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<mods:title id="036EAD2B47A50C70826E2037E14EBF62">Taxonomic review of the Oriental flower beetle Coilodera penicillata species complex (Coleoptera: Scarabaeidae: Cetoniinae)</mods:title>
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<treatment id="03CD6B0AFFDEFC472E81FDF0F716FF36" ID-DOI="http://doi.org/10.5281/zenodo.6040979" ID-GBIF-Taxon="137453743" ID-Zenodo-Dep="6040979" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36" httpUri="http://treatment.plazi.org/id/03CD6B0AFFDEFC472E81FDF0F716FF36" lastPageId="13" lastPageNumber="523" pageId="10" pageNumber="521">
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<taxonomicName id="4C64A19FFFDEFC402E81FDF0F018FE15" authority="Hope, 1831" authorityName="Hope" authorityYear="1831" box="[151,551,476,502]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="521" phylum="Arthropoda" rank="species" species="penicillata">
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<emphasis id="B910060EFFDEFC402E81FDF0F3ABFE15" bold="true" box="[151,404,476,502]" italics="true" pageId="10" pageNumber="521">Coilodera penicillata</emphasis>
<bibRefCitation id="EFF5A7EDFFDEFC402F8DFDF0F018FE15" author="Hope" box="[411,551,476,502]" pageId="10" pageNumber="521" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope, 1831</bibRefCitation>
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(
<figureCitation id="135FC699FFDEFC402E89FDD2F319FDF4" box="[159,294,510,535]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="10" pageNumber="521">Figs. 21–24</figureCitation>
, 
<figureCitation id="135FC699FFDEFC402F24FDD2F3C6FDF4" box="[306,505,510,535]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">52–56, 63–64, 70</figureCitation>
, 
<figureCitation id="135FC699FFDEFC402C12FDD3F049FDF4" box="[516,630,510,535]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="10" pageNumber="521">79–80, 85</figureCitation>
, 
<figureCitation id="135FC699FFDEFC402C94FDD3F0E7FDF4" box="[642,728,511,535]" captionStart-0="FIGURES 95 – 99" captionStart-1="FIGURES 100 – 106" captionStartId-0="19.[151,264,1958,1981]" captionStartId-1="20.[151,264,1875,1898]" captionTargetBox-0="[175,1411,193,1938]" captionTargetBox-1="[187,1399,193,1849]" captionTargetId-0="figure@19.[175,1411,193,1938]" captionTargetId-1="figure@20.[187,1399,193,1849]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 95 – 99. Habitat of Coilodera. 95 – 97, C. mearesii: 95, habitat (Zhangmu, Nyalam, Xizang); 96 – 97, host plant (Hydrangea sp.). 98 – 99, C. penicillata: larva in cockroach galleries of Salganea raggei in rotten wood (Yingjiang, Yunnan)." captionText-1="FIGURES 100 – 106. Habitat of Coilodera. 100 – 102, C. penicillata: 100 – 101, closer view of habitat and newly emerged adult in pupal cell in rotten wood (Tongbiguan Nature Reserve, Yingjiang, Yunnan); 102, newly emerged adult and larva inhabit in rotten wood with Blattodea and Passalidae (Mount Xiaolushan, Lancang, Yunnan). 103 – 106, Host plants of C. nigroscutellaris: 103 – 104. Hydrangea sp. (Mount Dayaoshan, Guangxi, photograph by Lu Qiu); 105 – 106. Ampelopsis sp. (Mount Wuzhishan, Hainan Island)." httpUri-0="https://zenodo.org/record/1059361/files/figure.png" httpUri-1="https://zenodo.org/record/1059363/files/figure.png" pageId="10" pageNumber="521">98–102</figureCitation>
)
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<taxonomicName id="4C64A19FFFDEFC402E81FE6AF025FDBF" authority="Hope, 1831: 25" authorityName="Hope" authorityPageNumber="25" authorityYear="1831" box="[151,538,582,604]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="521" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDEFC402E81FE6AF34EFDBF" box="[151,369,582,604]" italics="true" pageId="10" pageNumber="521">Coilodera penicillata</emphasis>
<treatmentCitation id="0AC5FC0DFFDEFC402F6CFE6BF025FDBF" author="Hope" box="[378,538,582,604]" page="25" pageId="10" pageNumber="521" year="1831">
<bibRefCitation id="EFF5A7EDFFDEFC402F6CFE6BF3CAFDBF" author="Hope" box="[378,501,582,604]" pageId="10" pageNumber="521" refString="Hope, F. W. (1831) Synopsis of the new species of Nepal insects in the collection of Major General Hardwicke. Zoological Miscellany, 1, 21 - 32." type="journal article" year="1831">Hope, 1831</bibRefCitation>
: 25
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(type locality: Nepal); 
<treatmentCitation id="0AC5FC0DFFDEFC402D1BFE6AF184FDBF" author="Paulian" box="[781,955,582,604]" page="13" pageId="10" pageNumber="521" year="1960">
<bibRefCitation id="EFF5A7EDFFDEFC402D1BFE6AF1A8FDBF" author="Paulian" box="[781,919,582,604]" pageId="10" pageNumber="521" refString="Paulian, R. (1960) Coleopteres Scarabeides de L'Indochine (Rutelines et Cetonines) (Suite). Annales de la Societe entomologique de France, 129, 1 - 87." type="journal article" year="1960">Paulian 1960</bibRefCitation>
: 13
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(Chapa, northern Vietnam; Xieng Khouang &amp; Lakhon, Laos), fig. 290; 
<treatmentCitation id="0AC5FC0DFFDEFC402FD5FE49F044FD99" author="Nakane" box="[451,635,613,635]" page="112" pageId="10" pageNumber="521" year="1972">
<bibRefCitation id="EFF5A7EDFFDEFC402FD5FE49F073FD99" author="Nakane" box="[451,588,613,635]" pageId="10" pageNumber="521" refString="Nakane, T. (1972) Coleoptera of Nepal collected by the Hokkaido University scientific expedition to Nepal Himalaya: Scarabaeidae I. Cetoniinae, Valginae and Dynastinae. Kontyu, 40, 112 - 118." type="journal article" year="1972">Nakane 1972</bibRefCitation>
: 112
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(Nepal), fig. 1 (parameres), 6 (habitus); 
<treatmentCitation id="0AC5FC0DFFDEFC402A03FE49F6F7FD99" author="Antoine" box="[1045,1224,613,635]" page="18" pageId="10" pageNumber="521" year="1986">
<bibRefCitation id="EFF5A7EDFFDEFC402A03FE49F69CFD98" author="Antoine" box="[1045,1187,613,635]" pageId="10" pageNumber="521" refString="Antoine, P. (1986) Sur la tribu Taenioderini: remarques taxonomiques et description d'especes nouvelles (Coleoptera, Cetoniidae). Bulletin de la Societe Sciences Nat, 49, 9 - 20." type="journal article" year="1986">Antoine 1986</bibRefCitation>
: 18
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, fig. 10b, 10d ♂; 
<treatmentCitation id="0AC5FC0DFFDEFC402B6CFE4AF313FD79" author="Ma" page="573" pageId="10" pageNumber="521" year="1987">
<bibRefCitation id="EFF5A7EDFFDEFC402B6CFE4AF2C2FD79" author="Ma" pageId="10" pageNumber="521" refString="Ma, W. Z. (1987) Coleoptera: Cetoniidae. In: Huang, F. S. (ed.), Forest Insects of Yunnan. Yunnan Science &amp; Technology Press, Kunming, China, pp. 570 - 580. [In Chinese]" type="book chapter" year="1987">Ma 1987</bibRefCitation>
: 573
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(Lincang, Yunnan, China), plate 2, fig. 26; 
<treatmentCitation id="0AC5FC0DFFDEFC402CF5FEA8F1ECFD79" author="Sakai" box="[739,979,644,666]" page="35" pageId="10" pageNumber="521" year="1996">
<bibRefCitation id="EFF5A7EDFFDEFC402CF5FEA8F190FD79" author="Sakai" box="[739,943,644,666]" pageId="10" pageNumber="521" refString="Sakai, K. &amp; Ikeda, K. (1996) Check list of insects of Thailand 4. Family Scarabaeidae, subfamily Cetoniinae. Memoirs of the Faculty of Liberal Arts &amp; Education. Part II, Mathematics &amp; Natural Sciences, University of Yamanashi, 47, 34 - 43." type="journal article" year="1996">Sakai &amp; Ikeda 1996</bibRefCitation>
: 35
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(northern Thailand); 
<treatmentCitation id="0AC5FC0DFFDEFC402ABAFEA8F766FD79" author="Krajcik" box="[1196,1369,644,666]" page="88" pageId="10" pageNumber="521" year="1998">
<bibRefCitation id="EFF5A7EDFFDEFC402ABAFEA8F70CFD79" author="Krajcik" box="[1196,1331,644,666]" pageId="10" pageNumber="521" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, Czech Republic, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
: 88
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402B75FEA8F3AFFD5B" author="Sakai" page="357" pageId="10" pageNumber="521" year="1998">
<bibRefCitation id="EFF5A7EDFFDEFC402B75FEA8F361FD5B" author="Sakai" pageId="10" pageNumber="521" refString="Sakai, K. &amp; Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Volume 3. Mushi- Sha, Tokyo, Japan, 421 pp." type="book" year="1998">Sakai &amp; Nagai 1998</bibRefCitation>
: 357
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(Nepal), plate 136, fig. 1625-1 ♀; Smetana 2006: 306; 
<treatmentCitation id="0AC5FC0DFFDEFC402DCEFE8EF6FBFD5B" author="Ek-Amnuay" box="[984,1220,674,696]" page="246" pageId="10" pageNumber="521" year="2008">
<bibRefCitation id="EFF5A7EDFFDEFC402DCEFE8EF6ADFD5B" author="Ek-Amnuay" box="[984,1170,674,696]" pageId="10" pageNumber="521" refString="Ek-Amnuay, P. (2008) Beetles of Thailand (2 nd edition). Fascinating Insects. Folume 1. Amarin Printing and Publishing Public, Bangkok, Thailand, 495 pp." type="book" year="2008">Ek-Amnuay 2008</bibRefCitation>
: 246
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(northern Thailand), plate 95, figs. C69 ♂ &amp; ♀; 
<treatmentCitation id="0AC5FC0DFFDEFC402FF3FEEDF040FD34" author="Sakai" box="[485,639,705,727]" page="20" pageId="10" pageNumber="521" year="2008">
<bibRefCitation id="EFF5A7EDFFDEFC402FF3FEEDF065FD34" author="Sakai" box="[485,602,705,727]" pageId="10" pageNumber="521" refString="Sakai, K. (2008) Record of cetoniid beetles from the central and northern Laos (Coleoptera, Scarabaeidae). Saikaku-Tsushin, 16, 11 - 22." type="journal article" year="2008">Sakai 2008</bibRefCitation>
: 20
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(Xieng Khuang &amp; Phu Pan, Laos), fig. 60 ♂; 
<treatmentCitation id="0AC5FC0DFFDEFC402A40FEEDF73BFD34" author="Krajcik" box="[1110,1284,705,727]" page="53" pageId="10" pageNumber="521" year="2011">
<bibRefCitation id="EFF5A7EDFFDEFC402A40FEEDF6E0FD34" author="Krajcik" box="[1110,1247,705,727]" pageId="10" pageNumber="521" refString="Krajcik, M. (2011) Illustrated catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x. Supplement, 1 - 113." type="book chapter" year="2011">Krajčík 2011</bibRefCitation>
: 53
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; 
<treatmentCitation id="0AC5FC0DFFDEFC402B19FEEDF2DCFD15" author="Krajcik" page="76" pageId="10" pageNumber="521" year="2012">
<bibRefCitation id="EFF5A7EDFFDEFC402B19FEEDF7A6FD34" author="Krajcik" box="[1295,1433,705,727]" pageId="10" pageNumber="521" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">Krajčík 2012</bibRefCitation>
: 76
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; 
<treatmentCitation id="0AC5FC0DFFDEFC402EFBFECCF3AAFD15" author="Li" box="[237,405,736,758]" page="32" pageId="10" pageNumber="521" year="2013">
<bibRefCitation id="EFF5A7EDFFDEFC402EFBFECCF34CFD15" author="Li" box="[237,371,736,758]" pageId="10" pageNumber="521" refString="Li, J. K., Zhang, X. P., Gao, M. X., Zhang, L. M., Huang, L. R. &amp; Flutsch, G. (2013) Cetoniidae from Mount Gaoligongshan, Yunnan, China (Coleoptera, Scarabaeoidea, Cetoniidae). Cetoniimania, new series, 5, 30 - 40." type="journal article" year="2013">
Li 
<emphasis id="B910060EFFDEFC402F1CFECDF308FD15" box="[266,311,736,758]" italics="true" pageId="10" pageNumber="521">et al</emphasis>
2013
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(Mount Gaoligongshan, Yunnan, China), fig. 37 ♂; 
<treatmentCitation id="0AC5FC0DFFDEFC402DBFFECCF65BFD15" author="Bezdek" box="[937,1124,736,758]" page="401" pageId="10" pageNumber="521" year="2016">
<bibRefCitation id="EFF5A7EDFFDEFC402DBFFECCF60DFD15" author="Bezdek" box="[937,1074,736,758]" pageId="10" pageNumber="521" refString="Bezdek, A. (2016) Subfamily Cetoniinae, pp. 367 - 412. In: Lobl, I. &amp; Lobl, D. (eds), Catalogue of Palaearctic Coleoptera, Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, The Netherlands." type="journal article" year="2016">Bezděk 2016</bibRefCitation>
: 401
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.
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<taxonomicName id="4C64A19FFFDEFC402E81FED2F387FCF7" authority="Hope" authorityName="Hope" authorityYear="1831" box="[151,440,766,788]" class="Insecta" family="Scarabaeidae" genus="Coelodera" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="521" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDEFC402E81FED2F34AFCF7" box="[151,373,766,788]" italics="true" pageId="10" pageNumber="521">Coelodera penicillata</emphasis>
Hope
</taxonomicName>
: 
<treatmentCitation id="0AC5FC0DFFDEFC402FD4FED2F09BFCF7" author="Burmeister" box="[450,676,766,788]" page="321" pageId="10" pageNumber="521" year="1842">
<bibRefCitation id="EFF5A7EDFFDEFC402FD4FED2F04EFCF7" author="Burmeister" box="[450,625,766,788]" pageId="10" pageNumber="521" refString="Burmeister, H. C. C. (1842) Handbuch der Entomologie, Band III. Besondere Entomologie, Fortsetzung. Coleoptera Lamellicornia Melitophila. T. C. F. Enslin, Berlin, Germany, 828 pp." type="book" year="1842">Burmeister 1842</bibRefCitation>
: 321
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402CA6FED2F14BFCF7" author="Schoch" box="[688,884,766,788]" page="378" pageId="10" pageNumber="521" year="1896">
<bibRefCitation id="EFF5A7EDFFDEFC402CA6FED2F17CFCF7" author="Schoch" box="[688,835,766,788]" pageId="10" pageNumber="521" refString="Schoch, G. (1896 a) Nachtrag II &amp; III zu Schoch: Die Genera und Species meiner Cetoniden-sammlung. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 9, 378 - 401." type="journal article" year="1896" yearSuffix="a">Schoch 1896a</bibRefCitation>
: 378
</treatmentCitation>
(Assam, India); 
<treatmentCitation id="0AC5FC0DFFDEFC402A37FED2F6E2FCF7" author="Schoch" box="[1057,1245,766,788]" page="34" pageId="10" pageNumber="521" year="1896">
<bibRefCitation id="EFF5A7EDFFDEFC402A37FED2F689FCF7" author="Schoch" box="[1057,1206,766,788]" pageId="10" pageNumber="521" refString="Schoch, G. (1896 b) Catalogus systematicus Cetonidarum et Trichiidarum ad huc cognitarum. W. Junk, Zurich, Switzerland, 95 pp." type="book" year="1896" yearSuffix="b">Schoch 1896b</bibRefCitation>
: 34
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402AFFFED2F2CFFCD0" author="Schenkling" page="133" pageId="10" pageNumber="521" year="1921">
<bibRefCitation id="EFF5A7EDFFDEFC402AFFFED2F7A7FCF7" author="Schenkling" box="[1257,1432,766,788]" pageId="10" pageNumber="521" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III. Pars 72. W. Junk, Berlin, Germany, 431 pp." type="book" year="1921">Schenkling 1921</bibRefCitation>
: 133
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402EEDFF31F391FCD0" author="Miksic" box="[251,430,797,819]" page="700" pageId="10" pageNumber="521" year="1972">
<bibRefCitation id="EFF5A7EDFFDEFC402EEDFF31F341FCD0" author="Miksic" box="[251,382,797,819]" pageId="10" pageNumber="521" refString="Miksic, R. (1972). Revision der Gattung Coelodera Burmeister (73. Beitrag zur Kenntnis der Scarabaeiden). Revue suisse de Zoologie, 79, 675 - 707." type="journal article" year="1972">Mikšić 1972</bibRefCitation>
: 700
</treatmentCitation>
(Nepal; northeastern India; northern Myanmar; northern Vietnam), figs. 18–19; 
<treatmentCitation id="0AC5FC0DFFDEFC402AF4FF31F7A7FCD0" author="Miksic" box="[1250,1432,797,819]" page="268" pageId="10" pageNumber="521" year="1976">
<bibRefCitation id="EFF5A7EDFFDEFC402AF4FF31F75AFCD0" author="Miksic" box="[1250,1381,797,819]" pageId="10" pageNumber="521" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, Bosnia and Herzegovina, 444 pp." type="book" year="1976">Mikšić 1976</bibRefCitation>
: 268
</treatmentCitation>
, fig. 32d; 
<treatmentCitation id="0AC5FC0DFFDEFC402F35FF11F399FCB1" author="Ma" box="[291,422,828,850]" page="41" pageId="10" pageNumber="521" year="1995">
<bibRefCitation id="EFF5A7EDFFDEFC402F35FF11F3BEFCB1" author="Ma" box="[291,385,828,850]" pageId="10" pageNumber="521" refString="Ma, W. Z. (1995) Economic Insect Fauna of China, Folume 46 Coleoptera: Cetoniidae, Trichiidae, and Falgidae. Science and Technology Press, Beijing, China, 210 pp. [In Chinese]" type="book" year="1995">Ma 1995</bibRefCitation>
: 41
</treatmentCitation>
, fig. 33.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDBDA1CFFDEFC402E81FF77F0B7FC6C" blockId="10.[151,1437,582,972]" pageId="10" pageNumber="521">
<treatmentCitationGroup id="AB74FD32FFDEFC402E81FF77F0B7FC6C" pageId="10" pageNumber="521">
<taxonomicName id="4C64A19FFFDEFC402E81FF77F3EAFC93" authority="(Hope)" authorityName="Hope" box="[151,469,858,880]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="521" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDEFC402E81FF77F341FC93" box="[151,382,858,880]" italics="true" pageId="10" pageNumber="521">Macronota penicillata</emphasis>
(Hope)
</taxonomicName>
: 
<treatmentCitation id="0AC5FC0DFFDEFC402FF2FF76F08EFC93" author="Harold" box="[484,689,858,880]" page="1298" pageId="10" pageNumber="521" year="1869">
<bibRefCitation id="EFF5A7EDFFDEFC402FF2FF76F052FC93" author="Harold" box="[484,621,858,880]" pageId="10" pageNumber="521" refString="Harold, E. von (1869) Scarabaeidae, Tome IV. In: Gemminger, M. &amp; Harold, E. von (eds.), Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. E. H. Grummi, Munich, Germany, pp. 979 - 1346." type="book chapter" year="1869">Harold 1869</bibRefCitation>
: 1298
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402CD6FF76F145FC93" author="Gestro" box="[704,890,858,880]" page="843" pageId="10" pageNumber="521" year="1891">
<bibRefCitation id="EFF5A7EDFFDEFC402CD6FF76F17AFC93" author="Gestro" box="[704,837,858,880]" pageId="10" pageNumber="521" refString="Gestro, R. (1891) Viaggio di Leonardo Fea in Birmania regione vicine. XXXVII. Enumerazione delle Cetonie. Annali del Museo civico di Storia Naturale di Genova, Series 2, 10, 835 - 876." type="journal article" year="1891">Gestro 1891</bibRefCitation>
: 843
</treatmentCitation>
(Bhamo, northeastern 
<collectingCountry id="F3739A8CFFDEFC402A64FF77F6E6FC93" box="[1138,1241,859,880]" name="Myanmar" pageId="10" pageNumber="521">Myanmar</collectingCountry>
); 
<treatmentCitation id="0AC5FC0DFFDEFC402AE6FF77F7A4FC93" author="Arrow" box="[1264,1435,858,880]" page="44" pageId="10" pageNumber="521" year="1910">
<bibRefCitation id="EFF5A7EDFFDEFC402AE6FF77F74CFC93" author="Arrow" box="[1264,1395,858,880]" pageId="10" pageNumber="521" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, United Kingdom, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
: 44
</treatmentCitation>
(northeastern 
<collectingCountry id="F3739A8CFFDEFC402F44FF55F3B6FC6C" box="[338,393,889,911]" name="India" pageId="10" pageNumber="521">India</collectingCountry>
; Keren Hills, 
<collectingCountry id="F3739A8CFFDEFC402C00FF56F042FC6C" box="[534,637,890,911]" name="Myanmar" pageId="10" pageNumber="521">Myanmar</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDBDA1CFFDEFC402E81FFB5F339FC2F" blockId="10.[151,1437,582,972]" pageId="10" pageNumber="521">
<treatmentCitationGroup id="AB74FD32FFDEFC402E81FFB5F339FC2F" pageId="10" pageNumber="521">
<taxonomicName id="4C64A19FFFDEFC402E81FFB5F37CFC4D" ID-CoL="3X74H" authorityName="Gory &amp; Percheron" authorityYear="1833" box="[151,323,920,942]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="521" phylum="Arthropoda" rank="species" species="dives">
<emphasis id="B910060EFFDEFC402E81FFB5F37CFC4D" box="[151,323,920,942]" italics="true" pageId="10" pageNumber="521">Macronota dives</emphasis>
</taxonomicName>
(nec Gory &amp; Percheron): 
<treatmentCitation id="0AC5FC0DFFDEFC402C4FFFB4F120FC4D" author="Schaum" box="[601,799,920,942]" page="367" pageId="10" pageNumber="521" year="1844">
<bibRefCitation id="EFF5A7EDFFDEFC402C4FFFB4F0D5FC4D" author="Schaum" box="[601,746,920,942]" pageId="10" pageNumber="521" refString="Schaum, H. (1844) Observations critiques sur la famille des Lamellicornes melitophiles. Annales de la Societe entomologique de France, series 2, 2, 333 - 426." type="journal article" year="1844">Schaum 1844</bibRefCitation>
: 367
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402D3AFFB4F1DAFC4D" author="Schaum" box="[812,997,920,942]" page="43" pageId="10" pageNumber="521" year="1845">
<bibRefCitation id="EFF5A7EDFFDEFC402D3AFFB4F182FC4D" author="Schaum" box="[812,957,920,942]" pageId="10" pageNumber="521" refString="Schaum, H. (1845) Catalogue des especes connues qui entrent dans la famille des Lamillicornes melitophiles. Annales de la Societe entomologique de France, series 2 (3), 37 - 56." type="journal article" year="1845">Schaum 1845</bibRefCitation>
: 43
</treatmentCitation>
; 
<treatmentCitation id="0AC5FC0DFFDEFC402DE5FFB4F693FC4D" author="Schaum" box="[1011,1196,920,942]" page="27" pageId="10" pageNumber="521" year="1848">
<bibRefCitation id="EFF5A7EDFFDEFC402DE5FFB4F6BCFC4D" author="Schaum" box="[1011,1155,920,942]" pageId="10" pageNumber="521" refString="Schaum, H. (1848) Ferzeichnis der Lamellicornia, Melitophila. Dr, Schaum, Szczecin, Poland, 74 pp." type="book" year="1848">Schaum 1848</bibRefCitation>
: 27
</treatmentCitation>
. (misapplication of the name)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E8997FFDEFC402E81FFD5F1C1FB25" pageId="10" pageNumber="521" type="diagnosis">
<paragraph id="8BDBDA1CFFDEFC402E81FFD5F1C1FB25" blockId="10.[151,1437,1017,2007]" pageId="10" pageNumber="521">
<emphasis id="B910060EFFDEFC402E81FFD5F32EFBF1" bold="true" box="[151,273,1017,1042]" pageId="10" pageNumber="521">Diagnosis.</emphasis>
Tomentum on body surface orange-red. The length of male antennal club about 1.5 times as the length of antennomeres 2–7 combined. Posterior margin of the median tomentose area of pronotum notched. Scutellum usually with arrow-shaped tomentose area (
<figureCitation id="135FC699FFDEFC402CBCF86DF102FBB9" box="[682,829,1089,1114]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">Figs. 52–53</figureCitation>
). Sutural maculae and anteapical maculae usually separated. Apex of mesometasternal process slightly curved; mesometasternal suture not engraved, almost without setae (
<figureCitation id="135FC699FFDEFC402EC9F8A5F30BFB41" box="[223,308,1161,1186]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">Fig. 63</figureCitation>
). Median portion of abdomen usually glabrous. Outer teeth of metatibia longer than ventral teeth in both sexes (
<figureCitation id="135FC699FFDEFC402F0AF881F34FFB25" box="[284,368,1197,1222]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">Fig. 70</figureCitation>
). Basal piece of male genitalia distinctly convex (
<figureCitation id="135FC699FFDEFC402D8BF881F1CDFB25" box="[925,1010,1197,1222]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="10" pageNumber="521">Fig. 80</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E8997FFDEFC402ED1F8FDF374FA7D" pageId="10" pageNumber="521" type="description">
<paragraph id="8BDBDA1CFFDEFC402ED1F8FDF374FA7D" blockId="10.[151,1437,1017,2007]" pageId="10" pageNumber="521">
<emphasis id="B910060EFFDEFC402ED1F8FDF300FB09" bold="true" box="[199,319,1233,1258]" pageId="10" pageNumber="521">Variation.</emphasis>
Body length 18.5–22.4 mm; width 8.8–10.8 mm. Sutural maculae and anteapical maculae occasionally connected in some individuals from western 
<collectingRegion id="49A014FEFFDEFC402D76F8DAF186FAEE" box="[864,953,1270,1293]" country="China" name="Yunnan" pageId="10" pageNumber="521">Yunnan</collectingRegion>
, 
<collectingCountry id="F3739A8CFFDEFC402DDBF8D9F62FFAED" box="[973,1040,1269,1294]" name="China" pageId="10" pageNumber="521">China</collectingCountry>
(
<figureCitation id="135FC699FFDEFC402A3FF8D9F6B9FAED" box="[1065,1158,1269,1294]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="10" pageNumber="521">Fig. 22</figureCitation>
). Posthumeral maculae sometimes absent (
<figureCitation id="135FC699FFDEFC402F7BF935F3CAFAD1" box="[365,501,1305,1330]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="10" pageNumber="521">Figs. 21, 23</figureCitation>
). Most individuals have arrow-shaped tomentose area on their scutellum (
<figureCitation id="135FC699FFDEFC402B25F935F289FAB5" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="10" pageNumber="521">Figs. 21– 22</figureCitation>
, 
<figureCitation id="135FC699FFDEFC402ED7F911F334FAB5" box="[193,267,1341,1366]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">52–53</figureCitation>
), but that of the four individuals from Mount Daweishan (
<collectingRegion id="49A014FEFFDEFC402D80F912F1CFFAB5" box="[918,1008,1342,1366]" country="China" name="Yunnan" pageId="10" pageNumber="521">Yunnan</collectingRegion>
, 
<collectingCountry id="F3739A8CFFDEFC402DEDF911F67FFAB5" box="[1019,1088,1341,1366]" name="China" pageId="10" pageNumber="521">China</collectingCountry>
) and central 
<collectingCountry id="F3739A8CFFDEFC402AC4F911F70EFAB5" box="[1234,1329,1341,1366]" name="Vietnam" pageId="10" pageNumber="521">Vietnam</collectingCountry>
is almost glabrous (
<figureCitation id="135FC699FFDEFC402F1CF94DF354FA99" box="[266,363,1377,1402]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="10" pageNumber="521">Figs. 23</figureCitation>
, 
<figureCitation id="135FC699FFDEFC402F6CF94DF3FBFA99" box="[378,452,1377,1402]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">54–56</figureCitation>
). A Chinese specimen from western 
<collectingRegion id="49A014FEFFDEFC402D63F94EF1F4FA9A" box="[885,971,1378,1401]" country="China" name="Yunnan" pageId="10" pageNumber="521">Yunnan</collectingRegion>
is setiferous along the mesometasternal suture (
<figureCitation id="135FC699FFDEFC402EFDF9A9F300FA7D" box="[235,319,1413,1438]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="10" pageNumber="521">Fig. 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E8997FFDEFC412ED1F985F393F9C3" lastPageId="11" lastPageNumber="522" pageId="10" pageNumber="521" type="materials_examined">
<paragraph id="8BDBDA1CFFDEFC402ED1F985F6B9FA21" blockId="10.[151,1437,1017,2007]" box="[199,1158,1449,1474]" pageId="10" pageNumber="521">
<emphasis id="B910060EFFDEFC402ED1F985F34AFA21" bold="true" box="[199,373,1449,1474]" pageId="10" pageNumber="521">
<typeStatus id="54DF64BEFFDEFC402ED1F985F33DFA21" box="[199,258,1449,1474]" pageId="10" pageNumber="521">Type</typeStatus>
material.
</emphasis>
The 
<typeStatus id="54DF64BEFFDEFC402FB8F986F3E1FA21" box="[430,478,1450,1474]" pageId="10" pageNumber="521">type</typeStatus>
series cannot be located, and are perhaps lost (
<bibRefCitation id="EFF5A7EDFFDEFC402DFDF986F645FA21" author="Arrow" box="[1003,1146,1450,1474]" pageId="10" pageNumber="521" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, United Kingdom, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
).
</paragraph>
<paragraph id="8BDBDA1CFFDEFC412ED1F9E1F393F9C3" blockId="10.[151,1437,1017,2007]" lastBlockId="11.[151,1437,151,2029]" lastPageId="11" lastPageNumber="522" pageId="10" pageNumber="521">
<emphasis id="B910060EFFDEFC402ED1F9E1F3C9FA05" bold="true" box="[199,502,1485,1510]" pageId="10" pageNumber="521">Other material examined</emphasis>
(63♂♂, 109♀♀). 
<materialsCitation id="3B0CD041FFDEFC402CDEF9E1F1F9FA05" ID-GBIF-Occurrence="1701220361" box="[712,966,1485,1510]" collectionCode="MFNB" country="Nepal" location="NEPAL" pageId="10" pageNumber="521" specimenCount="1" specimenCount-male="1">
<emphasis id="B910060EFFDEFC402CDEF9E1F111FA05" bold="true" box="[712,814,1485,1510]" pageId="10" pageNumber="521">
<collectingCountry id="F3739A8CFFDEFC402CDEF9E1F116FA05" box="[712,809,1485,1510]" name="Nepal" pageId="10" pageNumber="521">NEPAL</collectingCountry>
:
</emphasis>
<specimenCount id="9D621195FFDEFC402D20F9E2F166FA05" box="[822,857,1486,1510]" pageId="10" pageNumber="521" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402D7FF9E2F181FA05" box="[873,958,1486,1510]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="521">MFNB</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDEFC402DC4F9E2F2CDF9EA" ID-GBIF-Occurrence="1701220347" collectionCode="NHMB" collectorName="Brit. Ind. &amp; Coll. Dr. Itzinger" country="Nepal" location="Nepal" pageId="10" pageNumber="521" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D621195FFDEFC402DC4F9E2F1CAFA05" box="[978,1013,1486,1510]" pageId="10" pageNumber="521" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402A13F9E2F661FA05" box="[1029,1118,1486,1510]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="521">NHMB</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402A65F9E1F686FA05" box="[1139,1209,1485,1510]" name="Nepal" pageId="10" pageNumber="521">Nepal</collectingCountry>
, 
<collectorName id="2691BFCAFFDEFC402AD2F9E1F711FA05" box="[1220,1326,1485,1510]" pageId="10" pageNumber="521">Brit. Ind.</collectorName>
, 
<collectorName id="2691BFCAFFDEFC402B2CF9E1F2CDF9EA" pageId="10" pageNumber="521">Coll. Dr. Itzinger</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDEFC402F17F9DDF614F9EA" ID-GBIF-Occurrence="1701220399" box="[257,1067,1520,1545]" collectionCode="NHMB" country="Nepal" elevation="2" latitude="27.05" location="Pakhribas" longLatPrecision="1239" longitude="87.3" pageId="10" pageNumber="521" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D621195FFDEFC402F17F9DDF306F9EA" box="[257,313,1521,1545]" pageId="10" pageNumber="521" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402F5AF9DDF39AF9EA" box="[332,421,1521,1545]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="521">NHMB</collectionCode>
), Kosi-#3, 
<location id="8EBB8CC7FFDEFC402C3CF9DCF0A4F9EA" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDEFC402C3CF9DCF0A4F9EA" box="[554,667,1520,1545]" country="Nepal" latitude="27.05" longLatPrecision="1239" longitude="87.3" name="Pakhribas" pageId="10" pageNumber="521">Pakhribas</location>
, 
<elevation id="00493D2FFFDEFC402CBDF9DDF16FF9EA" box="[683,848,1521,1545]" metricMagnitude="0" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="10" pageNumber="521" unit="m" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">
<quantity id="4C9C77F9FFDEFC402CBDF9DDF16FF9EA" box="[683,848,1521,1545]" metricMagnitude="0" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="10" pageNumber="521" unit="m" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1,700–1,900m</quantity>
</elevation>
, 
<geoCoordinate id="EE50BCDBFFDEFC402D49F9DCF180F9EA" box="[863,959,1520,1545]" direction="north" orientation="latitude" pageId="10" pageNumber="521" precision="925" value="27.05">27°03′N</geoCoordinate>
, 
<geoCoordinate id="EE50BCDBFFDEFC402DD9F9DCF614F9EA" box="[975,1067,1520,1545]" direction="east" orientation="longitude" pageId="10" pageNumber="521" precision="925" value="87.3">87°18′E</geoCoordinate>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDEFC402A2AF9DDF07FF9CE" ID-GBIF-Occurrence="1701220389" collectingDate="2001-05-27" collectingDateMax="2001-06" collectingDateMin="2001-05-27" collectionCode="NHMB" collectorName="Basel" country="Nepal" location="Nepal" pageId="10" pageNumber="521" specimenCount="1">
<collectingDate id="EF9E0534FFDEFC402A2AF9DDF6E7F9EA" box="[1084,1240,1521,1545]" pageId="10" pageNumber="521" value="2001-05-27" valueMax="2001-05-28" valueMin="2001-05-27">
<date id="FFDAFCDCFFDEFC402A2AF9DDF6E7F9EA" box="[1084,1240,1521,1545]" pageId="10" pageNumber="521" value="2001-05-27" valueMax="2001-05-28" valueMin="2001-05-27">27–28.V.2001</date>
</collectingDate>
// 
<collectionCode id="ED7542D9FFDEFC402AE9F9DDF76CF9EB" box="[1279,1363,1521,1544]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="521">NHMB</collectionCode>
<collectorName id="2691BFCAFFDEFC402B48F9DCF7A3F9EA" box="[1374,1436,1520,1545]" pageId="10" pageNumber="521">Basel</collectorName>
expedition to 
<collectingCountry id="F3739A8CFFDEFC402F2DFA38F341F9CE" box="[315,382,1556,1581]" name="Nepal" pageId="10" pageNumber="521">Nepal</collectingCountry>
May/ 
<collectingDate id="EF9E0534FFDEFC402FD3FA39F07FF9CE" box="[453,576,1557,1581]" pageId="10" pageNumber="521" value="2001-06">
<date id="FFDAFCDCFFDEFC402FD3FA39F07FF9CE" box="[453,576,1557,1581]" pageId="10" pageNumber="521" value="2001-06">June 2001</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDEFC402C59FA38F184F9CE" ID-GBIF-Occurrence="1701220385" box="[591,955,1556,1581]" collectionCode="MNHN" country="India" location="Sikkim" pageId="10" pageNumber="521" specimenCount="1" specimenCount-male="1" stateProvince="Sikkim">
<emphasis id="B910060EFFDEFC402C59FA38F129F9CE" bold="true" box="[591,790,1556,1581]" pageId="10" pageNumber="521">
<collectingCountry id="F3739A8CFFDEFC402C59FA38F09AF9CE" box="[591,677,1556,1581]" name="India" pageId="10" pageNumber="521">INDIA</collectingCountry>
: 
<collectingRegion id="49A014FEFFDEFC402CA1FA38F12EF9CE" box="[695,785,1556,1581]" country="India" name="Sikkim" pageId="10" pageNumber="521">Sikkim</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDEFC402D34FA39F17BF9CE" box="[802,836,1557,1581]" pageId="10" pageNumber="521" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402D4EFA39F18CF9CE" box="[856,947,1557,1581]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="521">MNHN</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDEFC402DDAFA38F7A7F9CE" ID-GBIF-Occurrence="1701220367" box="[972,1432,1556,1581]" country="India" location="Sikkim" pageId="10" pageNumber="521" specimenCount="1" stateProvince="Sikkim">
<collectingCountry id="F3739A8CFFDEFC402DDAFA38F65FF9CE" box="[972,1120,1556,1581]" name="India" pageId="10" pageNumber="521">British India</collectingCountry>
, 
<collectingRegion id="49A014FEFFDEFC402A66FA38F6FBF9CE" box="[1136,1220,1556,1581]" country="India" name="Sikkim" pageId="10" pageNumber="521">Sikkim</collectingRegion>
, Lachen-Dachung
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDEFC402E81FA14F03AF9B3" ID-GBIF-Occurrence="1701220384" box="[151,517,1591,1616]" collectingDate="1933-08" collectionCode="IRSNB" country="India" location="Sikkim" pageId="10" pageNumber="521" specimenCount="1" specimenCount-female="1" stateProvince="Sikkim">
<collectingDate id="EF9E0534FFDEFC402E81FA14F336F9B3" box="[151,265,1592,1616]" pageId="10" pageNumber="521" value="1933-08">
<date id="FFDAFCDCFFDEFC402E81FA14F336F9B3" box="[151,265,1592,1616]" pageId="10" pageNumber="521" value="1933-08">VIII.1933</date>
</collectingDate>
; 
<specimenCount id="9D621195FFDEFC402F02FA14F308F9B3" box="[276,311,1592,1616]" pageId="10" pageNumber="521" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402F53FA14F3A2F9B3" box="[325,413,1592,1616]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="521">IRSNB</collectionCode>
), 
<collectingRegion id="49A014FEFFDEFC402FA6FA1BF03AF9B3" box="[432,517,1591,1616]" country="India" name="Sikkim" pageId="10" pageNumber="521">Sikkim</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDEFC402C19FA1BF2EDF990" ID-GBIF-Occurrence="1701220371" collectionCode="SDEI" collectorName="L. Durel" country="India" location="West Bengal" pageId="10" pageNumber="521" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal">
<emphasis id="B910060EFFDEFC402C19FA1BF093F9B3" bold="true" box="[527,684,1591,1616]" pageId="10" pageNumber="521">
<collectingRegion id="49A014FEFFDEFC402C19FA1BF098F9B3" box="[527,679,1591,1616]" country="India" name="West Bengal" pageId="10" pageNumber="521">West Bengal</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDEFC402CA5FA14F0E9F9B3" box="[691,726,1592,1616]" pageId="10" pageNumber="521" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402CF3FA14F117F9B3" box="[741,808,1592,1616]" pageId="10" pageNumber="521">SDEI</collectionCode>
), British Bootan, Padong [Pedong, Darjeeling], 
<collectorName id="2691BFCAFFDEFC402B28FA14F7A3F9B3" box="[1342,1436,1591,1616]" pageId="10" pageNumber="521">L. Durel</collectorName>
1891
</materialsCitation>
; 2♀♀ (BMNH, Bourgoin Coll.), 3♂♂, 7♀♀ (MNHN), British Bootan, L. Durel 1898; 2♀♀ (BMNH, Bourgoin Coll.), 1♂, 1♀ (IRSNB), British Bootan, Padong, L. Durel 1913; 1♂, 1♀ (BMNH, Bourgoin Coll.), 5♀♀ (MNHN), British Bootan, Padong, L. Durel 1914; 2♂♂, 2♀♀ (BMNH, Bourgoin Coll.), British Bootan, Maria Basti, L. Durel; 1♂, 2♀♀ (BMNH, Bourgoin Coll.), Darjeeling, Mowis 1891; 1♂, 
<materialsCitation id="3B0CD041FFDEFC402A53FAEAF382F8E1" ID-GBIF-Occurrence="1701220374" collectionCode="BMNH" collectorName="Kurseong Div. Lat Panchar" country="India" elevation="1219" location="India" pageId="10" pageNumber="521" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D621195FFDEFC402A53FAEAF657F93D" box="[1093,1128,1734,1758]" pageId="10" pageNumber="521" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402A6EFAEAF6EDF93D" box="[1144,1234,1734,1758]" pageId="10" pageNumber="521">BMNH</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402AF0FAE9F71BF93D" box="[1254,1316,1733,1758]" name="India" pageId="10" pageNumber="521">India</collectingCountry>
: 
<collectorName id="2691BFCAFFDEFC402B39FAEAF368F8E1" pageId="10" pageNumber="521">Kurseong Div. Lat Panchar</collectorName>
, 
<elevation id="00493D2FFFDEFC402F75FAC5F386F8E1" box="[355,441,1769,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="10" pageNumber="521" unit="ft" value="4000.0">
<quantity id="4C9C77F9FFDEFC402F75FAC5F386F8E1" box="[355,441,1769,1794]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="10" pageNumber="521" unit="ft" value="4000.0">4000 ft</quantity>
</elevation>
.
</materialsCitation>
, 
<date id="FFDAFCDCFFDEFC402FDCFAC5F017F8E2" box="[458,552,1769,1793]" pageId="10" pageNumber="521" value="1934-06">VI.1934</date>
, H.G. Champion; 1♂ (MFNB), Mungphu [Mungpoo]; 1♀ (SDEI), Darjeeling; 1♀ (SDEI), Pedong, Desgodins; 1♂ (MSNG), Mungphu 1890, D. Atkinson; 1♂ (NHMB), Pedong bei, Darjeeling 1935, Slg, R. Oberthür, Eing. Nr.4.1956; 1♀ (NHMB), Darjeeling; 1♂, 
<materialsCitation id="3B0CD041FFDEFC402DCDFB1DF0E0F88F" ID-GBIF-Occurrence="1701220373" collectionCode="MNHN" country="India" location="Inde" pageId="10" pageNumber="521" specimenCount="7" specimenCount-female="7">
<specimenCount id="9D621195FFDEFC402DCDFB1DF62CF8AB" box="[987,1043,1841,1864]" pageId="10" pageNumber="521" type="female">7♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402A33FB1CF6BEF8AB" box="[1061,1153,1840,1864]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="521">MNHN</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402A81FB03F6F6F8AB" box="[1175,1225,1839,1864]" name="India" pageId="10" pageNumber="521">Inde</collectingCountry>
Anglaise, Pedong Region de Darjeeling. Chasseurs indigenes 1933
</materialsCitation>
; 3♂♂, 
<materialsCitation id="3B0CD041FFDEFC402D56FB78F075F86C" ID-GBIF-Occurrence="1701220380" collectionCode="MNHN" country="India" location="Inde" pageId="10" pageNumber="521" specimenCount="3" specimenCount-female="3">
<specimenCount id="9D621195FFDEFC402D56FB78F148F88F" box="[832,887,1876,1900]" pageId="10" pageNumber="521" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402D98FB78F1D6F88F" box="[910,1001,1876,1900]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="521">MNHN</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402A12FB7FF609F88F" box="[1028,1078,1875,1900]" name="India" pageId="10" pageNumber="521">Inde</collectingCountry>
Anglaise, Pedong District de Darjeeling. Chasseurs indigenes 1932
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDEFC402C4FFB5BF37AF850" ID-GBIF-Occurrence="1701220378" collectionCode="MNHN" country="India" location="Inde" pageId="10" pageNumber="521" specimenCount="3" specimenCount-female="3">
<specimenCount id="9D621195FFDEFC402C4FFB5BF0AEF86C" box="[601,657,1911,1935]" pageId="10" pageNumber="521" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402CB5FB5BF0C0F86C" box="[675,767,1911,1935]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="521">MNHN</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402D03FB5AF178F86C" box="[789,839,1910,1935]" name="India" pageId="10" pageNumber="521">Inde</collectingCountry>
Anglaise, Pedong Region de Darjeeling. Chasseurs indigenes 1934
</materialsCitation>
; 1♂, 
<materialsCitation id="3B0CD041FFDEFC402F92FBB7F767F850" ID-GBIF-Occurrence="1701220390" box="[388,1368,1946,1971]" collectionCode="MNHN" country="India" location="Inde" pageId="10" pageNumber="521" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D621195FFDEFC402F92FBB7F398F850" box="[388,423,1947,1971]" pageId="10" pageNumber="521" type="female">2♀</specimenCount>
(
<collectionCode id="ED7542D9FFDEFC402FA1FBB7F02DF850" box="[439,530,1947,1971]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="521">MNHN</collectionCode>
), 
<collectingCountry id="F3739A8CFFDEFC402C30FBB6F067F850" box="[550,600,1946,1971]" name="India" pageId="10" pageNumber="521">Inde</collectingCountry>
Anglaise, Pedong Region de Darjeeling. Chasseurs indigenes 1937
</materialsCitation>
; 2♂♂ (MNHN), Maria Basti, Bootang; 1♀ (MNHN), British Bootang, L. Durel, 1899; 5♀♀ (MNHN), British Bootang, Maria Basti, L. Durel; 1♀ (MNHN), Kurseong, R.P. Decoly, 1898; 
<materialsCitation id="3B0CD041FFDFFC412DA4FCB4F325FF37" ID-GBIF-Occurrence="1701220362" collectionCode="MNHN" collectorName="R. P. Bretaudeau" country="India" location="Environs de Kurseong" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D621195FFDFFC412DA4FCB4F1EAFF53" box="[946,981,152,176]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DFEFCB4F67BFF53" box="[1000,1092,152,176]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="522">MNHN</collectionCode>
), 
<location id="8EBB8CC7FFDFFC412A4CFCBBF760FF53" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412A4CFCBBF760FF53" box="[1114,1375,151,176]" country="India" name="Environs de Kurseong" pageId="11" pageNumber="522">Environs de Kurseong</location>
, 
<collectorName id="2691BFCAFFDFFC412B78FCB4F325FF37" pageId="11" pageNumber="522">R.P. Bretaudeau</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412F31FC97F391FF37" ID-GBIF-Occurrence="1701220377" box="[295,430,187,212]" country="India" location="Assam" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Assam">
<emphasis id="B910060EFFDFFC412F31FC97F3BFFF37" bold="true" box="[295,384,187,212]" pageId="11" pageNumber="522">
<collectingRegion id="49A014FEFFDFFC412F31FC97F344FF37" box="[295,379,187,212]" country="India" name="Assam" pageId="11" pageNumber="522">Assam</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412F9FFC90F391FF37" box="[393,430,188,212]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412FABFC90F65AFF37" ID-GBIF-Occurrence="1701220393" box="[445,1125,187,212]" collectionCode="BMNH" collectorName="Fry Coll. &amp; Polkai Mount &amp; Doherty" country="India" location="Assam" pageId="11" pageNumber="522" specimenCount="4" specimenCount-female="4" stateProvince="Assam">
<specimenCount id="9D621195FFDFFC412FABFC90F3CBFF37" box="[445,500,188,212]" pageId="11" pageNumber="522" type="female">4♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412C10FC90F061FF37" box="[518,606,188,212]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412C7DFC90F0E9FF37" box="[619,726,187,212]" pageId="11" pageNumber="522">Fry Coll.</collectorName>
), 
<collectingRegion id="49A014FEFFDFFC412CFAFC90F104FF37" box="[748,827,188,212]" country="India" name="Assam" pageId="11" pageNumber="522">Assam</collectingRegion>
, 
<collectorName id="2691BFCAFFDFFC412D5FFC97F1DAFF30" box="[841,997,187,212]" pageId="11" pageNumber="522">Polkai Mount</collectorName>
// 
<collectorName id="2691BFCAFFDFFC412A10FC97F65AFF37" box="[1030,1125,187,212]" pageId="11" pageNumber="522">Doherty</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A65FC97F378FF14" ID-GBIF-Occurrence="1701220370" collectionCode="MNHN" collectorName="Oberthur Coll." country="India" location="Meghalaya" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Meghalaya">
<emphasis id="B910060EFFDFFC412A65FC97F73EFF37" bold="true" box="[1139,1281,187,212]" pageId="11" pageNumber="522">
<collectingRegion id="49A014FEFFDFFC412A65FC97F6C3FF37" box="[1139,1276,187,212]" country="India" name="Meghalaya" pageId="11" pageNumber="522">Meghalaya</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412B1CFC90F713FF37" box="[1290,1324,188,212]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412B2BFC90F7A8FF37" box="[1341,1431,188,212]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="522">MNHN</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412E81FCF2F300FF14" box="[151,319,222,247]" pageId="11" pageNumber="522">Oberthür Coll.</collectorName>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412F42FCF3F056FF14" ID-GBIF-Occurrence="1701220375" box="[340,617,222,248]" country="India" location="Assam Reg. de Shillong" pageId="11" pageNumber="522" specimenCount="1" stateProvince="Assam">
<location id="8EBB8CC7FFDFFC412F42FCF3F056FF14" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412F42FCF3F056FF14" box="[340,617,222,248]" country="India" name="Assam Reg. de Shillong" pageId="11" pageNumber="522" stateProvince="Assam">
<collectingRegion id="49A014FEFFDFFC412F42FCF3F39FFF14" box="[340,416,223,247]" country="India" name="Assam" pageId="11" pageNumber="522">Assam</collectingRegion>
Reg. de Shillong
</location>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412C63FCF2F2D4FEF8" ID-GBIF-Occurrence="1701220379" collectionCode="BMNH" collectorName="G. B. Rogers Coll. &amp; Asaam" country="India" elevation="2" location="northern Manipur" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Manipur">
<emphasis id="B910060EFFDFFC412C63FCF2F0D5FF14" bold="true" box="[629,746,222,247]" pageId="11" pageNumber="522">
<collectingRegion id="49A014FEFFDFFC412C63FCF2F0DAFF14" box="[629,741,222,247]" country="India" name="Manipur" pageId="11" pageNumber="522">Manipur</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412CE4FCF3F12BFF14" box="[754,788,223,247]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412D32FCF3F142FF14" box="[804,893,223,247]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412D9FFCF3F662FF14" box="[905,1117,222,248]" pageId="11" pageNumber="522">G. B. Rogers Coll.</collectorName>
), 
<location id="8EBB8CC7FFDFFC412A64FCF2F705FF14" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412A64FCF2F705FF14" box="[1138,1338,222,247]" country="India" name="northern Manipur" pageId="11" pageNumber="522" stateProvince="Manipur">
northern 
<collectingRegion id="49A014FEFFDFFC412ACEFCF2F705FF14" box="[1240,1338,222,247]" country="India" name="Manipur" pageId="11" pageNumber="522">Manipur</collectingRegion>
</location>
, 
<collectorName id="2691BFCAFFDFFC412B51FCF3F7A7FF14" box="[1351,1432,223,247]" pageId="11" pageNumber="522">Asaam</collectorName>
, 
<elevation id="00493D2FFFDFFC412E81FD2EF2D4FEF8" box="[151,235,258,283]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="522" unit="ft" value="6.0">
<quantity id="4C9C77F9FFDFFC412E81FD2EF2D4FEF8" box="[151,235,258,283]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" pageId="11" pageNumber="522" unit="ft" value="6.0">6,000ft</quantity>
</elevation>
</materialsCitation>
; 2♀♀ (BMNH, Nevinson Coll.), 
<materialsCitation id="3B0CD041FFDFFC412C72FD2EF628FEF8" ID-GBIF-Occurrence="1701220364" box="[612,1047,258,283]" collectionCode="BMNH" country="India" location="northern Manipur" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Manipur">
<collectingCountry id="F3739A8CFFDFFC412C72FD2EF09EFEF8" box="[612,673,258,283]" name="India" pageId="11" pageNumber="522">India</collectingCountry>
; 
<specimenCount id="9D621195FFDFFC412CBBFD2FF0EFFEF8" box="[685,720,259,283]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412CC9FD2FF107FEF8" box="[735,824,259,283]" pageId="11" pageNumber="522">BMNH</collectionCode>
), 
<location id="8EBB8CC7FFDFFC412D5BFD2EF628FEF8" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412D5BFD2EF628FEF8" box="[845,1047,258,283]" country="India" name="northern Manipur" pageId="11" pageNumber="522" stateProvince="Manipur">
northern 
<collectingRegion id="49A014FEFFDFFC412DA5FD2EF628FEF8" box="[947,1047,258,283]" country="India" name="Manipur" pageId="11" pageNumber="522">Manipur</collectingRegion>
</location>
</materialsCitation>
; 1♀ (BMNH), 
<materialsCitation id="3B0CD041FFDFFC412AD5FD2EF752FEF8" ID-GBIF-Occurrence="1701220369" box="[1219,1389,258,283]" country="India" location="Manipur" pageId="11" pageNumber="522" specimenCount="3" specimenCount-male="3" stateProvince="Manipur">
<collectingRegion id="49A014FEFFDFFC412AD5FD2EF718FEF8" box="[1219,1319,258,283]" country="India" name="Manipur" pageId="11" pageNumber="522">Manipur</collectingRegion>
; 
<specimenCount id="9D621195FFDFFC412B24FD2FF752FEF8" box="[1330,1389,259,283]" pageId="11" pageNumber="522" type="male">3♂♂</specimenCount>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412B6FFD2FF0D0FEDD" ID-GBIF-Occurrence="1701220383" collectionCode="BMNH" collectorName="Fry Coll. &amp; Doherty" country="India" location="Manipur" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Manipur">
<specimenCount id="9D621195FFDFFC412B6FFD2FF7A3FEF8" box="[1401,1436,259,283]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412E88FD0AF2C8FEDD" box="[158,247,294,318]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412F10FD0BF34DFEDD" box="[262,370,293,319]" pageId="11" pageNumber="522">Fry Coll.</collectorName>
), east 
<collectingCountry id="F3739A8CFFDFFC412FD6FD09F3C3FEDD" box="[448,508,293,318]" name="India" pageId="11" pageNumber="522">India</collectingCountry>
, 
<collectingRegion id="49A014FEFFDFFC412C1DFD09F054FEDD" box="[523,619,293,318]" country="India" name="Manipur" pageId="11" pageNumber="522">Manipur</collectingRegion>
// 
<collectorName id="2691BFCAFFDFFC412C87FD09F0D0FEDD" box="[657,751,293,318]" pageId="11" pageNumber="522">Doherty</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B0CD041FFDFFC412CEAFD09F031FE82" ID-GBIF-Occurrence="1701220382" collectingDate="1988-12" collectionCode="MSNG" collectorName="Carin Cheba &amp; Taungoo &amp; L. Fea" country="Myanmar" elevation="1" location="MYANMAR" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1">
<emphasis id="B910060EFFDFFC412CEAFD09F1A2FEDD" bold="true" box="[764,925,293,318]" pageId="11" pageNumber="522">
<collectingCountry id="F3739A8CFFDFFC412CEAFD09F1A7FEDD" box="[764,920,293,318]" name="Myanmar" pageId="11" pageNumber="522">MYANMAR</collectingCountry>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412DBEFD0AF1F4FEDD" box="[936,971,294,318]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DCBFD0AF60BFEDD" box="[989,1076,294,318]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="11" pageNumber="522">MSNG</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412A5DFD09F6E3FEDD" box="[1099,1244,293,318]" pageId="11" pageNumber="522">Carin Chebà</collectorName>
[near 
<collectorName id="2691BFCAFFDFFC412B3CFD0BF7AEFEDC" box="[1322,1425,295,319]" pageId="11" pageNumber="522">Taungoo</collectorName>
], 900– 
<elevation id="00493D2FFFDFFC412EC6FD65F30FFE82" box="[208,304,329,353]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="522" unit="m" value="1.1">
<quantity id="4C9C77F9FFDFFC412EC6FD65F30FFE82" box="[208,304,329,353]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="522" unit="m" value="1.1">1,100 m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412F56FD66F3B1FE82" box="[320,398,330,353]" pageId="11" pageNumber="522">L. Fea</collectorName>
, V– 
<collectingDate id="EF9E0534FFDFFC412FD7FD65F031FE82" box="[449,526,329,353]" pageId="11" pageNumber="522" value="1988-12">
<date id="FFDAFCDCFFDFFC412FD7FD65F031FE82" box="[449,526,329,353]" pageId="11" pageNumber="522" value="1988-12">XII-88</date>
</collectingDate>
</materialsCitation>
; 1♂ (MSNG), Carin Asciuii Chebà, 
<quantity id="4C9C77F9FFDFFC412DA8FD65F653FE82" box="[958,1132,329,353]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="11" pageNumber="522" unit="m" value="1.25" valueMax="1.3" valueMin="1.2">1,200–1,300 m</quantity>
, L. Fea, 
<date id="FFDAFCDCFFDFFC412ACFFD65F738FE82" box="[1241,1287,329,353]" pageId="11" pageNumber="522" value="1988-01">I-88</date>
/ 
<taxonomicName id="4C64A19FFFDFFC412B32FD65F2E8FE64" authority="Hope" authorityName="Hope" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDFFC412B32FD65F7A3FE81" box="[1316,1436,329,354]" italics="true" pageId="11" pageNumber="522">penicillata</emphasis>
Hope
</taxonomicName>
; 1♀ (BMNH), Burma, 1920-132; 3♂♂, 
<materialsCitation id="3B0CD041FFDFFC412CD2FD5CF0B3FE48" ID-GBIF-Occurrence="1701220388" collectionCode="MNHN" collectorName="H te Brimanie &amp; Mogok &amp; Doherty" country="Myanmar" elevation="2" location="Mines de Rubis" pageId="11" pageNumber="522" specimenCount="3" specimenCount-female="3" stateProvince="Mandalay">
<specimenCount id="9D621195FFDFFC412CD2FD5CF0C3FE6B" box="[708,764,368,392]" pageId="11" pageNumber="522" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412D04FD5CF151FE6B" box="[786,878,368,392]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="522">MNHN</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412D91FD5CF620FE6B" box="[903,1055,364,392]" pageId="11" pageNumber="522">H te Brimanie</collectorName>
, 
<location id="8EBB8CC7FFDFFC412A27FD43F6D0FE6B" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412A27FD43F6D0FE6B" box="[1073,1263,367,392]" country="Myanmar" name="Mines de Rubis" pageId="11" pageNumber="522" stateProvince="Mandalay">Mines de Rubis</location>
[near 
<collectorName id="2691BFCAFFDFFC412B52FD43F7A7FE6B" box="[1348,1432,367,392]" pageId="11" pageNumber="522">Mogok</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412E81FDBFF332FE4F" box="[151,269,403,428]" country="Myanmar" name="Mandalay" pageId="11" pageNumber="522">Mandalay</collectingRegion>
], 
<elevation id="00493D2FFFDFFC412F35FDBFF3ECFE48" box="[291,467,403,427]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="11" pageNumber="522" unit="m" value="1.75" valueMax="2.3" valueMin="1.2">
<quantity id="4C9C77F9FFDFFC412F35FDBFF3ECFE48" box="[291,467,403,427]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="11" pageNumber="522" unit="m" value="1.75" valueMax="2.3" valueMin="1.2">1,200–2,300 m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412FF2FDBFF07EFE4F" box="[484,577,403,428]" pageId="11" pageNumber="522">Doherty</collectorName>
, 1880
</materialsCitation>
. 
<materialsCitation id="3B0CD041FFDFFC412C8DFDBFF61EFE2D" ID-GBIF-Occurrence="1701220381" collectingDate="2016-06" collectionCode="QCCC" collectorName="Yarlung Tsangpo Grand Canyon Nature &amp; Beibeng &amp; Wei Hu" country="China" county="Medog County" elevation="2" location="Medog County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Xizang">
<emphasis id="B910060EFFDFFC412C8DFDBFF154FE4F" bold="true" box="[667,875,403,428]" pageId="11" pageNumber="522">
<collectingCountry id="F3739A8CFFDFFC412C8DFDBFF0C3FE4F" box="[667,764,403,428]" name="China" pageId="11" pageNumber="522">CHINA</collectingCountry>
: 
<collectingRegion id="49A014FEFFDFFC412D1BFDBFF15AFE4F" box="[781,869,403,428]" country="China" name="Xizang" pageId="11" pageNumber="522">Xizang</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412D61FDBFF1A5FE48" box="[887,922,403,427]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DB8FDBFF63FFE48" box="[942,1024,403,427]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412A01FDBFF649FE4F" box="[1047,1142,403,428]" pageId="11" pageNumber="522" value="2016-06">
<date id="FFDAFCDCFFDFFC412A01FDBFF649FE4F" box="[1047,1142,403,428]" pageId="11" pageNumber="522" value="2016-06">VI.2016</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412A91FDBFF379FE2D" pageId="11" pageNumber="522">Yarlung Tsangpo Grand Canyon Nature</collectorName>
Reserve, Hanmi, 
<collectorName id="2691BFCAFFDFFC412C09FD9AF040FE2C" box="[543,639,438,463]" pageId="11" pageNumber="522">Beibeng</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412C86FD9AF101FE2C" box="[656,830,438,463]" pageId="11" pageNumber="522">Mêdog County</collectingCounty>
, 
<elevation id="00493D2FFFDFFC412D59FD9BF18DFE2D" box="[847,946,439,463]" metricMagnitude="0" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="522" unit="m" value="2.2">
<quantity id="4C9C77F9FFDFFC412D59FD9BF18DFE2D" box="[847,946,439,463]" metricMagnitude="0" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="522" unit="m" value="2.2">2,200 m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412DD5FD9AF61EFE2D" box="[963,1057,438,463]" pageId="11" pageNumber="522">Wei Hu</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A27FD9AF3D7FE10" ID-GBIF-Occurrence="1701220392" collectionCode="BMNH" collectorName="Sima" country="China" location="Yunnan" pageId="11" pageNumber="522" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan">
<emphasis id="B910060EFFDFFC412A27FD9AF6A7FE2C" bold="true" box="[1073,1176,438,463]" pageId="11" pageNumber="522">
<collectingRegion id="49A014FEFFDFFC412A27FD9AF6ACFE2C" box="[1073,1171,438,463]" country="China" name="Yunnan" pageId="11" pageNumber="522">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412AB2FD9BF6E2FE2C" box="[1188,1245,439,463]" pageId="11" pageNumber="522" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412AE7FD9BF775FE2C" box="[1265,1354,439,463]" pageId="11" pageNumber="522">BMNH</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412B74FD9AF7A3FE2C" box="[1378,1436,438,463]" pageId="11" pageNumber="522">Sima</collectorName>
[Xima, Yingjiang], 1914-145
</materialsCitation>
; 1♂ (BMNH), Zhehou, Jingdong, 
<date id="FFDAFCDCFFDFFC412D6DFDF6F1CEFE10" box="[891,1009,474,499]" pageId="11" pageNumber="522" value="1978-05-15">15.V.1978</date>
; 1♀ (BMNH), 
<date id="FFDAFCDCFFDFFC412AB0FDF6F724FE10" box="[1190,1307,474,499]" pageId="11" pageNumber="522" value="1985-05-24">24.V.1985</date>
, Menghun, Xishuangbanna, 
<quantity id="4C9C77F9FFDFFC412F42FDD2F03FFDF6" box="[340,512,510,534]" metricMagnitude="0" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="11" pageNumber="522" unit="m" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1,200–1,400 m</quantity>
; 1870; 
<materialsCitation id="3B0CD041FFDFFC412C42FDD2F30AFDDA" ID-GBIF-Occurrence="1701220372" collectingDate="2015-05-21" collectionCode="SWU" collectorName="Jian-Yue Qiu" country="China" county="Hekou County" location="Mount Daweishan Nature Reserve" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412C42FDD2F048FDF5" box="[596,631,510,534]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412C91FDD2F0F4FDF5" box="[647,715,510,534]" country="South Korea" httpUri="http://grbio.org/cool/71hc-n1fq" name="Sungshin Women's University" pageId="11" pageNumber="522">SWU</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412CC8FDD1F16CFDF5" box="[734,851,509,534]" pageId="11" pageNumber="522" value="2015-05-21">
<date id="FFDAFCDCFFDFFC412CC8FDD1F16CFDF5" box="[734,851,509,534]" pageId="11" pageNumber="522" value="2015-05-21">21.V.2015</date>
</collectingDate>
, 
<location id="8EBB8CC7FFDFFC412D49FDD2F6D8FDF6" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412D49FDD2F6D8FDF6" box="[863,1255,509,534]" country="China" county="Hekou County" name="Mount Daweishan Nature Reserve" pageId="11" pageNumber="522" stateProvince="Yunnan">Mount Daweishan Nature Reserve</location>
, 
<collectingCounty id="62BAA290FFDFFC412AE4FDD1F7A6FDF5" box="[1266,1433,509,534]" pageId="11" pageNumber="522">Hekou County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412E81FE0CF30AFDDA" box="[151,309,544,569]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
</materialsCitation>
; 1♂, 
<materialsCitation id="3B0CD041FFDFFC412F96FE0DF078FDBE" ID-GBIF-Occurrence="1701220419" collectingDate="2015-05-21" collectingDateMax="2015-06" collectingDateMin="2015-05-21" collectionCode="SWU" collectorName="Jian-Yue Qiu" country="China" county="Hekou County" location="Mount Daweishan Nature Reserve" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412F96FE0DF39CFDDA" box="[384,419,545,569]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412FAFFE0DF3C2FDDA" box="[441,509,545,569]" country="South Korea" httpUri="http://grbio.org/cool/71hc-n1fq" name="Sungshin Women's University" pageId="11" pageNumber="522">SWU</collectionCode>
), 
<location id="8EBB8CC7FFDFFC412C0EFE0DF18CFDDB" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412C0EFE0DF18CFDDB" box="[536,947,544,569]" country="China" county="Hekou County" name="Mount Daweishan Nature Reserve" pageId="11" pageNumber="522" stateProvince="Yunnan">Mount Daweishan Nature Reserve</location>
, 
<collectingCounty id="62BAA290FFDFFC412DD0FE0CF64CFDDA" box="[966,1139,544,569]" pageId="11" pageNumber="522">Hekou County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412A93FE0CF71CFDDA" box="[1157,1315,544,569]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
, pupae in 
<collectingDate id="EF9E0534FFDFFC412E81FE68F333FDBE" box="[151,268,580,605]" pageId="11" pageNumber="522" value="2015-05-21">
<date id="FFDAFCDCFFDFFC412E81FE68F333FDBE" box="[151,268,580,605]" pageId="11" pageNumber="522" value="2015-05-21">21.V.2015</date>
</collectingDate>
, adults emerged in 
<collectingDate id="EF9E0534FFDFFC412FF1FE68F078FDBE" box="[487,583,580,605]" pageId="11" pageNumber="522" value="2015-06">
<date id="FFDAFCDCFFDFFC412FF1FE68F078FDBE" box="[487,583,580,605]" pageId="11" pageNumber="522" value="2015-06">VI.2015</date>
</collectingDate>
</materialsCitation>
; 2♂♂, 
<materialsCitation id="3B0CD041FFDFFC412C8CFE69F377FD63" ID-GBIF-Occurrence="1701220418" collectingDate="2014-05" collectionCode="QCCC" collectorName="Manjiu &amp; Manwan &amp; Lincang &amp; Zi-Chun Xiong" country="China" county="Yunxian County" location="Yunxian County" pageId="11" pageNumber="522" specimenCount="7" specimenCount-female="7" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412C8CFE69F0EDFDBF" box="[666,722,581,604]" pageId="11" pageNumber="522" type="female">7♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412CF7FE69F10CFDBE" box="[737,819,581,605]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412D51FE69F1A5FDBE" box="[839,922,581,605]" pageId="11" pageNumber="522" value="2014-05">
<date id="FFDAFCDCFFDFFC412D51FE69F1A5FDBE" box="[839,922,581,605]" pageId="11" pageNumber="522" value="2014-05">V.2014</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412DB3FE68F1C6FDBE" box="[933,1017,580,605]" pageId="11" pageNumber="522">Manjiu</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412A13FE69F655FDBF" box="[1029,1130,581,604]" pageId="11" pageNumber="522">Manwan</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412A63FE68F711FDBE" box="[1141,1326,580,605]" pageId="11" pageNumber="522">Yunxian County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412B2CFE68F7A7FDBE" box="[1338,1432,580,605]" pageId="11" pageNumber="522">Lincang</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412E81FE4BF377FD63" box="[151,328,615,640]" pageId="11" pageNumber="522">Zi-Chun Xiong</collectorName>
</materialsCitation>
; 3♀♀ (QCCC), F1 generation of the female from Lincang, Zi-Chun Xiong, eggs in 
<date id="FFDAFCDCFFDFFC412B13FE44F767FD63" box="[1285,1368,616,640]" pageId="11" pageNumber="522" value="2014-05">V.2014</date>
, adult emerged in 
<date id="FFDAFCDCFFDFFC412F3FFEA3F3B5FD44" box="[297,394,655,679]" pageId="11" pageNumber="522" value="2014-09">IX.2014</date>
; 
<materialsCitation id="3B0CD041FFDFFC412F8AFEA3F6CDFD44" ID-GBIF-Occurrence="1701220407" box="[412,1266,654,679]" collectingDate="2013-08-10" collectionCode="QCCC" collectorName="Huguo &amp; Guang-Lin Xie" country="China" county="Longchuan County" location="Longchuan County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412F8AFEA3F380FD44" box="[412,447,655,679]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412FC3FEA3F018FD44" box="[469,551,655,679]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412C54FEA3F0E8FD44" box="[578,727,655,679]" pageId="11" pageNumber="522" value="2013-08-10">
<date id="FFDAFCDCFFDFFC412C54FEA3F0E8FD44" box="[578,727,655,679]" pageId="11" pageNumber="522" value="2013-08-10">10.VIII.2013</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412CFFFEA3F107FD44" box="[745,824,655,679]" pageId="11" pageNumber="522">Huguo</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412D5DFEA2F614FD44" box="[843,1067,654,679]" pageId="11" pageNumber="522">Longchuan County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412A2BFEA2F6CDFD44" box="[1085,1266,654,679]" pageId="11" pageNumber="522">Guang-Lin Xie</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412B13FEA3F1E2FD2A" ID-GBIF-Occurrence="1701220391" collectingDate="2015-06-12" collectionCode="QCCC" collectorName="Kangjiaba &amp; Wumulong &amp; Mao Ye" country="China" county="Lincang County" location="Lincang County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412B13FEA3F717FD44" box="[1285,1320,655,679]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412B28FEA3F7AFFD44" box="[1342,1424,655,679]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412E81FE9DF326FD29" box="[151,281,689,714]" pageId="11" pageNumber="522" value="2015-06-12">
<date id="FFDAFCDCFFDFFC412E81FE9DF326FD29" box="[151,281,689,714]" pageId="11" pageNumber="522" value="2015-06-12">12.VI.2015</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412F3CFE9DF3A2FD29" box="[298,413,689,714]" pageId="11" pageNumber="522">Kangjiaba</collectorName>
reservoir, 
<collectorName id="2691BFCAFFDFFC412C09FE9DF0A0FD29" box="[543,671,689,714]" pageId="11" pageNumber="522">Wumulong</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412CB9FE9DF154FD29" box="[687,875,689,714]" pageId="11" pageNumber="522">Lincang County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412D6DFE9EF1E2FD2A" box="[891,989,690,713]" pageId="11" pageNumber="522">Mao Ye</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412DFBFE9EF616FD0E" ID-GBIF-Occurrence="1701220413" collectingDate="2016-02-18" collectionCode="QCCC" collectorName="Tongbiguan Nature Reserve &amp; Dehong" country="China" county="Yingjiang County" elevation="2" location="Yingjiang County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412DFBFE9EF62FFD29" box="[1005,1040,690,714]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A32FE9EF649FD29" box="[1060,1142,690,714]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412A99FE9DF739FD29" box="[1167,1286,689,714]" pageId="11" pageNumber="522" value="2016-02-18">
<date id="FFDAFCDCFFDFFC412A99FE9DF739FD29" box="[1167,1286,689,714]" pageId="11" pageNumber="522" value="2016-02-18">18.II.2016</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412B01FE9DF375FD0E" pageId="11" pageNumber="522">Tongbiguan Nature Reserve</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412F4EFEF8F01BFD0E" box="[344,548,724,749]" pageId="11" pageNumber="522">Yingjiang County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412C24FEF8F0B2FD0E" box="[562,653,724,749]" pageId="11" pageNumber="522">Dehong</collectorName>
, 
<elevation id="00493D2FFFDFFC412C8DFEF9F0C3FD0E" box="[667,764,725,749]" metricMagnitude="0" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="522" unit="m" value="2.1">
<quantity id="4C9C77F9FFDFFC412C8DFEF9F0C3FD0E" box="[667,764,725,749]" metricMagnitude="0" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="522" unit="m" value="2.1">2,100 m</quantity>
</elevation>
, Jian-Yue Qiu &amp; Hao Xu
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A21FEF9F38DFCD4" ID-GBIF-Occurrence="1701220395" collectingDate="2016-02-18" collectingDateMax="2016-05" collectingDateMin="2016-02-18" collectionCode="QCCC" collectorName="Bangzhongdazhai &amp; Rd &amp; Nongzhang &amp; Dehong" country="China" county="Yingjiang County" elevation="1" location="Yingjiang County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412A21FEF9F665FD0E" box="[1079,1114,725,749]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A7AFEF9F681FD0E" box="[1132,1214,725,749]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412AC2FEF8F7A2FD0E" box="[1236,1437,724,749]" pageId="11" pageNumber="522">Bangzhongdazhai</collectorName>
(S233 
<collectorName id="2691BFCAFFDFFC412EF2FED7F337FCF7" box="[228,264,763,788]" pageId="11" pageNumber="522">Rd</collectorName>
), 
<collectorName id="2691BFCAFFDFFC412F09FED7F39CFCF7" box="[287,419,763,788]" pageId="11" pageNumber="522">Nongzhang</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412FA7FED7F042FCF7" box="[433,637,763,788]" pageId="11" pageNumber="522">Yingjiang County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412C9DFED7F0D8FCF7" box="[651,743,763,788]" pageId="11" pageNumber="522">Dehong</collectorName>
, 
<elevation id="00493D2FFFDFFC412CE3FED0F169FCF0" box="[757,854,764,788]" metricMagnitude="0" metricUnit="m" metricValue="1.32" pageId="11" pageNumber="522" unit="m" value="1.32">
<quantity id="4C9C77F9FFDFFC412CE3FED0F169FCF0" box="[757,854,764,788]" metricMagnitude="0" metricUnit="m" metricValue="1.32" pageId="11" pageNumber="522" unit="m" value="1.32">1,320 m</quantity>
</elevation>
, Jian-Yue Qiu &amp; Hao Xu, 3nd instar in 
<collectingDate id="EF9E0534FFDFFC412B36FED7F7A7FCF7" box="[1312,1432,763,788]" pageId="11" pageNumber="522" value="2016-02-18">
<date id="FFDAFCDCFFDFFC412B36FED7F7A7FCF7" box="[1312,1432,763,788]" pageId="11" pageNumber="522" value="2016-02-18">18.II.2016</date>
</collectingDate>
, adult emerged in 
<collectingDate id="EF9E0534FFDFFC412F49FF32F38DFCD4" box="[351,434,798,823]" pageId="11" pageNumber="522" value="2016-05">
<date id="FFDAFCDCFFDFFC412F49FF32F38DFCD4" box="[351,434,798,823]" pageId="11" pageNumber="522" value="2016-05">V.2016</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412FA9FF33F7A8FCD4" ID-GBIF-Occurrence="1701220397" box="[447,1431,798,823]" collectingDate="2015-06" collectionCode="QCCC" collectorName="Xinping &amp; Gasha &amp; Huang-Shun Xi" country="China" elevation="2" location="Mount Ailaoshan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412FA9FF33F3DDFCD4" box="[447,482,799,823]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412FE5FF33F07AFCD4" box="[499,581,799,823]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412C4CFF32F086FCD4" box="[602,697,798,823]" pageId="11" pageNumber="522" value="2015-06">
<date id="FFDAFCDCFFDFFC412C4CFF32F086FCD4" box="[602,697,798,823]" pageId="11" pageNumber="522" value="2015-06">VI.2015</date>
</collectingDate>
, 
<location id="8EBB8CC7FFDFFC412CD0FF0CF1B0FCD4" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412CD0FF0CF1B0FCD4" box="[710,911,798,823]" country="China" name="Mount Ailaoshan" pageId="11" pageNumber="522" stateProvince="Yunnan">Mount Ailaoshan</location>
, 
<collectorName id="2691BFCAFFDFFC412D8DFF32F1C8FCD4" box="[923,1015,798,823]" pageId="11" pageNumber="522">Xinping</collectorName>
to 
<collectorName id="2691BFCAFFDFFC412A09FF32F657FCD4" box="[1055,1128,798,823]" pageId="11" pageNumber="522">Gasha</collectorName>
, 
<elevation id="00493D2FFFDFFC412A62FF33F6EBFCD4" box="[1140,1236,799,823]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="522" unit="m" value="1.9">
<quantity id="4C9C77F9FFDFFC412A62FF33F6EBFCD4" box="[1140,1236,799,823]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="522" unit="m" value="1.9">1,900 m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412AF4FF32F7A8FCD4" box="[1250,1431,798,823]" pageId="11" pageNumber="522">Huang-Shun Xi</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412E81FF6EF625FCB8" ID-GBIF-Occurrence="1701220394" box="[151,1050,834,859]" collectingDate="1984-05-12" collectionCode="NKUM" collectorName="Fangjiajing &amp; Liu" country="China" county="Jingdong County" location="Mount Ailaoshan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412E81FF6EF285FCB9" box="[151,186,834,858]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412EDCFF6EF31AFCB9" box="[202,293,834,858]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="11" pageNumber="522">NKUM</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412F2CFF6EF390FCB9" box="[314,431,834,858]" pageId="11" pageNumber="522" value="1984-05-12">
<date id="FFDAFCDCFFDFFC412F2CFF6EF390FCB9" box="[314,431,834,858]" pageId="11" pageNumber="522" value="1984-05-12">12.V.1984</date>
</collectingDate>
, 
<location id="8EBB8CC7FFDFFC412FABFF6FF0BAFCB8" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412FABFF6FF0BAFCB8" box="[445,645,834,859]" country="China" county="Jingdong County" name="Mount Ailaoshan" pageId="11" pageNumber="522" stateProvince="Yunnan">Mount Ailaoshan</location>
, 
<collectorName id="2691BFCAFFDFFC412C87FF6EF12BFCB8" box="[657,788,834,859]" pageId="11" pageNumber="522">Fangjiajing</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412D37FF6EF1DBFCB9" box="[801,996,834,859]" pageId="11" pageNumber="522">Jingdong County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412DE6FF6EF625FCB8" box="[1008,1050,834,859]" pageId="11" pageNumber="522">Liu</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A31FF6EF67DFC9E" ID-GBIF-Occurrence="1701220406" collectingDate="2017-01-30" collectionCode="QCCC" collectorName="Zhutang &amp; Puer &amp; Jian-Yue Qiu &amp; Hao Xu" country="China" county="Lancang County" location="Lancang County" pageId="11" pageNumber="522" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412A31FF6EF660FCB9" box="[1063,1119,834,858]" pageId="11" pageNumber="522" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A66FF6EF6FDFCB9" box="[1136,1218,834,858]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412AC1FF6EF77BFCB9" box="[1239,1348,834,858]" pageId="11" pageNumber="522" value="2017-01-30">
<date id="FFDAFCDCFFDFFC412AC1FF6EF77BFCB9" box="[1239,1348,834,858]" pageId="11" pageNumber="522" value="2017-01-30">30.I.2017</date>
</collectingDate>
, Mount Xiaolushan, Cizhuhe, 
<collectorName id="2691BFCAFFDFFC412F8FFF49F3C6FC9D" box="[409,505,869,894]" pageId="11" pageNumber="522">Zhutang</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412C1FFF4AF0F6FC9D" box="[521,713,870,894]" pageId="11" pageNumber="522">Lancang County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412CCEFF4AF132FC9E" box="[728,781,870,893]" pageId="11" pageNumber="522">Puer</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412D0BFF49F18AFC9D" box="[797,949,869,894]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
&amp; 
<collectorName id="2691BFCAFFDFFC412DF4FF4AF67DFC9E" box="[994,1090,870,893]" pageId="11" pageNumber="522">Hao Xu</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A44FF4AF350FC26" ID-GBIF-Occurrence="1701220404" collectingDate="2017-01-31" collectingDateMax="2017-05" collectingDateMin="2017-01-31" collectionCode="QCCC" collectorName="Bulong Nature Reserve &amp; Bulangshan &amp; Xishuanbanna &amp; Jian-Yue Qiu &amp; Hao Xu" country="China" county="Menghai County" location="Menghai County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412A44FF4AF64AFC9D" box="[1106,1141,870,894]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A9EFF4AF6E5FC9D" box="[1160,1242,870,894]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412AE4FF49F2CBFC42" pageId="11" pageNumber="522">Bulong Nature Reserve</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412F16FFA5F3B8FC41" box="[256,391,905,930]" pageId="11" pageNumber="522">Bulangshan</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412F82FFA5F06CFC42" box="[404,595,905,930]" pageId="11" pageNumber="522">Menghai County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412C76FFA5F13CFC41" box="[608,771,905,930]" pageId="11" pageNumber="522">Xishuanbanna</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412D06FFA5F19AFC41" box="[784,933,905,930]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
&amp; 
<collectorName id="2691BFCAFFDFFC412DDDFFA6F618FC42" box="[971,1063,906,929]" pageId="11" pageNumber="522">Hao Xu</collectorName>
, mature larva in 
<collectingDate id="EF9E0534FFDFFC412AFFFFA5F768FC42" box="[1257,1367,905,929]" pageId="11" pageNumber="522" value="2017-01-31">
<date id="FFDAFCDCFFDFFC412AFFFFA5F768FC42" box="[1257,1367,905,929]" pageId="11" pageNumber="522" value="2017-01-31">31.I.2017</date>
</collectingDate>
, adult emerged in 
<collectingDate id="EF9E0534FFDFFC412F0DFF81F350FC26" box="[283,367,941,965]" pageId="11" pageNumber="522" value="2017-05">
<date id="FFDAFCDCFFDFFC412F0DFF81F350FC26" box="[283,367,941,965]" pageId="11" pageNumber="522" value="2017-05">V.2017</date>
</collectingDate>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412F6CFF81F161FC0B" ID-GBIF-Occurrence="1701220414" collectingDate="2017-02-01" collectingDateMax="2017-05" collectingDateMin="2017-02-01" collectionCode="QCCC" collectorName="Yiwu &amp; Xishuanbanna &amp; Jian-Yue Qiu &amp; Hao Xu" country="China" county="Mengla County" location="Mengla County" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="9D621195FFDFFC412F6CFF81F3A2FC26" box="[378,413,941,965]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412FBAFF81F3C1FC26" box="[428,510,941,965]" pageId="11" pageNumber="522">QCCC</collectionCode>
), Xujialiangzi, Manla, 
<collectorName id="2691BFCAFFDFFC412D16FF80F17FFC26" box="[768,832,940,965]" pageId="11" pageNumber="522">Yiwu</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412D5DFF80F1C3FC26" box="[843,1020,940,965]" pageId="11" pageNumber="522">Mengla County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412A11FF80F695FC26" box="[1031,1194,940,965]" pageId="11" pageNumber="522">Xishuanbanna</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412AA0FF80F776FC26" box="[1206,1353,940,965]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
&amp; 
<collectorName id="2691BFCAFFDFFC412B7BFF81F283FC0B" pageId="11" pageNumber="522">Hao Xu</collectorName>
, larva in pupal cell in 
<collectingDate id="EF9E0534FFDFFC412FD1FFFCF00FFC0B" box="[455,560,976,1000]" pageId="11" pageNumber="522" value="2017-02-01">
<date id="FFDAFCDCFFDFFC412FD1FFFCF00FFC0B" box="[455,560,976,1000]" pageId="11" pageNumber="522" value="2017-02-01">1.II.2017</date>
</collectingDate>
, adult emerged in 
<collectingDate id="EF9E0534FFDFFC412D1CFFFCF161FC0B" box="[778,862,976,1000]" pageId="11" pageNumber="522" value="2017-05">
<date id="FFDAFCDCFFDFFC412D1CFFFCF161FC0B" box="[778,862,976,1000]" pageId="11" pageNumber="522" value="2017-05">V.2017</date>
</collectingDate>
</materialsCitation>
; 1♂ (KIZ), 
<materialsCitation id="3B0CD041FFDFFC412DFDFFFCF61AFBEF" ID-GBIF-Occurrence="1701220376" collectingDate="2017-05-06" collectionCode="QCCC" collectorName="Zhong-Cheng Tian" country="China" county="Lvchun County" elevation="1" location="Mount Huanglianshan Nature Reserve" pageId="11" pageNumber="522" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan">
<collectingDate id="EF9E0534FFDFFC412DFDFFFCF66CFC0A" box="[1003,1107,976,1001]" pageId="11" pageNumber="522" value="2017-05-06">
<date id="FFDAFCDCFFDFFC412DFDFFFCF66CFC0A" box="[1003,1107,976,1001]" pageId="11" pageNumber="522" value="2017-05-06">6.V.2017</date>
</collectingDate>
, erfu, 
<location id="8EBB8CC7FFDFFC412A89FFFDF373FBE8" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412A89FFFDF373FBE8" country="China" county="Lvchun County" name="Mount Huanglianshan Nature Reserve" pageId="11" pageNumber="522" stateProvince="Yunnan">Mount Huanglianshan Nature Reserve</location>
, 
<collectingCounty id="62BAA290FFDFFC412F4BFFDFF02CFBEF" box="[349,531,1011,1036]" pageId="11" pageNumber="522">Lvchun County</collectingCounty>
, 
<elevation id="00493D2FFFDFFC412C35FFD8F0B8FBE8" box="[547,647,1012,1036]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="522" unit="m" value="1.3">
<quantity id="4C9C77F9FFDFFC412C35FFD8F0B8FBE8" box="[547,647,1012,1036]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="522" unit="m" value="1.3">1,300 m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412C8EFFDFF14CFBEF" box="[664,883,1011,1036]" pageId="11" pageNumber="522">Zhong-Cheng Tian</collectorName>
<specimenCount id="9D621195FFDFFC412D69FFD8F188FBEF" box="[895,951,1012,1036]" pageId="11" pageNumber="522" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DDDFFD8F622FBEF" box="[971,1053,1012,1036]" pageId="11" pageNumber="522">QCCC</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412A20FFD8F6C6FBCC" ID-GBIF-Occurrence="1701220402" collectingDate="2017-02-01" collectionCode="NHMB" collectorName="Yiwu &amp; Xishuanbanna &amp; Jian-Yue Qiu &amp; Hao Xu." country="Thailand" county="Mengla County" location="Mengla County" pageId="11" pageNumber="522" specimenCount="3" specimenCount-male="3">
<collectingDate id="EF9E0534FFDFFC412A20FFD8F6A1FBEF" box="[1078,1182,1012,1036]" pageId="11" pageNumber="522" value="2017-02-01">
<date id="FFDAFCDCFFDFFC412A20FFD8F6A1FBEF" box="[1078,1182,1012,1036]" pageId="11" pageNumber="522" value="2017-02-01">1.II.2017</date>
</collectingDate>
, Xujialiangzi, Manla, 
<collectorName id="2691BFCAFFDFFC412E81F83AF2E8FBCC" box="[151,215,1046,1071]" pageId="11" pageNumber="522">Yiwu</collectorName>
, 
<collectingCounty id="62BAA290FFDFFC412EF1F83AF3A1FBCC" box="[231,414,1046,1071]" pageId="11" pageNumber="522">Mengla County</collectingCounty>
, 
<collectorName id="2691BFCAFFDFFC412FB8F83AF06EFBCC" box="[430,593,1046,1071]" pageId="11" pageNumber="522">Xishuanbanna</collectorName>
, 
<collectorName id="2691BFCAFFDFFC412C75F83AF0C3FBCC" box="[611,764,1046,1071]" pageId="11" pageNumber="522">Jian-Yue Qiu</collectorName>
&amp; 
<collectorName id="2691BFCAFFDFFC412D3DF834F1AEFBCC" box="[811,913,1048,1071]" pageId="11" pageNumber="522">Hao Xu.</collectorName>
<emphasis id="B910060EFFDFFC412D8BF83AF602FBCC" bold="true" box="[925,1085,1046,1071]" pageId="11" pageNumber="522">
<collectingCountry id="F3739A8CFFDFFC412D8BF83AF607FBCC" box="[925,1080,1046,1071]" name="Thailand" pageId="11" pageNumber="522">THAILAND</collectingCountry>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412A5CF83BF6BDFBCC" box="[1098,1154,1047,1071]" pageId="11" pageNumber="522" type="male">3♂♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A81F83BF6CFFBCC" box="[1175,1264,1047,1071]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="11" pageNumber="522">NHMB</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412B1CF83AF10CFBB0" ID-GBIF-Occurrence="1701220409" collectingDate="1990-06-07" collectionCode="BMNH" collectorName="G. Minet. &amp; Bourgoin Coll." country="Laos" location="Chiang Mai" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai">
<collectingRegion id="49A014FEFFDFFC412B1CF83AF7A7FBCC" box="[1290,1432,1046,1071]" country="Thailand" name="Chiang Mai" pageId="11" pageNumber="522">Chiang Mai</collectingRegion>
, 
<collectingDate id="EF9E0534FFDFFC412E81F817F334FBB0" box="[151,267,1083,1107]" pageId="11" pageNumber="522" value="1990-06-07">
<date id="FFDAFCDCFFDFFC412E81F817F334FBB0" box="[151,267,1083,1107]" pageId="11" pageNumber="522" value="1990-06-07">7.VI.1990</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412F01F817F3BDFBB0" box="[279,386,1082,1107]" pageId="11" pageNumber="522">G. Minet.</collectorName>
<emphasis id="B910060EFFDFFC412F9FF816F3E0FBB0" bold="true" box="[393,479,1082,1107]" pageId="11" pageNumber="522">
<collectingCountry id="F3739A8CFFDFFC412F9FF816F3E6FBB0" box="[393,473,1082,1107]" name="Laos" pageId="11" pageNumber="522">LAOS</collectingCountry>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412FF1F817F035FBB0" box="[487,522,1083,1107]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412C0FF817F04DFBB0" box="[537,626,1083,1107]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412C68F816F115FBB0" box="[638,810,1082,1107]" pageId="11" pageNumber="522">Bourgoin Coll.</collectorName>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412D29F816F7A9FBB0" ID-GBIF-Occurrence="1701220387" box="[831,1430,1082,1107]" collectingDate="1915-05-18" collectorName="R. Vitalis" country="Laos" location="Xieng Khouang" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Xieng Khouang">
<collectingRegion id="49A014FEFFDFFC412D29F816F1C9FBB0" box="[831,1014,1082,1107]" country="Laos" name="Xiangkhouang" pageId="11" pageNumber="522">Xieng Khouang</collectingRegion>
, 
<collectingDate id="EF9E0534FFDFFC412A17F817F6A6FBB0" box="[1025,1177,1082,1107]" pageId="11" pageNumber="522" value="1915-05-18">
<date id="FFDAFCDCFFDFFC412A17F817F6A6FBB0" box="[1025,1177,1082,1107]" pageId="11" pageNumber="522" value="1915-05-18">18 May 1915</date>
</collectingDate>
// 
<collectingCountry id="F3739A8CFFDFFC412AAFF817F6CFFBB1" box="[1209,1264,1083,1106]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
<collectorName id="2691BFCAFFDFFC412AE1F817F759FBB0" box="[1271,1382,1082,1107]" pageId="11" pageNumber="522">R. Vitalis</collectorName>
; 
<specimenCount id="9D621195FFDFFC412B64F817F7A9FBB0" box="[1394,1430,1083,1107]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
</materialsCitation>
, 2♀♀ (BMNH, Bourgoin Coll.), Muong Houoc, 
<date id="FFDAFCDCFFDFFC412CAEF872F112FB95" box="[696,813,1118,1142]" pageId="11" pageNumber="522" value="1920-05-10">10.V.1920</date>
; 1♂ (BMNH, Bourgoin Coll.), Pou Bia, 
<date id="FFDAFCDCFFDFFC412AE8F872F753FB95" box="[1278,1388,1118,1142]" pageId="11" pageNumber="522" value="1919-01-23">23.I.1919</date>
; 1♂ (BMNH, Bourgoin Coll.), Muong Pak, 
<date id="FFDAFCDCFFDFFC412C66F8AEF0D1FB79" box="[624,750,1154,1178]" pageId="11" pageNumber="522" value="1918-12-04">4.XII.1918</date>
; 1♂ (BMNH, Bourgoin Coll.), Pou Mi, 
<date id="FFDAFCDCFFDFFC412ACDF8AEF757FB79" box="[1243,1384,1154,1178]" pageId="11" pageNumber="522" value="1918-12-27">27.XII.1918</date>
; 1♂ (BMNH, Bourgoin Coll.), Nam Tha, 
<date id="FFDAFCDCFFDFFC412C21F889F08AFB5E" box="[567,693,1189,1213]" pageId="11" pageNumber="522" value="1918-04-10">10.IV.1918</date>
; 
<materialsCitation id="3B0CD041FFDFFC412CD7F889F702FB5D" ID-GBIF-Occurrence="1701220400" box="[705,1341,1189,1214]" collectingDate="1960-05" collectionCode="MNHN" collectorName="Baudon" country="Laos" location="Xieng Khouang" pageId="11" pageNumber="522" specimenCount="2" specimenCount-male="2" stateProvince="Xieng Khouang">
<specimenCount id="9D621195FFDFFC412CD7F889F0C6FB5E" box="[705,761,1189,1213]" pageId="11" pageNumber="522" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412D1EF889F15CFB5E" box="[776,867,1189,1213]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="522">MNHN</collectionCode>
), 
<collectingRegion id="49A014FEFFDFFC412D60F889F614FB5D" box="[886,1067,1189,1214]" country="Laos" name="Xiangkhouang" pageId="11" pageNumber="522">Xieng Khouang</collectingRegion>
, 
<collectingDate id="EF9E0534FFDFFC412A20F889F6B7FB5D" box="[1078,1160,1189,1214]" pageId="11" pageNumber="522" value="1960-05">
<date id="FFDAFCDCFFDFFC412A20F889F6B7FB5D" box="[1078,1160,1189,1214]" pageId="11" pageNumber="522" value="1960-05">V.1960</date>
</collectingDate>
, 
<collectingCountry id="F3739A8CFFDFFC412A85F88AF6F6FB5E" box="[1171,1225,1190,1213]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
(
<collectorName id="2691BFCAFFDFFC412ACEF889F70AFB5D" box="[1240,1333,1189,1214]" pageId="11" pageNumber="522">Baudon</collectorName>
)
</materialsCitation>
; 1♂, 1♀ (MNHN), Ban Van Eua, 
<materialsCitation id="3B0CD041FFDFFC412FAEF8E5F1E2FB02" ID-GBIF-Occurrence="1701220415" box="[440,989,1224,1249]" collectingDate="1967-04-30" collectionCode="BMNH" collectorName="Bourgoin Coll." country="Laos" location="Laos" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F3739A8CFFDFFC412FAEF8E5F3CEFB03" box="[440,497,1225,1248]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
, 
<collectingDate id="EF9E0534FFDFFC412FE9F8E4F040FB02" box="[511,639,1224,1249]" pageId="11" pageNumber="522" value="1967-04-30">
<date id="FFDAFCDCFFDFFC412FE9F8E4F040FB02" box="[511,639,1224,1249]" pageId="11" pageNumber="522" value="1967-04-30">30.IV.1967</date>
</collectingDate>
; 
<specimenCount id="9D621195FFDFFC412C9AF8E5F090FB02" box="[652,687,1225,1249]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412CD7F8E5F125FB02" box="[705,794,1225,1249]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412D31F8E4F1EBFB02" box="[807,980,1224,1249]" pageId="11" pageNumber="522">Bourgoin Coll.</collectorName>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412DFDF8E5F0D2FAE7" ID-GBIF-Occurrence="1701220386" collectingDate="1918-04-28" collectionCode="IRSNB" collectorName="Nam Long" country="Laos" determinerName="A. Bourgoin" location="Laos" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D621195FFDFFC412DFDF8E5F631FB02" box="[1003,1038,1225,1249]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412A36F8E5F647FB02" box="[1056,1144,1225,1249]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="11" pageNumber="522">IRSNB</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412A9BF8E5F734FB02" box="[1165,1291,1225,1249]" pageId="11" pageNumber="522">Nam Long</collectorName>
, 
<collectingDate id="EF9E0534FFDFFC412B0FF8E5F7A8FB02" box="[1305,1431,1225,1249]" pageId="11" pageNumber="522" value="1918-04-28">
<date id="FFDAFCDCFFDFFC412B0FF8E5F7A8FB02" box="[1305,1431,1225,1249]" pageId="11" pageNumber="522" value="1918-04-28">28.IV.1918</date>
</collectingDate>
, 
<collectingCountry id="F3739A8CFFDFFC412E81F8C0F2F2FAE0" box="[151,205,1260,1283]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
// 
<taxonomicName id="4C64A19FFFDFFC412EFAF8C1F3D9FAE7" box="[236,486,1259,1284]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDFFC412EFAF8C1F3D9FAE7" box="[236,486,1259,1284]" italics="true" pageId="11" pageNumber="522">Macronota penicillata</emphasis>
</taxonomicName>
det. 
<determinerName id="7E0736D8FFDFFC412C08F8C0F095FAE7" box="[542,682,1259,1284]" pageId="11" pageNumber="522">A. Bourgoin</determinerName>
1925
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412CEFF8C0F699FAE0" ID-GBIF-Occurrence="1701220401" box="[761,1190,1259,1284]" collectionCode="IRSNB" collectorName="Pakson" country="Laos" elevation="1" location="Laos" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D621195FFDFFC412CEFF8C0F123FAE7" box="[761,796,1260,1284]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412D3DF8C0F1BCFAE7" box="[811,899,1260,1284]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="11" pageNumber="522">IRSNB</collectionCode>
), 
<collectorName id="2691BFCAFFDFFC412D81F8C7F1D7FAE7" box="[919,1000,1259,1284]" pageId="11" pageNumber="522">Pakson</collectorName>
(
<collectingCountry id="F3739A8CFFDFFC412DE1F8C0F60DFAE7" box="[1015,1074,1260,1284]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
), 
<elevation id="00493D2FFFDFFC412A50F8C0F699FAE0" box="[1094,1190,1260,1284]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="522" unit="m" value="1.1">
<quantity id="4C9C77F9FFDFFC412A50F8C0F699FAE0" box="[1094,1190,1260,1284]" metricMagnitude="0" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="522" unit="m" value="1.1">1,100 m</quantity>
</elevation>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412AA4F8C0F0F3FACB" ID-GBIF-Occurrence="1701220398" collectingDate="2001-04-26" collectingDateMax="2001-05-11" collectingDateMin="2001-04-26" collectionCode="BMNH" collectorName="Krajcik Coll. &amp; Ban Saluei" country="Laos" location="Hua Phan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Hua Phan">
<specimenCount id="9D621195FFDFFC412AA4F8C0F6EAFAE7" box="[1202,1237,1260,1284]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412AF2F8C0F702FAE7" box="[1252,1341,1260,1284]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412B5EF8C7F2EEFACB" pageId="11" pageNumber="522">Krajčík Coll.</collectorName>
), 
<collectingDate id="EF9E0534FFDFFC412EFAF923F391FACB" box="[236,430,1295,1320]" pageId="11" pageNumber="522" value="2001-04-26" valueMax="2001-05-11" valueMin="2001-04-26">
<date id="FFDAFCDCFFDFFC412EFAF923F391FACB" box="[236,430,1295,1320]" pageId="11" pageNumber="522" value="2001-04-26" valueMax="2001-05-11" valueMin="2001-04-26">26.IV–11.V.2001</date>
</collectingDate>
, 
<collectingRegion id="49A014FEFFDFFC412FD6F93CF008FACB" box="[448,567,1295,1320]" country="Laos" name="Houaphan" pageId="11" pageNumber="522">Hua Phan</collectingRegion>
, 
<collectorName id="2691BFCAFFDFFC412C5EF93CF0F3FACB" box="[584,716,1295,1320]" pageId="11" pageNumber="522">Ban Saluei</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412CCBF923F3BFFAA8" ID-GBIF-Occurrence="1701220403" collectionCode="QCCC" collectorName="D. Hauck" country="Laos" elevation="2" latitude="20.25" location="Phu Phan" longLatPrecision="1268" longitude="104.03333" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Phu Phan">
<collectingRegion id="49A014FEFFDFFC412CCBF923F171FACB" box="[733,846,1295,1320]" inferredBySuffix="true" pageId="11" pageNumber="522">Phu Phan</collectingRegion>
Mountain, 
<geoCoordinate id="EE50BCDBFFDFFC412DCAF923F602FACB" box="[988,1085,1295,1320]" direction="north" orientation="latitude" pageId="11" pageNumber="522" precision="925" value="20.25">20°15′N</geoCoordinate>
, 
<geoCoordinate id="EE50BCDBFFDFFC412A58F923F685FACB" box="[1102,1210,1295,1320]" direction="east" orientation="longitude" pageId="11" pageNumber="522" precision="925" value="104.03333">104°02′E</geoCoordinate>
, 
<elevation id="00493D2FFFDFFC412ADDF923F750FACB" box="[1227,1391,1295,1320]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="522" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">
<quantity id="4C9C77F9FFDFFC412ADDF923F750FACB" box="[1227,1391,1295,1320]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="522" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1,500–2,000m</quantity>
</elevation>
, 
<collectorName id="2691BFCAFFDFFC412B97F93CF2DBFAA8" pageId="11" pageNumber="522">D. Hauck</collectorName>
; 
<specimenCount id="9D621195FFDFFC412EE4F91FF32BFAA8" box="[242,276,1331,1355]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412F30F91FF347FAA8" box="[294,376,1331,1355]" pageId="11" pageNumber="522">QCCC</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412F98F91FF0C4FAA9" ID-GBIF-Occurrence="1701220412" box="[398,763,1330,1355]" collectingDate="2014-06" country="Laos" elevation="2" location="Mount Phan" pageId="11" pageNumber="522" specimenCount="1" stateProvince="Mount Phan">
<collectingDate id="EF9E0534FFDFFC412F98F91FF3D2FAA8" box="[398,493,1331,1355]" pageId="11" pageNumber="522" value="2014-06">
<date id="FFDAFCDCFFDFFC412F98F91FF3D2FAA8" box="[398,493,1331,1355]" pageId="11" pageNumber="522" value="2014-06">VI.2014</date>
</collectingDate>
, 
<collectingRegion id="49A014FEFFDFFC412FEAF91FF0B4FAA8" box="[508,651,1330,1355]" inferredBySuffix="true" pageId="11" pageNumber="522">Mount Phan</collectingRegion>
, 
<elevation id="00493D2FFFDFFC412C8FF91EF0C4FAA9" box="[665,763,1330,1355]" metricMagnitude="0" metricUnit="m" metricValue="2.06" pageId="11" pageNumber="522" unit="m" value="2.06">
<quantity id="4C9C77F9FFDFFC412C8FF91EF0C4FAA9" box="[665,763,1330,1355]" metricMagnitude="0" metricUnit="m" metricValue="2.06" pageId="11" pageNumber="522" unit="m" value="2.06">2,060 m</quantity>
</elevation>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412D1FF91FF753FAA8" ID-GBIF-Occurrence="1701220405" box="[777,1388,1330,1355]" collectingDate="VII. 2011" collectionCode="QCCC" country="Laos" location="Mount Phu Pane" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Hua Phan">
<specimenCount id="9D621195FFDFFC412D1FF91FF113FAA8" box="[777,812,1331,1355]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412D28F91FF1AFFAA8" box="[830,912,1331,1355]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412DB0F91FF627FAA8" box="[934,1048,1331,1355]" pageId="11" pageNumber="522">
<date id="FFDAFCDCFFDFFC412DB0F91FF627FAA8" box="[934,1048,1331,1355]" pageId="11" pageNumber="522">VII.2011</date>
</collectingDate>
, 
<location id="8EBB8CC7FFDFFC412A30F91FF6D6FAA9" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412A30F91FF6D6FAA9" box="[1062,1257,1330,1355]" country="Laos" name="Mount Phu Pane" pageId="11" pageNumber="522" stateProvince="Hua Phan">Mount Phu Pane</location>
, 
<collectingRegion id="49A014FEFFDFFC412AE1F91FF753FAA8" box="[1271,1388,1330,1355]" country="Laos" name="Houaphan" pageId="11" pageNumber="522">Hua Phan</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412B6FF91FF3C8FA8C" ID-GBIF-Occurrence="1701220410" collectingDate="2014-04" collectionCode="QCCC" country="Laos" location="Mount Phan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Mount Phan">
<specimenCount id="9D621195FFDFFC412B6FF91FF7A3FAA8" box="[1401,1436,1331,1355]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412E89F97BF2CEFA8C" box="[159,241,1367,1391]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412F12F97BF360FA8C" box="[260,351,1367,1391]" pageId="11" pageNumber="522" value="2014-04">
<date id="FFDAFCDCFFDFFC412F12F97BF360FA8C" box="[260,351,1367,1391]" pageId="11" pageNumber="522" value="2014-04">IV.2014</date>
</collectingDate>
, 
<collectingRegion id="49A014FEFFDFFC412F7DF97BF3C8FA8C" box="[363,503,1366,1391]" inferredBySuffix="true" pageId="11" pageNumber="522">Mount Phan</collectingRegion>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412C14F97BF133FA8C" ID-GBIF-Occurrence="1701220396" box="[514,780,1366,1391]" country="Laos" elevation="2" location="Hua Phan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Hua Phan">
<collectingRegion id="49A014FEFFDFFC412C14F97BF04CFA8C" box="[514,627,1366,1391]" country="Laos" name="Houaphan" pageId="11" pageNumber="522">Hua Phan</collectingRegion>
, 
<elevation id="00493D2FFFDFFC412C68F97AF0E2FA8D" box="[638,733,1366,1391]" metricMagnitude="0" metricUnit="m" metricValue="2.026" pageId="11" pageNumber="522" unit="m" value="2.026">
<quantity id="4C9C77F9FFDFFC412C68F97AF0E2FA8D" box="[638,733,1366,1391]" metricMagnitude="0" metricUnit="m" metricValue="2.026" pageId="11" pageNumber="522" unit="m" value="2.026">2,026 m</quantity>
</elevation>
; 
<specimenCount id="9D621195FFDFFC412CFEF97BF133FA8C" box="[744,780,1367,1391]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
</materialsCitation>
, 1♀ (QCCC), 
<date id="FFDAFCDCFFDFFC412DA6F97AF61AFA8C" box="[944,1061,1366,1391]" pageId="11" pageNumber="522" value="2008-05-15">15.V.2008</date>
, Phou Pan, 
<materialsCitation id="3B0CD041FFDFFC412ABFF97BF767FA8C" ID-GBIF-Occurrence="1701220424" box="[1193,1368,1366,1391]" country="Laos" location="Houa Phan" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Houa Phan">
<collectingRegion id="49A014FEFFDFFC412ABFF97BF717FA8C" box="[1193,1320,1366,1391]" inferredBySuffix="true" pageId="11" pageNumber="522">Houa Phan</collectingRegion>
; 
<specimenCount id="9D621195FFDFFC412B25F97BF767FA8C" box="[1331,1368,1367,1391]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412B72F97BF0B8FA72" ID-GBIF-Occurrence="1701220423" collectingDate="2013-05" collectionCode="QCCC" collectorName="Sam Neua" country="Laos" location="Mount Pan" pageId="11" pageNumber="522" specimenCount="3" specimenCount-female="3" stateProvince="Houa Phan">
<specimenCount id="9D621195FFDFFC412B72F97BF7A3FA8C" box="[1380,1436,1367,1391]" pageId="11" pageNumber="522" type="female">3♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412E89F956F2CEFA71" box="[159,241,1402,1426]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412F1DF956F362FA71" box="[267,349,1402,1426]" pageId="11" pageNumber="522" value="2013-05">
<date id="FFDAFCDCFFDFFC412F1DF956F362FA71" box="[267,349,1402,1426]" pageId="11" pageNumber="522" value="2013-05">V.2013</date>
</collectingDate>
, 
<location id="8EBB8CC7FFDFFC412F66F956F3CAFA72" LSID="urn:lsid:plazi:treatment:03CD6B0AFFDEFC472E81FDF0F716FF36:8EBB8CC7FFDFFC412F66F956F3CAFA72" box="[368,501,1402,1425]" country="Laos" name="Mount Pan" pageId="11" pageNumber="522" stateProvince="Houa Phan">Mount Pan</location>
, 
<collectorName id="2691BFCAFFDFFC412C1EF956F0B8FA72" box="[520,647,1402,1426]" pageId="11" pageNumber="522">Sam Neua</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B0CD041FFDFFC412C8EF955F14EFA71" ID-GBIF-Occurrence="1701220420" box="[664,881,1401,1426]" country="Vietnam" location="VIETNAM" pageId="11" pageNumber="522" specimenCount="3" specimenCount-male="3">
<emphasis id="B910060EFFDFFC412C8EF955F117FA71" bold="true" box="[664,808,1401,1426]" pageId="11" pageNumber="522">
<collectingCountry id="F3739A8CFFDFFC412C8EF955F11BFA71" box="[664,804,1401,1426]" name="Vietnam" pageId="11" pageNumber="522">VIETNAM</collectingCountry>
:
</emphasis>
<specimenCount id="9D621195FFDFFC412D20F956F14EFA71" box="[822,881,1402,1426]" pageId="11" pageNumber="522" type="male">3♂♂</specimenCount>
</materialsCitation>
, 
<materialsCitation id="3B0CD041FFDFFC412D92F956F2C1FA55" ID-GBIF-Occurrence="1701220411" collectingDate="1992-05-16" collectingDateMax="1992-05-22" collectingDateMin="1992-05-16" collectionCode="QCCC" collectorName="Sapa" country="Vietnam" location="Lao Cai Province" pageId="11" pageNumber="522" specimenCount="2" specimenCount-female="2" stateProvince="Lao Cai Province">
<specimenCount id="9D621195FFDFFC412D92F956F183FA71" box="[900,956,1402,1426]" pageId="11" pageNumber="522" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DC4F956F61BFA71" box="[978,1060,1402,1426]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412A28F955F6E2FA71" box="[1086,1245,1401,1426]" pageId="11" pageNumber="522" value="1992-05-16" valueMax="1992-05-22" valueMin="1992-05-16">
<date id="FFDAFCDCFFDFFC412A28F955F6E2FA71" box="[1086,1245,1401,1426]" pageId="11" pageNumber="522" value="1992-05-16" valueMax="1992-05-22" valueMin="1992-05-16">16–22.V.1992</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412AF9F956F717FA71" box="[1263,1320,1402,1426]" pageId="11" pageNumber="522">Sapa</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412B2DF956F2C1FA55" country="Vietnam" name="Lao Cai" pageId="11" pageNumber="522">
Lao 
<collectingRegion id="49A014FEFFDFFC412B63F955F2C1FA55" inferredBySuffix="true" pageId="11" pageNumber="522">Cai Province</collectingRegion>
</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412F1AF9B2F193FA55" ID-GBIF-Occurrence="1701220422" box="[268,940,1437,1462]" collectingDate="2015-06-01" collectionCode="QCCC" collectorName="Kon &amp; Kinh &amp; Van Dang" country="Vietnam" location="Gia Lai" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai">
<specimenCount id="9D621195FFDFFC412F1AF9B2F311FA55" box="[268,302,1438,1462]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412F56F9B2F3ADFA55" box="[320,402,1438,1462]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412FB1F9B1F024FA55" box="[423,539,1437,1462]" pageId="11" pageNumber="522" value="2015-06-01">
<date id="FFDAFCDCFFDFFC412FB1F9B1F024FA55" box="[423,539,1437,1462]" pageId="11" pageNumber="522" value="2015-06-01">1.VI.2015</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412C3FF9B2F066FA56" box="[553,601,1438,1461]" pageId="11" pageNumber="522">Kon</collectorName>
ka 
<collectorName id="2691BFCAFFDFFC412C91F9B1F0FDFA55" box="[647,706,1437,1462]" pageId="11" pageNumber="522">Kinh</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412CD9F9B1F116FA55" box="[719,809,1437,1462]" country="Vietnam" name="Gia Lai" pageId="11" pageNumber="522">Gia Lai</collectingRegion>
, 
<collectorName id="2691BFCAFFDFFC412D21F9B2F193FA55" box="[823,940,1438,1462]" pageId="11" pageNumber="522">Van Dang</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412DACF9B2F364FA3A" ID-GBIF-Occurrence="1701220408" collectingDate="2015-07-01" collectionCode="QCCC" collectorName="Ngoc Linh &amp; Van Dang" country="Vietnam" location="Kon Tum" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Kon Tum">
<specimenCount id="9D621195FFDFFC412DACF9B2F1E3FA55" box="[954,988,1438,1462]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412DF8F9B2F67FFA55" box="[1006,1088,1438,1462]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412A43F9B1F6ECFA55" box="[1109,1235,1437,1462]" pageId="11" pageNumber="522" value="2015-07-01">
<date id="FFDAFCDCFFDFFC412A43F9B1F6ECFA55" box="[1109,1235,1437,1462]" pageId="11" pageNumber="522" value="2015-07-01">1.VII.2015</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412AF6F9B2F761FA55" box="[1248,1374,1437,1462]" pageId="11" pageNumber="522">Ngoc Linh</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412B7AF9B2F2F1FA3B" country="Vietnam" name="Kon Tum" pageId="11" pageNumber="522">Kon Tum</collectingRegion>
, 
<collectorName id="2691BFCAFFDFFC412EF7F9EDF364FA3A" box="[225,347,1473,1497]" pageId="11" pageNumber="522">Van Dang</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412F78F9EDF64AFA3B" ID-GBIF-Occurrence="1701220417" box="[366,1141,1472,1497]" collectingDate="2016-04" collectionCode="QCCC" collectorName="Mucangchai" country="Vietnam" elevation="2" location="Yen Bai Province" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Yen Bai Province">
<specimenCount id="9D621195FFDFFC412F78F9EDF3AEFA3A" box="[366,401,1473,1497]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412FBEF9EDF3C5FA3A" box="[424,506,1473,1497]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412C03F9ECF04EFA3A" box="[533,625,1472,1497]" pageId="11" pageNumber="522" value="2016-04">
<date id="FFDAFCDCFFDFFC412C03F9ECF04EFA3A" box="[533,625,1472,1497]" pageId="11" pageNumber="522" value="2016-04">IV.2016</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412C92F9ECF12EFA3A" box="[644,785,1472,1497]" pageId="11" pageNumber="522">Mucangchai</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412D32F9EDF1C3FA3A" box="[804,1020,1472,1497]" country="Vietnam" name="Yen Bai" pageId="11" pageNumber="522">
Yen 
<collectingRegion id="49A014FEFFDFFC412D49F9ECF1C3FA3A" box="[863,1020,1472,1497]" inferredBySuffix="true" pageId="11" pageNumber="522">Bai Province</collectingRegion>
</collectingRegion>
, 
<elevation id="00493D2FFFDFFC412A18F9EDF64AFA3B" box="[1038,1141,1473,1497]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="522" unit="m" value="1.7">
<quantity id="4C9C77F9FFDFFC412A18F9EDF64AFA3B" box="[1038,1141,1473,1497]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="522" unit="m" value="1.7">1,700 m</quantity>
</elevation>
</materialsCitation>
; 
<materialsCitation id="3B0CD041FFDFFC412A9FF9EDF054FA1F" ID-GBIF-Occurrence="1701220416" collectingDate="2016-07" collectionCode="QCCC" collectorName="Mucangchai" country="Vietnam" elevation="2" location="Yen Bai Province" pageId="11" pageNumber="522" specimenCount="1" specimenCount-female="1" stateProvince="Yen Bai Province">
<specimenCount id="9D621195FFDFFC412A9FF9EDF693FA3A" box="[1161,1196,1473,1497]" pageId="11" pageNumber="522" type="female">1♀</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412AD4F9EDF72BFA3A" box="[1218,1300,1473,1497]" pageId="11" pageNumber="522">QCCC</collectionCode>
), 
<collectingDate id="EF9E0534FFDFFC412B39F9ECF7A7FA3A" box="[1327,1432,1472,1497]" pageId="11" pageNumber="522" value="2016-07">
<date id="FFDAFCDCFFDFFC412B39F9ECF7A7FA3A" box="[1327,1432,1472,1497]" pageId="11" pageNumber="522" value="2016-07">VII.2016</date>
</collectingDate>
, 
<collectorName id="2691BFCAFFDFFC412E81F9C8F31AFA1E" box="[151,293,1508,1533]" pageId="11" pageNumber="522">Mucangchai</collectorName>
, 
<collectingRegion id="49A014FEFFDFFC412F27F9C9F3C2FA1E" box="[305,509,1508,1533]" country="Vietnam" name="Yen Bai" pageId="11" pageNumber="522">
Yen 
<collectingRegion id="49A014FEFFDFFC412F70F9C8F3C2FA1E" box="[358,509,1508,1533]" inferredBySuffix="true" pageId="11" pageNumber="522">Bai Province</collectingRegion>
</collectingRegion>
, 
<elevation id="00493D2FFFDFFC412C1CF9C8F054FA1F" box="[522,619,1508,1532]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="522" unit="m" value="1.7">
<quantity id="4C9C77F9FFDFFC412C1CF9C8F054FA1F" box="[522,619,1508,1532]" metricMagnitude="0" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="522" unit="m" value="1.7">1,700 m</quantity>
</elevation>
</materialsCitation>
; 1♂ (BMNH, Bourgoin Coll.), Chapa, 
<date id="FFDAFCDCFFDFFC412A3AF9C8F6B3FA1E" box="[1068,1164,1508,1533]" pageId="11" pageNumber="522" value="1916-06">VI.1916</date>
; 
<materialsCitation id="3B0CD041FFDFFC412A8CF9C8F393F9C3" ID-GBIF-Occurrence="1701220421" collectionCode="BMNH" collectorName="Bourgoin Coll. &amp; Lao-Kay." country="Vietnam" location="Tonkin" pageId="11" pageNumber="522" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D621195FFDFFC412A8CF9C8F683FA1F" box="[1178,1212,1508,1532]" pageId="11" pageNumber="522" type="male">1♂</specimenCount>
(
<collectionCode id="ED7542D9FFDFFC412ADBF9C8F719FA1F" box="[1229,1318,1508,1532]" pageId="11" pageNumber="522">BMNH</collectionCode>
, 
<collectorName id="2691BFCAFFDFFC412B25F9C8F2EEF9C3" pageId="11" pageNumber="522">Bourgoin Coll.</collectorName>
), 
<collectingCountry id="F3739A8CFFDFFC412EF3FA2BF308F9C3" box="[229,311,1543,1568]" name="Vietnam" pageId="11" pageNumber="522">Tonkin</collectingCountry>
, 
<collectorName id="2691BFCAFFDFFC412F54FA24F393F9C3" box="[322,428,1544,1568]" pageId="11" pageNumber="522">Lao-Kay.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E8997FFDFFC412ED1FA07F706F8FB" pageId="11" pageNumber="522" type="distribution">
<paragraph id="8BDBDA1CFFDFFC412ED1FA07F384F984" blockId="11.[151,1437,151,2029]" pageId="11" pageNumber="522">
<emphasis id="B910060EFFDFFC412ED1FA07F35FF9A7" bold="true" box="[199,352,1579,1604]" pageId="11" pageNumber="522">Distribution.</emphasis>
<collectingCountry id="F3739A8CFFDFFC412F7CFA07F38EF9A7" box="[362,433,1579,1604]" name="China" pageId="11" pageNumber="522">China</collectingCountry>
: 
<collectingRegion id="49A014FEFFDFFC412FABFA07F031F9A7" box="[445,526,1579,1604]" country="China" name="Xizang" pageId="11" pageNumber="522">Xizang</collectingRegion>
(
<emphasis id="B910060EFFDFFC412C37FA07F095F9A7" bold="true" box="[545,682,1579,1604]" pageId="11" pageNumber="522">new record</emphasis>
), 
<collectingRegion id="49A014FEFFDFFC412CD2FA00F122F9A0" box="[708,797,1580,1603]" country="China" name="Yunnan" pageId="11" pageNumber="522">Yunnan</collectingRegion>
; 
<collectingCountry id="F3739A8CFFDFFC412D3CFA07F1B3F9A7" box="[810,908,1579,1604]" name="Vietnam" pageId="11" pageNumber="522">Vietnam</collectingCountry>
; 
<collectingCountry id="F3739A8CFFDFFC412D8CFA00F1EBF9A0" box="[922,980,1580,1603]" name="Laos" pageId="11" pageNumber="522">Laos</collectingCountry>
; northern 
<collectingCountry id="F3739A8CFFDFFC412A5CFA07F68EF9A7" box="[1098,1201,1579,1604]" name="Thailand" pageId="11" pageNumber="522">Thailand</collectingCountry>
; northern 
<collectingCountry id="F3739A8CFFDFFC412B31FA00F7A8F9A7" box="[1319,1431,1580,1604]" name="Myanmar" pageId="11" pageNumber="522">Myanmar</collectingCountry>
; northeastern 
<collectingCountry id="F3739A8CFFDFFC412F3EFA62F35AF984" box="[296,357,1614,1639]" name="India" pageId="11" pageNumber="522">India</collectingCountry>
; 
<collectingCountry id="F3739A8CFFDFFC412F67FA62F388F984" box="[369,439,1614,1639]" name="Nepal" pageId="11" pageNumber="522">Nepal</collectingCountry>
.
</paragraph>
<paragraph id="8BDBDA1CFFDFFC412ED1FA5EF706F8FB" blockId="11.[151,1437,151,2029]" pageId="11" pageNumber="522">
<emphasis id="B910060EFFDFFC412ED1FA5EF3B9F968" bold="true" box="[199,390,1650,1675]" pageId="11" pageNumber="522">Natural history.</emphasis>
This species usually inhabit the mountainous area at elevations of 
<quantity id="4C9C77F9FFDFFC412A6CFA5EF718F969" box="[1146,1319,1650,1675]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="11" pageNumber="522" unit="m" value="1.75" valueMax="2.5" valueMin="1.0">1,000–2,500 m</quantity>
. The label data indicated that the adult activity from May to August. This species overwinter as mature larvae or newly emerged adults: some fresh adults were found in 
<date id="FFDAFCDCFFDFFC412CA9FA94F148F932" box="[703,887,1720,1745]" pageId="11" pageNumber="522" value="2014-09">September 2014</date>
(Mr. Zi-Chun Xiong, by artificial breeding), and many larvae with a fresh adult were captured in decayed wood in 
<date id="FFDAFCDCFFDFFC412D69FAF0F61CF916" box="[895,1059,1756,1781]" pageId="11" pageNumber="522" value="2016-02">February 2016</date>
within the cockroach galleries of 
<taxonomicName id="4C64A19FFFDFFC412E81FAD3F3ECF8FB" authority="Roth, 1979" authorityName="Roth" authorityYear="1979" box="[151,467,1791,1816]" class="Magnoliopsida" family="Adoxaceae" genus="Salgunea" kingdom="Plantae" order="Dipsacales" pageId="11" pageNumber="522" phylum="Tracheophyta" rank="species" species="raggei">
<emphasis id="B910060EFFDFFC412E81FAD3F370F8FB" box="[151,335,1791,1816]" italics="true" pageId="11" pageNumber="522">Salgunea raggei</emphasis>
Roth, 1979
</taxonomicName>
(
<taxonomicName id="4C64A19FFFDFFC412FF7FAD3F06DF8FB" authorityName="Brunner von Wattenwyl" authorityYear="1882" box="[481,594,1791,1816]" class="Insecta" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="522" phylum="Arthropoda" rank="order">Blattodea</taxonomicName>
: 
<taxonomicName id="4C64A19FFFDFFC412C4BFAD3F0E4F8FB" box="[605,731,1791,1816]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="522" phylum="Arthropoda" rank="family">Blaberidae</taxonomicName>
: 
<taxonomicName id="4C64A19FFFDFFC412CF3FAD3F14BF8FB" box="[741,884,1791,1816]" class="Insecta" family="Blaberidae" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="522" phylum="Arthropoda" rank="subFamily" subFamily="Panesthiinae">Panesthiinae</taxonomicName>
) (
<figureCitation id="135FC699FFDFFC412D9DFAD3F62BF8FB" box="[907,1044,1791,1816]" captionStart="FIGURES 95 – 99" captionStartId="19.[151,264,1958,1981]" captionTargetBox="[175,1411,193,1938]" captionTargetId="figure@19.[175,1411,193,1938]" captionTargetPageId="19" captionText="FIGURES 95 – 99. Habitat of Coilodera. 95 – 97, C. mearesii: 95, habitat (Zhangmu, Nyalam, Xizang); 96 – 97, host plant (Hydrangea sp.). 98 – 99, C. penicillata: larva in cockroach galleries of Salganea raggei in rotten wood (Yingjiang, Yunnan)." httpUri="https://zenodo.org/record/1059361/files/figure.png" pageId="11" pageNumber="522">Figs. 98–99</figureCitation>
), or 
<taxonomicName id="4C64A19FFFDFFC412A50FAD3F684F8FB" authorityName="Leach" authorityYear="1815" box="[1094,1211,1791,1816]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
(
<figureCitation id="135FC699FFDFFC412ADCFAD3F712F8FB" box="[1226,1325,1791,1816]" captionStart="FIGURES 100 – 106" captionStartId="20.[151,264,1875,1898]" captionTargetBox="[187,1399,193,1849]" captionTargetId="figure@20.[187,1399,193,1849]" captionTargetPageId="20" captionText="FIGURES 100 – 106. Habitat of Coilodera. 100 – 102, C. penicillata: 100 – 101, closer view of habitat and newly emerged adult in pupal cell in rotten wood (Tongbiguan Nature Reserve, Yingjiang, Yunnan); 102, newly emerged adult and larva inhabit in rotten wood with Blattodea and Passalidae (Mount Xiaolushan, Lancang, Yunnan). 103 – 106, Host plants of C. nigroscutellaris: 103 – 104. Hydrangea sp. (Mount Dayaoshan, Guangxi, photograph by Lu Qiu); 105 – 106. Ampelopsis sp. (Mount Wuzhishan, Hainan Island)." httpUri="https://zenodo.org/record/1059363/files/figure.png" pageId="11" pageNumber="522">Fig. 102</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E8997FFDFFC472ED1FB0FF716FF36" lastPageId="13" lastPageNumber="524" pageId="11" pageNumber="522" type="discussion">
<paragraph id="8BDBDA1CFFDFFC412ED1FB0FF6C4F80E" blockId="11.[151,1437,151,2029]" pageId="11" pageNumber="522">
<emphasis id="B910060EFFDFFC412ED1FB0FF304F8DF" bold="true" box="[199,315,1827,1852]" pageId="11" pageNumber="522">Remarks.</emphasis>
<taxonomicName id="4C64A19FFFDFFC412F55FB0FF00AF8DF" authorityName="Hope" authorityYear="1831" box="[323,565,1827,1852]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDFFC412F55FB0FF00AF8DF" box="[323,565,1827,1852]" italics="true" pageId="11" pageNumber="522">Coilodera penicillata</emphasis>
</taxonomicName>
is similar to both 
<taxonomicName id="4C64A19FFFDFFC412D13FB08F1E8F8DF" authorityName="Moser" authorityYear="1902" box="[773,983,1827,1852]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="nigroscutellaris">
<emphasis id="B910060EFFDFFC412D13FB08F1E8F8DF" box="[773,983,1827,1852]" italics="true" pageId="11" pageNumber="522">C. nigroscutellaris</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C64A19FFFDFFC412A06FB08F695F8DF" authorityName="Moser" authorityYear="1910" box="[1040,1194,1828,1852]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFDFFC412A06FB08F695F8DF" box="[1040,1194,1828,1852]" italics="true" pageId="11" pageNumber="522">C. formosana</emphasis>
</taxonomicName>
, but the scutellum of the latter two species are glabrous (scutellum usually tomentose in 
<taxonomicName id="4C64A19FFFDFFC412DBAFB6BF676F8BC" authorityName="Hope" authorityYear="1831" box="[940,1097,1862,1887]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDFFC412DBAFB6BF676F8BC" box="[940,1097,1862,1887]" italics="true" pageId="11" pageNumber="522">C. penicillata</emphasis>
</taxonomicName>
), and the distal maculae and sutural maculae on the elytra of 
<taxonomicName id="4C64A19FFFDFFC412C1AFB47F098F860" authorityName="Moser" authorityYear="1910" box="[524,679,1899,1923]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFDFFC412C1AFB47F098F860" box="[524,679,1899,1923]" italics="true" pageId="11" pageNumber="522">C. formosana</emphasis>
</taxonomicName>
always connected (those of the former two species unconnected). These diagnostic characters were adopted when 
<bibRefCitation id="EFF5A7EDFFDFFC412CF5FBA1F1B5F845" author="Miksic" box="[739,906,1933,1958]" pageId="11" pageNumber="522" refString="Miksic, R. (1972). Revision der Gattung Coelodera Burmeister (73. Beitrag zur Kenntnis der Scarabaeiden). Revue suisse de Zoologie, 79, 675 - 707." type="journal article" year="1972">Mikšić (1972)</bibRefCitation>
revised the genus, but the latter two were recognized as two subspecies of 
<taxonomicName id="4C64A19FFFDFFC412C15FB9EF0A4F829" authorityName="Hope" authorityYear="1831" box="[515,667,1969,1994]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="522" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFDFFC412C15FB9EF0A4F829" box="[515,667,1969,1994]" italics="true" pageId="11" pageNumber="522">C. penicillata</emphasis>
</taxonomicName>
, and this treatment was widely accepted (
<bibRefCitation id="EFF5A7EDFFDFFC412A7AFB9DF6C2F829" author="Miksic" box="[1132,1277,1969,1994]" pageId="11" pageNumber="522" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, Bosnia and Herzegovina, 444 pp." type="book" year="1976">Mikšić 1976</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFDFFC412B1FFB9DF2EDF80E" author="Miyake" pageId="11" pageNumber="522" refString="Miyake, Y., Nakamura, S. &amp; Kojima, K. (1991) The Scarabaeoidea of Taiwan preserved in Hiwa Museum for Natural History, with descriptions of two new species. Miscellaneous Reports of the Hiwa Museum for Natural History, 29, 1 - 41." type="journal article" year="1991">
Miyake 
<emphasis id="B910060EFFDFFC412B72FB9EF7A3F829" box="[1380,1436,1969,1994]" italics="true" pageId="11" pageNumber="522">et al.</emphasis>
1991
</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFDFFC412ECBFBF8F34CF80E" author="Krajcik" box="[221,371,2004,2029]" pageId="11" pageNumber="522" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, Czech Republic, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
, 
<bibRefCitation id="EFF5A7EDFFDFFC412F68FBF9F387F80E" author="Krajcik" box="[382,440,2005,2029]" pageId="11" pageNumber="522" refString="Krajcik, M. (2011) Illustrated catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x. Supplement, 1 - 113." type="book chapter" year="2011">2011</bibRefCitation>
, 
<bibRefCitation id="EFF5A7EDFFDFFC412FD2FBF9F3C0F80E" author="Krajcik" box="[452,511,2005,2029]" pageId="11" pageNumber="522" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">2012</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFDFFC412C1CFBF8F0CEF80E" author="Sakai" box="[522,753,2004,2029]" pageId="11" pageNumber="522" refString="Sakai, K. &amp; Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Volume 3. Mushi- Sha, Tokyo, Japan, 421 pp." type="book" year="1998">Sakai &amp; Nagai 1998</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFDFFC412CEBFBF9F1A2F80E" author="Yu" box="[765,925,2004,2029]" pageId="11" pageNumber="522" refString="Yu, C. K., Kobayashi, H. &amp; Chu, Y. I. (1998) The Scarabaeidae of Taiwan. Mu Sheng Co., Taipei, Taiwan, 263 pp." type="book" year="1998">
Yu 
<emphasis id="B910060EFFDFFC412D35FBFAF164F80E" box="[803,859,2004,2029]" italics="true" pageId="11" pageNumber="522">et al.</emphasis>
1998
</bibRefCitation>
; Smetana 2006; 
<bibRefCitation id="EFF5A7EDFFDFFC412A41FBF8F6D0F80E" author="Bezdek" box="[1111,1263,2004,2029]" pageId="11" pageNumber="522" refString="Bezdek, A. (2016) Subfamily Cetoniinae, pp. 367 - 412. In: Lobl, I. &amp; Lobl, D. (eds), Catalogue of Palaearctic Coleoptera, Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, The Netherlands." type="journal article" year="2016">Bezděk 2016</bibRefCitation>
).
</paragraph>
<caption id="DF1B8A94FFD8FC462E81F9DAF301F96B" httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="12" pageNumber="523" targetBox="[151,1436,198,1500]" targetPageId="12">
<paragraph id="8BDBDA1CFFD8FC462E81F9DAF301F96B" blockId="12.[151,1436,1526,1672]" pageId="12" pageNumber="523">
<emphasis id="B910060EFFD8FC462E81F9DAF369F9EF" bold="true" box="[151,342,1526,1549]" pageId="12" pageNumber="523">FIGURES 21–33.</emphasis>
Habitus of 
<taxonomicName id="4C64A19FFFD8FC462FC4F9DBF006F9EE" authorityName="Hope" authorityYear="1831" box="[466,569,1527,1549]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B910060EFFD8FC462FC4F9DBF006F9EE" box="[466,569,1527,1549]" italics="true" pageId="12" pageNumber="523">Coilodera</emphasis>
</taxonomicName>
. 21–24, 
<taxonomicName id="4C64A19FFFD8FC462C83F9D4F11FF9EE" authorityName="Hope" authorityYear="1831" box="[661,800,1527,1549]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFD8FC462C83F9D4F11FF9EE" box="[661,800,1527,1549]" italics="true" pageId="12" pageNumber="523">C. penicillata</emphasis>
</taxonomicName>
(male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25–28, 
<taxonomicName id="4C64A19FFFD8FC462C95FA3BF132F9CF" authorityName="Moser" authorityYear="1910" box="[643,781,1558,1580]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFD8FC462C95FA3BF132F9CF" box="[643,781,1558,1580]" italics="true" pageId="12" pageNumber="523">C. formosana</emphasis>
</taxonomicName>
: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29–33, 
<taxonomicName id="4C64A19FFFD8FC462C17FA19F081F9A9" authorityName="Moser" authorityYear="1902" box="[513,702,1588,1610]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="nigroscutellaris">
<emphasis id="B910060EFFD8FC462C17FA19F081F9A9" box="[513,702,1588,1610]" italics="true" pageId="12" pageNumber="523">C. nigroscutellaris</emphasis>
</taxonomicName>
: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male).
</paragraph>
</caption>
<paragraph id="8BDBDA1CFFD8FC462ED1FA83F76DF87C" blockId="12.[151,1436,1711,2024]" pageId="12" pageNumber="523">
The character of distal maculae connecting with sutural maculae appears in all three species discussed above (
<figureCitation id="135FC699FFD8FC462E89FAFFF3B8F90F" box="[159,391,1747,1772]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="12" pageNumber="523">Figs. 22, 25–26, 30</figureCitation>
), but the differences of the scutellum, the mesometasternal suture, and the parameres are evident and consistent. Consequently, we regarded them as three independent species: scutellum usually clad with arrow-shape yellow tomentum in 
<taxonomicName id="4C64A19FFFD8FC462C00FB30F08EF8D7" authorityName="Hope" authorityYear="1831" box="[534,689,1819,1844]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFD8FC462C00FB30F08EF8D7" box="[534,689,1819,1844]" italics="true" pageId="12" pageNumber="523">C. penicillata</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462CD7FB37F176F8D7" box="[705,841,1819,1844]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="12" pageNumber="523">Figs. 21–22</figureCitation>
) (except for a few individuals, see variation above), while glabrous in 
<taxonomicName id="4C64A19FFFD8FC462F71FB6CF005F8BB" authorityName="Moser" authorityYear="1902" box="[359,570,1855,1880]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="nigroscutellaris">
<emphasis id="B910060EFFD8FC462F71FB6CF005F8BB" box="[359,570,1855,1880]" italics="true" pageId="12" pageNumber="523">C. nigroscutellaris</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462C5AFB13F0EBF8B4" box="[588,724,1855,1880]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="12" pageNumber="523">Figs. 29–31</figureCitation>
) and 
<taxonomicName id="4C64A19FFFD8FC462D01FB6CF18DF8B4" authorityName="Moser" authorityYear="1910" box="[791,946,1855,1879]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFD8FC462D01FB6CF18DF8B4" box="[791,946,1855,1879]" italics="true" pageId="12" pageNumber="523">C. formosana</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462DD5FB13F670F8BB" box="[963,1103,1855,1880]" captionStart="FIGURES 21 – 33" captionStartId="12.[151,264,1526,1549]" captionTargetBox="[151,1436,198,1500]" captionTargetId="figure@12.[151,1436,193,1506]" captionTargetPageId="12" captionText="FIGURES 21 – 33. Habitus of Coilodera. 21 – 24, C. penicillata (male): 21 &amp; 24, Hua Phan, northeastern Laos; 22, Yunxian, Yunnan; 23, Mount Daweishan, Yunnan. 25 – 28, C. formosana: 25 &amp; 28, Nantou, Taiwan (male); 26, Taitung, Taiwan (female); 27, Taitung, Taiwan (male). 29 – 33, C. nigroscutellaris: 29, Mount Dayaoshan, Guangxi (male); 30, Mount Wuzhishan, Hainan Island (female); 31, Mount Wuyishan, Fujian (female); 32, Mount Shengtangshan, Guangxi (male); 33, Mount Maoershan, Guangxi (male)." httpUri="https://zenodo.org/record/1059341/files/figure.png" pageId="12" pageNumber="523">Figs. 25–26</figureCitation>
); mesometasternal suture not engraved in 
<taxonomicName id="4C64A19FFFD8FC462F35FB48F383F89F" authorityName="Hope" authorityYear="1831" box="[291,444,1891,1916]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFD8FC462F35FB48F383F89F" box="[291,444,1891,1916]" italics="true" pageId="12" pageNumber="523">C. penicillata</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462FDAFB4FF06BF89F" box="[460,596,1891,1916]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="12" pageNumber="523">Figs. 63–64</figureCitation>
) and 
<taxonomicName id="4C64A19FFFD8FC462C85FB48F15AF89F" authorityName="Moser" authorityYear="1902" box="[659,869,1891,1916]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="nigroscutellaris">
<emphasis id="B910060EFFD8FC462C85FB48F15AF89F" box="[659,869,1891,1916]" italics="true" pageId="12" pageNumber="523">C. nigroscutellaris</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462D63FB4FF1F6F89F" box="[885,969,1891,1916]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="12" pageNumber="523">Fig. 66</figureCitation>
), while deeply engraved in 
<taxonomicName id="4C64A19FFFD8FC462B15FB48F7A3F89F" authorityName="Moser" authorityYear="1910" box="[1283,1436,1892,1916]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFD8FC462B15FB48F7A3F89F" box="[1283,1436,1892,1916]" italics="true" pageId="12" pageNumber="523">C. formosana</emphasis>
</taxonomicName>
(
<figureCitation id="135FC699FFD8FC462E89FBABF2CCF843" box="[159,243,1927,1952]" captionStart="FIGURES 49 – 72" captionStartId="15.[151,264,1812,1835]" captionTargetBox="[151,1436,195,1791]" captionTargetId="figure@15.[151,1436,195,1791]" captionTargetPageId="15" captionText="FIGURES 49 – 72. Scutellum, mesometasternal process, and male metatibia of Coilodera. 49 – 58, scutellum: 49, C. dives. 50, C. grandimaculata. 51, C. mearesii. 52 – 56, C. penicillata: 52 – 53, Lincang, Yunnan; 54, Gia Lai, central Vietnam; 55 – 56, Mount Daweishan, Yunnan. 57, C. formosana. 58, C. nigroscutellaris. 59 – 66, mesometasternal process. 59, C. dives. 60 – 61, C. grandimaculata: 60, male holotype; 61, female paratype. 62, C. mearesii. 63 – 64, C. penicillata: 63. Hua Phan, northeastern Laos; 64, Jingdong, Yunnan. 65, C. formosana; 66, C. nigroscutellaris. 67 – 72, male metatibia. 67, C. dives. 68, C. grandimaculata. 69, C. mearesii. 70, C. penicillata. 71, C. formosana. 72, C. nigroscutellaris." httpUri="https://zenodo.org/record/1059349/files/figure.png" pageId="12" pageNumber="523">Fig. 65</figureCitation>
); parameres and basal piece distinctly different between the three species as showed in 
<figureCitation id="135FC699FFD8FC462AD0FBABF771F87C" box="[1222,1358,1927,1952]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="12" pageNumber="523">Figs. 79–84</figureCitation>
.
</paragraph>
<paragraph id="8BDBDA1CFFD8FC472ED1FB87F716FF36" blockId="12.[151,1436,1711,2024]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="524" pageId="12" pageNumber="523">
The distribution of 
<taxonomicName id="4C64A19FFFD8FC462FA7FB87F098F827" authorityName="Hope" authorityYear="1831" box="[433,679,1963,1988]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B910060EFFD8FC462FA7FB87F098F827" box="[433,679,1963,1988]" italics="true" pageId="12" pageNumber="523">Coilodera penicillata</emphasis>
</taxonomicName>
ranges from 
<collectingCountry id="F3739A8CFFD8FC462D5BFB87F1AFF827" box="[845,912,1963,1988]" name="Nepal" pageId="12" pageNumber="523">Nepal</collectingCountry>
and northeastern 
<collectingCountry id="F3739A8CFFD8FC462A7FFB87F69CF827" box="[1129,1187,1963,1988]" name="India" pageId="12" pageNumber="523">India</collectingCountry>
to northern 
<collectingCountry id="F3739A8CFFD8FC462B2BFB87F7A3F827" box="[1341,1436,1963,1988]" name="Vietnam" pageId="12" pageNumber="523">Vietnam</collectingCountry>
(
<bibRefCitation id="EFF5A7EDFFD8FC462E88FBE3F311F804" author="Arrow" box="[158,302,1999,2023]" pageId="12" pageNumber="523" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, United Kingdom, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC462F2DFBE3F3F3F804" author="Miksic" box="[315,460,1999,2024]" pageId="12" pageNumber="523" refString="Miksic, R. (1972). Revision der Gattung Coelodera Burmeister (73. Beitrag zur Kenntnis der Scarabaeiden). Revue suisse de Zoologie, 79, 675 - 707." type="journal article" year="1972">Mikšić 1972</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC462FCCFBE3F04CF804" author="Nakane" box="[474,627,1999,2024]" pageId="12" pageNumber="523" refString="Nakane, T. (1972) Coleoptera of Nepal collected by the Hokkaido University scientific expedition to Nepal Himalaya: Scarabaeidae I. Cetoniinae, Valginae and Dynastinae. Kontyu, 40, 112 - 118." type="journal article" year="1972">Nakane 1972</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC462C97FBE3F156F80B" author="Sakai" box="[641,873,1999,2024]" pageId="12" pageNumber="523" refString="Sakai, K. &amp; Ikeda, K. (1996) Check list of insects of Thailand 4. Family Scarabaeidae, subfamily Cetoniinae. Memoirs of the Faculty of Liberal Arts &amp; Education. Part II, Mathematics &amp; Natural Sciences, University of Yamanashi, 47, 34 - 43." type="journal article" year="1996">Sakai &amp; Ikeda 1996</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC462D61FBE3F631F804" author="Krajcik" box="[887,1038,1999,2024]" pageId="12" pageNumber="523" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, Czech Republic, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
, 
<bibRefCitation id="EFF5A7EDFFD8FC462A0DFBE3F668F804" author="Krajcik" box="[1051,1111,1999,2023]" pageId="12" pageNumber="523" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">2012</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC462A72FBE3F76FF804" author="Sakai" box="[1124,1360,1999,2024]" pageId="12" pageNumber="523" refString="Sakai, K. &amp; Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Volume 3. Mushi- Sha, Tokyo, Japan, 421 pp." type="book" year="1998">Sakai &amp; Nagai 1998</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD8FC472B48FBE3F2ECFF53" author="Sakai" lastPageId="13" lastPageNumber="524" pageId="12" pageNumber="523" refString="Sakai, K. (2008) Record of cetoniid beetles from the central and northern Laos (Coleoptera, Scarabaeidae). Saikaku-Tsushin, 16, 11 - 22." type="journal article" year="2008">Sakai 2008</bibRefCitation>
; 
<bibRefCitation id="EFF5A7EDFFD9FC472EC9FCBBF347FF53" author="Bezdek" box="[223,376,151,176]" pageId="13" pageNumber="524" refString="Bezdek, A. (2016) Subfamily Cetoniinae, pp. 367 - 412. In: Lobl, I. &amp; Lobl, D. (eds), Catalogue of Palaearctic Coleoptera, Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, The Netherlands." type="journal article" year="2016">Bezděk 2016</bibRefCitation>
), and the record from 
<collectingRegion id="49A014FEFFD9FC472C6FFCB4F0EDFF4C" box="[633,722,152,175]" country="China" name="Yunnan" pageId="13" pageNumber="524">Yunnan</collectingRegion>
, 
<collectingCountry id="F3739A8CFFD9FC472CC8FCBBF11EFF53" box="[734,801,151,176]" name="China" pageId="13" pageNumber="524">China</collectingCountry>
was reported by 
<bibRefCitation id="EFF5A7EDFFD9FC472DFFFCB4F659FF53" author="Ma" box="[1001,1126,152,176]" pageId="13" pageNumber="524" refString="Ma, W. Z. (1987) Coleoptera: Cetoniidae. In: Huang, F. S. (ed.), Forest Insects of Yunnan. Yunnan Science &amp; Technology Press, Kunming, China, pp. 570 - 580. [In Chinese]" type="book chapter" year="1987">Ma (1987)</bibRefCitation>
. 
<taxonomicName id="4C64A19FFFD9FC472A67FCBBF7A3FF53" authorityName="Moser" authorityYear="1902" box="[1137,1436,151,176]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="species" species="nigroscutellaris">
<emphasis id="B910060EFFD9FC472A67FCBBF7A3FF53" box="[1137,1436,151,176]" italics="true" pageId="13" pageNumber="524">Coilodera nigroscutellaris</emphasis>
</taxonomicName>
occurs in northern 
<collectingCountry id="F3739A8CFFD9FC472F7CFC90F3F6FF36" box="[362,457,188,213]" name="Vietnam" pageId="13" pageNumber="524">Vietnam</collectingCountry>
and southern 
<collectingCountry id="F3739A8CFFD9FC472C70FC90F096FF36" box="[614,681,188,213]" name="China" pageId="13" pageNumber="524">China</collectingCountry>
and 
<taxonomicName id="4C64A19FFFD9FC472CC9FC91F147FF37" authorityName="Moser" authorityYear="1910" box="[735,888,188,212]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B910060EFFD9FC472CC9FC91F147FF37" box="[735,888,188,212]" italics="true" pageId="13" pageNumber="524">C. formosana</emphasis>
</taxonomicName>
is endemic to 
<collectingCountry id="F3739A8CFFD9FC472A0DFC90F652FF36" box="[1051,1133,188,213]" name="Taiwan" pageId="13" pageNumber="524">Taiwan</collectingCountry>
Island (
<figureCitation id="135FC699FFD9FC472ADEFC90F724FF36" box="[1224,1307,188,213]" captionStart="FIGURES 73 – 85" captionStartId="16.[151,264,1779,1802]" captionTargetBox="[151,1436,193,1758]" captionTargetId="figure@16.[151,1436,193,1758]" captionTargetPageId="16" captionText="FIGURES 73 – 85. Male genitalia and distribution of Coilodera. 73 – 84, Male genitalia: 73 – 74, C. dives. 75 – 76, C. grandimaculata, holotype. 77 – 78, C. mearesii. 79 – 80, C. penicillata. 81 – 82, C. formosana. 83 – 84, C. nigroscutellaris. 85, distribution of Coilodera." httpUri="https://zenodo.org/record/1059353/files/figure.png" pageId="13" pageNumber="524">Fig. 85</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>