004" box="[1201,1436,1334,1356]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="thailandicus">
<emphasis id="B910E933232AFF83FB7A5649FAF5FAC4" box="[1201,1344,1334,1356]" italics="true" pageId="15" pageNumber="134">P. thailandicus</emphasis>
Williams
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30902D84232AFF83FF5C562DFA29FA08" pageId="15" pageNumber="134">
<paragraph id="8BDB3521232AFF83FF5C562DFA29FAE5" blockId="15.[151,1437,249,1415]" box="[151,1436,1363,1386]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C562DFA29FAE5" box="[151,1436,1363,1386]" pageId="15" pageNumber="134">
17(15) Dorsum with additional cerarii forming 6 longitudinal rows........................
<taxonomicName id="4C644EA2232AFF83FB88562AFA29FAE5" ID-CoL="6WP6W" authority="Marotta &amp; Tranfaglia" authorityName="Marotta &amp; Tranfaglia" authorityYear="1993" box="[1091,1436,1364,1386]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="marsicanus">
<emphasis id="B910E933232AFF83FB88562AFB7CFAE6" box="[1091,1225,1364,1385]" italics="true" pageId="15" pageNumber="134">P. marsicanus</emphasis>
Marotta &amp; Tranfaglia
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BDB3521232AFF83FF5C560FFA29FA08" blockId="15.[151,1437,249,1415]" box="[151,1436,1393,1415]" pageId="15" pageNumber="134">
<keyLead id="30959614232AFF83FF5C560FFA29FA08" box="[151,1436,1393,1415]" pageId="15" pageNumber="134">
- Dorsum with additional cerarii forming 2 longitudinal rows..............................
<taxonomicName id="4C644EA2232AFF83FB4E560CFA29FA08" ID-CoL="4QL9K" authority="(Borchsenius)" baseAuthorityName="Borchsenius" baseAuthorityYear="1948" box="[1157,1436,1393,1415]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="kondarensis">
<emphasis id="B910E933232AFF83FB4E560CFABAFA09" box="[1157,1295,1393,1415]" italics="true" pageId="15" pageNumber="134">P. kondarensis</emphasis>
(Borchsenius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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