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<document ID-DOI="10.11646/zootaxa.5239.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7635225" ID-ZooBank="BBE62A0D-2587-4EAD-9241-84546DE5D618" approvalRequired="15" approvalRequired_for_taxonomicNames="3" approvalRequired_for_textStreams="12" checkinTime="1676283593745" checkinUser="plazi" docAuthor="Kalúz, Stanislav &amp; Ermilov, Sergey G." docDate="2023" docId="03A65F3BFFA7FFF67AB9FEE6BB06F931" docLanguage="en" docName="zootaxa.5239.4.4.pdf" docOrigin="Zootaxa 5239 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cunaxa asiatica Kalúz &amp; Ermilov 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="527" masterDocId="FF9F2743FFA5FFF07A2EFFADBF74FF8C" masterDocTitle="Two new species of Cunaxa (Acari, Prostigmata, Cunaxidae) from South-East Asia with a world key to the genus" masterLastPageNumber="536" masterPageNumber="521" pageNumber="523" updateTime="1676284219807" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Two new species of Cunaxa (Acari, Prostigmata, Cunaxidae) from South-East Asia with a world key to the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kalúz, Stanislav</mods:namePart>
<mods:affiliation>Institute of Zoology, Slovak Academy of Sciences, Dúbravskácesta 9, SK- 845 06</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ermilov, Sergey G.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0913-131X</mods:nameIdentifier>
<mods:affiliation>Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Semakova 10, 625003 Tyumen, Russia. ermilovacari @ yandex. ru; https: // orcid. org / 0000 - 0002 - 0913 - 131 X</mods:affiliation>
<mods:nameIdentifier type="email">ermilovacari@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5239</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
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<mods:start>521</mods:start>
<mods:end>536</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5239.4.4</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5239.4.4</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7635225</mods:identifier>
<mods:identifier type="ZooBank">BBE62A0D-2587-4EAD-9241-84546DE5D618</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7635268" ID-Zenodo-Dep="7635268" LSID="urn:lsid:plazi:treatment:03A65F3BFFA7FFF67AB9FEE6BB06F931" httpUri="http://treatment.plazi.org/id/03A65F3BFFA7FFF67AB9FEE6BB06F931" lastPageId="6" lastPageNumber="527" pageId="2" pageNumber="523">
<subSubSection box="[151,449,331,357]" pageId="2" pageNumber="523" type="nomenclature">
<paragraph blockId="2.[151,449,331,393]" box="[151,449,331,357]" pageId="2" pageNumber="523">
<heading bold="true" box="[151,449,331,357]" fontSize="11" level="1" pageId="2" pageNumber="523" reason="1">
<emphasis bold="true" box="[151,449,331,357]" pageId="2" pageNumber="523">
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[151,348,331,357]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis bold="true" box="[151,348,331,357]" italics="true" pageId="2" pageNumber="523">Cunaxa asiatica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[355,449,331,357]" pageId="2" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,274,367,393]" pageId="2" pageNumber="523" type="description">
<paragraph blockId="2.[151,449,331,393]" box="[151,274,367,393]" pageId="2" pageNumber="523">
(
<figureCitation box="[159,266,367,393]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1994,2018]" captionStartId-1="3.[151,250,1941,1965]" captionStartId-2="4.[151,250,1946,1970]" captionStartId-3="5.[151,250,1929,1953]" captionTargetBox-0="[154,1412,434,1953]" captionTargetBox-1="[168,1419,437,1896]" captionTargetBox-2="[170,1422,539,1915]" captionTargetBox-3="[166,1426,190,1895]" captionTargetId-0="figure-63@2.[150,1437,417,1971]" captionTargetId-1="figure-129@3.[151,1436,417,1917]" captionTargetId-2="figure-240@4.[151,1436,525,1923]" captionTargetId-3="figure-17@5.[151,1436,181,1905]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Cunaxa asiatica sp. nov., female (holotype), dorsal view of idiosoma." captionText-1="FIGURE 2. Cunaxa asiatica sp. nov., female (holotype), ventral view of idiosoma." captionText-2="FIGURE 3. Cunaxa asiatica sp. nov., female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view)." captionText-3="FIGURE 4. Cunaxa asiatica sp. nov., female (holotype), legs IIV (left side, dorsal view)." figureDoi-0="http://doi.org/10.5281/zenodo.7635227" figureDoi-1="http://doi.org/10.5281/zenodo.7635231" figureDoi-2="http://doi.org/10.5281/zenodo.7635233" figureDoi-3="http://doi.org/10.5281/zenodo.7635237" httpUri-0="https://zenodo.org/record/7635227/files/figure.png" httpUri-1="https://zenodo.org/record/7635231/files/figure.png" httpUri-2="https://zenodo.org/record/7635233/files/figure.png" httpUri-3="https://zenodo.org/record/7635237/files/figure.png" pageId="2" pageNumber="523">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635227" ID-Zenodo-Dep="7635227" box="[151,992,1994,2019]" httpUri="https://zenodo.org/record/7635227/files/figure.png" pageId="2" pageNumber="523" startId="2.[151,250,1994,2018]" targetBox="[154,1412,434,1953]" targetPageId="2">
<paragraph blockId="2.[151,992,1994,2019]" box="[151,992,1994,2019]" pageId="2" pageNumber="523">
<emphasis bold="true" box="[151,269,1994,2018]" pageId="2" pageNumber="523">FIGURE 1</emphasis>
.
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[281,447,1995,2018]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[281,447,1995,2018]" italics="true" pageId="2" pageNumber="523">Cunaxa asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[453,540,1995,2019]" pageId="2" pageNumber="523">
<taxonomicNameLabel box="[453,536,1995,2019]" pageId="2" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female (holotype), dorsal view of idiosoma.
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="524" type="diagnosis">
<paragraph blockId="3.[151,1436,150,393]" pageId="3" pageNumber="524">
<emphasis bold="true" box="[151,266,150,176]" pageId="3" pageNumber="524">Diagnosis</emphasis>
(female). Dorsal shields smooth. Proterosomal shield subtriangular, medial hysterosomal shield present, bearing setae
<emphasis box="[312,406,187,219]" italics="true" pageId="3" pageNumber="524">
c
<subScript attach="right" box="[325,333,205,219]" fontSize="6" pageId="3" pageNumber="524">2</subScript>
, c
<subScript attach="left" box="[363,371,205,219]" fontSize="6" pageId="3" pageNumber="524">1</subScript>
e
<subScript attach="right" box="[398,406,205,219]" fontSize="6" pageId="3" pageNumber="524">1</subScript>
</emphasis>
. Palpal telofemur with short and straight apophysis, and with two relatively long palpal tibiotarsal setae. Venter with 4 pairs of sts between coxae III and genital region, and with a pair of paragenital setae; 2 sts and 1 sts present on basifemora III and IV, respectively. Leg setal formulae: coxae IIV: 3-1-3-1 sts; basifemora 4-4-2-1 sts; telofemora 4-4-4-4 sts; proterosomal and hysterosomal shields present, medial hysterosomal shield with
<emphasis box="[151,241,331,363]" italics="true" pageId="3" pageNumber="524">
c
<subScript attach="right" box="[164,172,349,363]" fontSize="6" pageId="3" pageNumber="524">2</subScript>
, c
<subScript attach="left" box="[198,206,349,363]" fontSize="6" pageId="3" pageNumber="524">1</subScript>
e
<subScript attach="right" box="[233,241,349,363]" fontSize="6" pageId="3" pageNumber="524">1</subScript>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="527" pageId="3" pageNumber="524" type="description">
<paragraph blockId="3.[151,1436,150,393]" box="[199,1298,366,393]" pageId="3" pageNumber="524">
<emphasis bold="true" box="[199,345,366,392]" pageId="3" pageNumber="524">Description.</emphasis>
FEMALE (
<typeStatus box="[478,580,367,393]" pageId="3" pageNumber="524">holotype</typeStatus>
,
<specimenCount box="[592,715,367,393]" pageId="3" pageNumber="524" type="generic" typeStatus="paratype">1 paratype</specimenCount>
).
<emphasis box="[733,791,367,393]" italics="true" pageId="3" pageNumber="524">Body</emphasis>
length 354 (330377), width 249 (242256).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635231" ID-Zenodo-Dep="7635231" box="[151,1000,1941,1965]" httpUri="https://zenodo.org/record/7635231/files/figure.png" pageId="3" pageNumber="524" startId="3.[151,250,1941,1965]" targetBox="[168,1419,437,1896]" targetPageId="3">
<paragraph blockId="3.[151,1000,1941,1965]" box="[151,1000,1941,1965]" pageId="3" pageNumber="524">
<emphasis bold="true" box="[151,269,1941,1965]" pageId="3" pageNumber="524">FIGURE 2</emphasis>
.
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[281,447,1941,1965]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[281,447,1941,1965]" italics="true" pageId="3" pageNumber="524">Cunaxa asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[453,540,1941,1965]" pageId="3" pageNumber="524">
<taxonomicNameLabel box="[453,536,1941,1965]" pageId="3" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female (holotype), ventral view of idiosoma.
</paragraph>
</caption>
<paragraph blockId="4.[151,1437,151,507]" pageId="4" pageNumber="525">
<emphasis box="[199,289,151,176]" italics="true" pageId="4" pageNumber="525">Dorsum</emphasis>
(
<figureCitation box="[308,376,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1994,2018]" captionTargetBox="[154,1412,434,1953]" captionTargetId="figure-63@2.[150,1437,417,1971]" captionTargetPageId="2" captionText="FIGURE 1. Cunaxa asiatica sp. nov., female (holotype), dorsal view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.7635227" httpUri="https://zenodo.org/record/7635227/files/figure.png" pageId="4" pageNumber="525">Fig. 1</figureCitation>
). Propodosoma with subtriangular shield. Propodosomal shield 78 (7779) long, 204 (196 212) wide, bearing a pair of anterior (
<emphasis box="[582,604,187,212]" italics="true" pageId="4" pageNumber="525">at</emphasis>
) and posterior (
<emphasis box="[787,809,187,212]" italics="true" pageId="4" pageNumber="525">pt</emphasis>
) setose trichobothria and also two pairs of tactile setae (
<emphasis box="[161,252,223,249]" italics="true" pageId="4" pageNumber="525">mps, lps</emphasis>
). Anterior trichobothrium 196 (190202), posterior trichobothrium 253 (250256) long, distance between bases of
<emphasis box="[248,352,259,284]" italics="true" pageId="4" pageNumber="525">mpsmps</emphasis>
and
<emphasis box="[405,484,259,285]" italics="true" pageId="4" pageNumber="525">lpslps</emphasis>
47 (4351) and 119 (118120), respectively. Propodosomal and hysterosomal shields separated. Six pairs of tactile setae present on dorsal hysterosoma:
<emphasis box="[924,980,295,327]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[936,944,313,327]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
h
<subScript attach="left" box="[972,980,313,327]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
,
<emphasis box="[997,1017,295,327]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[1009,1017,313,327]" fontSize="6" pageId="4" pageNumber="525">2</subScript>
</emphasis>
. Proterosomal shield bears
<emphasis box="[1337,1392,295,327]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="right" box="[1350,1358,313,327]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
e
<subScript attach="left" box="[1384,1392,313,327]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
,
<emphasis box="[1409,1429,295,327]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[1421,1429,313,327]" fontSize="6" pageId="4" pageNumber="525">2</subScript>
</emphasis>
, setae
<emphasis box="[214,266,331,363]" italics="true" pageId="4" pageNumber="525">
f
<subScript attach="left" box="[221,229,349,363]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
h
<subScript attach="none" box="[258,266,349,363]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
are situated on small platelets. Length of setae
<emphasis box="[790,810,331,363]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[802,810,349,363]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
49 (4552),
<emphasis box="[951,971,331,363]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[963,971,349,363]" fontSize="6" pageId="4" pageNumber="525">2</subScript>
</emphasis>
23 (2125),
<emphasis box="[1114,1136,331,363]" italics="true" pageId="4" pageNumber="525">
d
<subScript attach="left" box="[1128,1136,349,363]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
22 (2222),
<emphasis box="[1279,1300,331,363]" italics="true" pageId="4" pageNumber="525">
e
<subScript attach="left" box="[1292,1300,349,363]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
25 (2525),
<emphasis box="[151,167,367,399]" italics="true" pageId="4" pageNumber="525">
f
<subScript attach="left" box="[159,167,385,399]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
40 (3841) and
<emphasis box="[358,380,367,399]" italics="true" pageId="4" pageNumber="525">
h
<subScript attach="left" box="[372,380,385,399]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
39 (3740). Distance between bases of setae:
<emphasis box="[914,969,367,399]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="left" box="[926,934,385,399]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
c
<subScript attach="left" box="[961,969,385,399]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
90 (8891) about two times longer than length of
<emphasis box="[259,280,403,435]" italics="true" pageId="4" pageNumber="525">
c
<subScript attach="right" box="[272,280,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
;
<emphasis box="[295,353,403,435]" italics="true" pageId="4" pageNumber="525">
d
<subScript attach="left" box="[309,317,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
d
<subScript attach="left" box="[345,353,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
80 (7881) about three times longer than length of
<emphasis box="[936,958,403,435]" italics="true" pageId="4" pageNumber="525">
d
<subScript attach="left" box="[950,958,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
;
<emphasis box="[973,1028,403,435]" italics="true" pageId="4" pageNumber="525">
e
<subScript attach="left" box="[985,993,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
e
<subScript attach="left" box="[1020,1028,421,435]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
71 (7071) about three times longer than length of
<emphasis box="[311,331,439,471]" italics="true" pageId="4" pageNumber="525">
e
<subScript attach="left" box="[323,331,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
;
<emphasis box="[346,397,439,471]" italics="true" pageId="4" pageNumber="525">
f
<subScript attach="left" box="[353,361,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
f
<subScript attach="left" box="[389,397,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
31 (3032), slightly shorter than length of
<emphasis box="[871,887,439,471]" italics="true" pageId="4" pageNumber="525">
f
<subScript attach="left" box="[879,887,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
;
<emphasis box="[901,959,439,471]" italics="true" pageId="4" pageNumber="525">
h
<subScript attach="left" box="[915,923,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
h
<subScript attach="left" box="[951,959,457,471]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
63 (6065) nearly twice longer than length of
<emphasis box="[182,204,475,507]" italics="true" pageId="4" pageNumber="525">
h
<subScript attach="left" box="[196,204,493,507]" fontSize="6" pageId="4" pageNumber="525">1</subScript>
</emphasis>
. Cupules
<emphasis box="[316,344,475,500]" italics="true" pageId="4" pageNumber="525">im</emphasis>
present.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635233" ID-Zenodo-Dep="7635233" httpUri="https://zenodo.org/record/7635233/files/figure.png" pageId="4" pageNumber="525" startId="4.[151,250,1946,1970]" targetBox="[170,1422,539,1915]" targetPageId="4">
<paragraph blockId="4.[151,1436,1946,2006]" pageId="4" pageNumber="525">
<emphasis bold="true" box="[151,268,1946,1970]" pageId="4" pageNumber="525">FIGURE 3</emphasis>
.
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[279,443,1947,1970]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[279,443,1947,1970]" italics="true" pageId="4" pageNumber="525">Cunaxa asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[448,527,1946,1970]" pageId="4" pageNumber="525">
<taxonomicNameLabel box="[448,527,1946,1970]" pageId="4" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7635237" ID-Zenodo-Dep="7635237" box="[151,1076,1929,1953]" httpUri="https://zenodo.org/record/7635237/files/figure.png" pageId="5" pageNumber="526" startId="5.[151,250,1929,1953]" targetBox="[166,1426,190,1895]" targetPageId="5">
<paragraph blockId="5.[151,1076,1929,1953]" box="[151,1076,1929,1953]" pageId="5" pageNumber="526">
<emphasis bold="true" box="[151,275,1929,1953]" pageId="5" pageNumber="526">FIGURE 4.</emphasis>
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[281,447,1930,1953]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="526" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[281,447,1930,1953]" italics="true" pageId="5" pageNumber="526">Cunaxa asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[453,540,1929,1953]" pageId="5" pageNumber="526">
<taxonomicNameLabel box="[453,536,1929,1953]" pageId="5" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female (holotype), legs IIV (left side, dorsal view).
</paragraph>
</caption>
<paragraph blockId="6.[151,1437,151,1731]" pageId="6" pageNumber="527">
<emphasis box="[199,271,151,176]" italics="true" pageId="6" pageNumber="527">Venter</emphasis>
(
<figureCitation box="[289,362,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1941,1965]" captionTargetBox="[168,1419,437,1896]" captionTargetId="figure-129@3.[151,1436,417,1917]" captionTargetPageId="3" captionText="FIGURE 2. Cunaxa asiatica sp. nov., female (holotype), ventral view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.7635231" httpUri="https://zenodo.org/record/7635231/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
). Coxal plates very weakly sclerotized, coxae III and IIIIV contiguous and finely striated. Setal formula of coxae IIV: 3-1-3-1sts. Venter of hysterosoma with a pair of simple centromedial setae 20 (2020) between coxae II and III on longitudinally striated integument, and with 3 pairs of hysterogastral setae 19 (1820) on transversally striated integument arranged anteriorly to genital platelets. A pair of aggenital setae 19 (1820) present close to anterior part of genital platelets. A pair of longitudinally striated genital platelets 48 (4650) long and 27 (2727) wide, each bearing four pairs of simple setae, setae increasing in length caudally:
<emphasis box="[1249,1271,331,363]" italics="true" pageId="6" pageNumber="527">
g
<subScript attach="left" box="[1263,1271,349,363]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
</emphasis>
16 (1516);
<emphasis box="[1414,1436,331,363]" italics="true" pageId="6" pageNumber="527">
g
<subScript attach="left" box="[1428,1436,349,363]" fontSize="6" pageId="6" pageNumber="527">2</subScript>
</emphasis>
16 (1616);
<emphasis box="[289,311,367,399]" italics="true" pageId="6" pageNumber="527">
g
<subScript attach="left" box="[303,311,385,399]" fontSize="6" pageId="6" pageNumber="527">3</subScript>
</emphasis>
17 (1717);
<emphasis box="[453,475,367,399]" italics="true" pageId="6" pageNumber="527">
g
<subScript attach="left" box="[467,475,385,399]" fontSize="6" pageId="6" pageNumber="527">4</subScript>
</emphasis>
17 (1717), and a pair of adjacent
<emphasis box="[862,895,367,399]" italics="true" pageId="6" pageNumber="527">
ps
<subScript attach="left" box="[887,895,385,399]" fontSize="6" pageId="6" pageNumber="527">2</subScript>
</emphasis>
setae 14 (1315), close to anal platelets. Genital papillae not visible.
</paragraph>
<paragraph blockId="6.[151,1437,151,1731]" pageId="6" pageNumber="527">
<taxonomicName authorityName="Krefft" authorityYear="1871" box="[199,342,439,465]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="527" phylum="Nematoda" rank="genus">
<emphasis box="[199,342,439,465]" italics="true" pageId="6" pageNumber="527">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation box="[359,484,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1946,1970]" captionTargetBox="[170,1422,539,1915]" captionTargetId="figure-240@4.[151,1436,525,1923]" captionTargetPageId="4" captionText="FIGURE 3. Cunaxa asiatica sp. nov., female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7635233" httpUri="https://zenodo.org/record/7635233/files/figure.png" pageId="6" pageNumber="527">Fig. 3AC</figureCitation>
). Five-segmented palp 203 (195210) long (including palpal trochanter) with palpal tibiotarsus slightly apically curved. (
<emphasis box="[565,618,475,501]" italics="true" pageId="6" pageNumber="527">Palp</emphasis>
(
<figureCitation box="[635,758,475,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1946,1970]" captionTargetBox="[170,1422,539,1915]" captionTargetId="figure-240@4.[151,1436,525,1923]" captionTargetPageId="4" captionText="FIGURE 3. Cunaxa asiatica sp. nov., female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7635233" httpUri="https://zenodo.org/record/7635233/files/figure.png" pageId="6" pageNumber="527">Fig. 3A, D</figureCitation>
) relatively slender; palpal chaetotaxy as follows: trochanter without setae; basifemur 47 (4747) long with 1 dorsomedial simple seta 10 (1011) long; inner surface of telofemur 32 (3232) with 1 lateromedial apophysis 12 (1212) and dorsodistal seta 11 (1112); inner ventral surface of genu 40 (3841) long with 1 laterodistal spine-like seta 11 (1112), dorsally with 1 laterodistal seta 11 (1111), and ventrally with mediodistal seta 19 (1819); tibiotarsus 69 (6871) inner surface with 1 long simple proximal seta 32 (3033) and medially with 1 stout spine-like seta 20 (1921), mediodorsal simple seta 17 (1618)], outer surface with 1 dorsodistal simple seta 8 (78), dorsodistal solenidion 8 (78) and terminating with small short claw.
<emphasis box="[151,261,727,753]" italics="true" pageId="6" pageNumber="527">Chelicera</emphasis>
(
<figureCitation box="[277,366,727,753]" captionStart="FIGURE 3" captionStartId="4.[151,250,1946,1970]" captionTargetBox="[170,1422,539,1915]" captionTargetId="figure-240@4.[151,1436,525,1923]" captionTargetPageId="4" captionText="FIGURE 3. Cunaxa asiatica sp. nov., female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7635233" httpUri="https://zenodo.org/record/7635233/files/figure.png" pageId="6" pageNumber="527">Fig. 3C</figureCitation>
). Slender, 128 (126132) long and 18 (1718) wide at widest part; cheliceral segment I as well as the proximal part of segment II smooth, a distal seta 10 (911) present.
<emphasis box="[971,1125,763,789]" italics="true" pageId="6" pageNumber="527">Subcapitulum</emphasis>
(
<figureCitation box="[1140,1225,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1946,1970]" captionTargetBox="[170,1422,539,1915]" captionTargetId="figure-240@4.[151,1436,525,1923]" captionTargetPageId="4" captionText="FIGURE 3. Cunaxa asiatica sp. nov., female (paratype).A—palp (dorsal view); B—subcapitulum (ventral view); C—chelicera (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7635233" httpUri="https://zenodo.org/record/7635233/files/figure.png" pageId="6" pageNumber="527">Fig. 3B</figureCitation>
) robust, 145 (144 145) long, with nearly rounded basal part 78 (7679) long and 85 (8387) wide, subcapitulum distally cone-shaped with two pairs of short adoral setae and four pairs of hypognathal setae (
<emphasis box="[966,994,835,861]" italics="true" pageId="6" pageNumber="527">hg</emphasis>
). Setae
<emphasis box="[1086,1122,835,867]" italics="true" pageId="6" pageNumber="527">
hg
<subScript attach="left" box="[1114,1122,853,867]" fontSize="6" pageId="6" pageNumber="527">4</subScript>
</emphasis>
48 (4351) long, about two times longer than
<emphasis box="[356,392,871,903]" italics="true" pageId="6" pageNumber="527">
hg
<subScript attach="left" box="[384,392,889,903]" fontSize="6" pageId="6" pageNumber="527">2</subScript>
</emphasis>
24 (2225), more than two times longer than
<emphasis box="[917,953,871,903]" italics="true" pageId="6" pageNumber="527">
hg
<subScript attach="left" box="[945,953,889,903]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
</emphasis>
22 (2222) and less than two times longer than
<emphasis box="[206,242,907,939]" italics="true" pageId="6" pageNumber="527">
hg
<subScript attach="left" box="[234,242,925,939]" fontSize="6" pageId="6" pageNumber="527">3</subScript>
</emphasis>
29 (2831). Coxal region of subcapitulum smooth.
</paragraph>
<paragraph blockId="6.[151,1437,151,1731]" pageId="6" pageNumber="527">
<emphasis box="[199,252,943,968]" italics="true" pageId="6" pageNumber="527">Legs</emphasis>
(
<figureCitation box="[269,341,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,250,1929,1953]" captionTargetBox="[166,1426,190,1895]" captionTargetId="figure-17@5.[151,1436,181,1905]" captionTargetPageId="5" captionText="FIGURE 4. Cunaxa asiatica sp. nov., female (holotype), legs IIV (left side, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7635237" httpUri="https://zenodo.org/record/7635237/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
). All legs smooth, legs IIII shorter than leg IV. Chaetotaxy IIV (excluding coxae) as follows: trochanters 1-1-1-1 sts; basifemora 4-4-2-1 sts; telofemora 4-4-4-4 sts; genu I—3
<emphasis box="[1084,1117,979,1005]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 4 sts; genu II—1
<emphasis box="[1327,1360,979,1005]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 5 sts; genu III—1
<emphasis box="[289,322,1015,1041]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 5 sts; genu IV—1
<emphasis box="[539,572,1015,1041]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 5sts; tibia I—1
<emphasis box="[758,791,1015,1041]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, (1
<emphasis box="[834,867,1015,1041]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 1 mst), 4 sts; tibia II—1
<emphasis box="[1153,1186,1015,1041]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
, 5 sts; tibia III—1
<emphasis box="[1397,1430,1015,1041]" italics="true" pageId="6" pageNumber="527">bsl</emphasis>
, 5 sts; tibia IV—1 smooth T, 4 sts; tarsus I—2
<emphasis box="[657,690,1051,1077]" italics="true" pageId="6" pageNumber="527">asl</emphasis>
,
<emphasis box="[703,745,1051,1077]" italics="true" pageId="6" pageNumber="527">fam</emphasis>
, 1
<emphasis box="[778,818,1051,1077]" italics="true" pageId="6" pageNumber="527">dtsl</emphasis>
, 11 sts, pretarsus with 1 dorsal and 1 ventral sts on each lobe; tarsus II—1
<emphasis box="[354,387,1087,1113]" italics="true" pageId="6" pageNumber="527">bsl</emphasis>
, 1
<emphasis box="[422,462,1087,1113]" italics="true" pageId="6" pageNumber="527">dtsl</emphasis>
, 12 sts, pretarsus with 1 dorsal and 1 ventral sts on each lobe; tarsus III—12 sts, 1
<emphasis box="[1389,1436,1087,1113]" italics="true" pageId="6" pageNumber="527">dtsl,</emphasis>
pretarsus with 1 dorsal and 1 ventral sts on each lobe; tarsus IV—10 sts, 1
<emphasis box="[979,1019,1123,1149]" italics="true" pageId="6" pageNumber="527">dtsl</emphasis>
, pretarsus with 1 dorsal and 1 ventral sts on each lobe. Length and width of leg segments IIV (coxa and trochanter not measured): basifemur 70 (6575) long, 29 (2830) wide; telofemur 28 (2532) long, 24 (2225]) wide; genu 27 (2528) long, 20 (2020) wide; tibia 37 (3440) long, 17 (1518) wide; tarsus 126 (125127) long, 16 (1517) wide at proximal part; total lengths of legs IIV: 288 (283293), 271 (265277), 303 (294311), 339 (336342).
</paragraph>
<paragraph blockId="6.[151,1437,151,1731]" box="[199,674,1302,1329]" pageId="6" pageNumber="527">
<emphasis bold="true" box="[199,549,1302,1329]" pageId="6" pageNumber="527">Male and preimaginal stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="527" type="materials_examined">
<paragraph blockId="6.[151,1437,151,1731]" pageId="6" pageNumber="527">
<emphasis bold="true" box="[199,435,1338,1364]" pageId="6" pageNumber="527">Material examined.</emphasis>
<materialsCitation collectingDate="2013-12-06" collectingDateMax="2013-12-07" collectingDateMin="2013-12-06" collectorName="A. E. Anichkin &amp; S. G. Ermilov" country="Vietnam" latitude="12.183333" location="Bi Dup - Nui Ba National Park" longLatPrecision="1293" longitude="108.683334" pageId="6" pageNumber="527" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="holotype">
<emphasis bold="true" box="[444,562,1339,1365]" pageId="6" pageNumber="527">
<typeStatus box="[444,557,1339,1365]" pageId="6" pageNumber="527">Holotype</typeStatus>
:
</emphasis>
<collectingCountry box="[572,670,1339,1365]" name="Vietnam" pageId="6" pageNumber="527">Vietnam</collectingCountry>
,
<collectingRegion box="[683,917,1339,1365]" country="Vietnam" name="Lam Dong" pageId="6" pageNumber="527">Lam Dong Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03A65F3BFFA7FFF67AB9FEE6BB06F931:8ED0B8F6FFA3FFF6798DFA96BA7BFAD9" box="[931,1295,1339,1365]" country="Vietnam" latitude="12.183333" longLatPrecision="1293" longitude="108.683334" name="Bi Dup - Nui Ba National Park" pageId="6" pageNumber="527" stateProvince="Lam Dong">Bi Dup—Nui Ba National Park</location>
, mixed pine dominated forest, 12º10
<geoCoordinate box="[456,550,1375,1401]" degrees="12" direction="north" minutes="11" orientation="latitude" pageId="6" pageNumber="527" precision="925" value="12.183333">12º11N</geoCoordinate>
, 108º40
<geoCoordinate box="[673,778,1375,1401]" degrees="108" direction="east" minutes="41" orientation="longitude" pageId="6" pageNumber="527" precision="925" value="108.683334">108º41E</geoCoordinate>
, soil and litter (sifter),
<date box="[1065,1307,1375,1401]" pageId="6" pageNumber="527" value="2013-12-06" valueMax="2013-12-07" valueMin="2013-12-06">
<collectingDate box="[1065,1307,1375,1401]" pageId="6" pageNumber="527" value="2013-12-06" valueMax="2013-12-07" valueMin="2013-12-06">67 December 2013</collectingDate>
</date>
, coll.
<collectorName pageId="6" pageNumber="527">A.E. Anichkin</collectorName>
and
<collectorName box="[316,468,1411,1437]" pageId="6" pageNumber="527">S.G. Ermilov</collectorName>
<emphasis bold="true" box="[468,496,1411,1437]" pageId="6" pageNumber="527"></emphasis>
<specimenCount box="[496,600,1411,1437]" pageId="6" pageNumber="527" type="female">1 female</specimenCount>
</materialsCitation>
.
<materialsCitation collectingDate="2013-12-02" collectorName="A. E. Anichkin &amp; S. G. Ermilov" country="Vietnam" latitude="11.383333" location="Biosphere Reserve" longLatPrecision="1295" longitude="107.36667" pageId="6" pageNumber="527" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai" typeStatus="paratype">
<emphasis bold="true" box="[613,731,1411,1437]" pageId="6" pageNumber="527">
<typeStatus box="[613,726,1411,1437]" pageId="6" pageNumber="527">Paratype</typeStatus>
:
</emphasis>
<collectingCountry box="[741,839,1411,1437]" name="Vietnam" pageId="6" pageNumber="527">Vietnam</collectingCountry>
,
<collectingRegion box="[853,1078,1411,1437]" country="Vietnam" name="Dong Nai" pageId="6" pageNumber="527">Dong Nai Province</collectingRegion>
,
<collectingRegion box="[1093,1205,1411,1437]" country="Vietnam" name="Dong Nai" pageId="6" pageNumber="527">Dong Nai</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03A65F3BFFA7FFF67AB9FEE6BB06F931:8ED0B8F6FFA3FFF67E91FA2EBAECFA11" box="[1215,1432,1411,1437]" country="Vietnam" latitude="11.383333" longLatPrecision="1295" longitude="107.36667" name="Biosphere Reserve" pageId="6" pageNumber="527" stateProvince="Dong Nai">Biosphere Reserve</location>
,
<geoCoordinate box="[151,254,1447,1473]" degrees="11" direction="north" minutes="23" orientation="latitude" pageId="6" pageNumber="527" precision="925" value="11.383333">11°23 N</geoCoordinate>
,
<geoCoordinate box="[265,379,1447,1473]" degrees="107" direction="east" minutes="22" orientation="longitude" pageId="6" pageNumber="527" precision="925" value="107.36667">107°22 E</geoCoordinate>
, forest plantations (mainly of
<taxonomicName box="[714,915,1447,1473]" class="Magnoliopsida" family="Dipterocarpaceae" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="527" phylum="Tracheophyta" rank="family">Dipterocarpaceae</taxonomicName>
), soil and litter (sifter),
<date box="[1176,1377,1447,1473]" pageId="6" pageNumber="527" value="2013-12-02">
<collectingDate box="[1176,1377,1447,1473]" pageId="6" pageNumber="527" value="2013-12-02">2 December 2013</collectingDate>
</date>
, coll.
<collectorName box="[151,312,1483,1509]" pageId="6" pageNumber="527">A.E. Anichkin</collectorName>
and
<collectorName box="[367,519,1483,1509]" pageId="6" pageNumber="527">S.G. Ermilov</collectorName>
<specimenCount box="[549,644,1483,1509]" pageId="6" pageNumber="527" type="female">1 female</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="6.[151,1437,151,1731]" box="[199,1180,1518,1545]" pageId="6" pageNumber="527">
<emphasis bold="true" box="[199,396,1518,1545]" pageId="6" pageNumber="527">Type depository.</emphasis>
<materialsCitation box="[403,1176,1519,1545]" collectionCode="SNMB" pageId="6" pageNumber="527" specimenCount="2" typeStatus="holotype">
The
<typeStatus box="[453,551,1519,1545]" pageId="6" pageNumber="527">holotype</typeStatus>
and
<typeStatus box="[606,702,1519,1545]" pageId="6" pageNumber="527">paratype</typeStatus>
are deposited in the collections of
<collectionCode box="[1093,1176,1519,1545]" country="Germany" httpUri="http://grbio.org/cool/jw6i-g070" name="Staatliches Naturhistorisches Museum" pageId="6" pageNumber="527">SNMB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,1234,1555,1581]" pageId="6" pageNumber="527" type="etymology">
<paragraph blockId="6.[151,1437,151,1731]" box="[199,1234,1555,1581]" pageId="6" pageNumber="527">
<emphasis bold="true" box="[199,334,1555,1581]" pageId="6" pageNumber="527">Etymology.</emphasis>
The specific epithet refers to the continent, where the species was found—Asia.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="527" type="discussion">
<paragraph blockId="6.[151,1437,151,1731]" pageId="6" pageNumber="527">
<emphasis bold="true" box="[199,315,1591,1617]" pageId="6" pageNumber="527">Remarks.</emphasis>
The new species,
<taxonomicName authority="Kalúz &amp; Ermilov, 2023" authorityName="Kalúz &amp; Ermilov" authorityYear="2023" box="[529,714,1591,1617]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[529,714,1591,1617]" italics="true" pageId="6" pageNumber="527">Cunaxa asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[722,821,1591,1617]" pageId="6" pageNumber="527">
<taxonomicNameLabel box="[722,815,1591,1617]" pageId="6" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
is morphologically most similar to
<taxonomicName authority="(Chaudri, 1980)" baseAuthorityName="Chaudri" baseAuthorityYear="1980" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="doxa">
<emphasis box="[1232,1320,1591,1617]" italics="true" pageId="6" pageNumber="527">C. doxa</emphasis>
(Chaudri, 1980)
</taxonomicName>
in having the medial hysterosomal shield smooth and bearing
<emphasis box="[959,1014,1627,1659]" italics="true" pageId="6" pageNumber="527">
c
<subScript attach="right" box="[972,980,1645,1659]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
e
<subScript attach="left" box="[1006,1014,1645,1659]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
</emphasis>
,
<emphasis box="[1033,1053,1627,1659]" italics="true" pageId="6" pageNumber="527">
c
<subScript attach="left" box="[1045,1053,1645,1659]" fontSize="6" pageId="6" pageNumber="527">2</subScript>
</emphasis>
. It differs from the latter by the presence of 1sts on basifemora IV, finger-like palpofemoral apophysis, and smooth setae
<emphasis box="[1174,1229,1663,1695]" italics="true" pageId="6" pageNumber="527">
c
<subScript attach="left" box="[1186,1194,1681,1695]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
e
<subScript attach="left" box="[1221,1229,1681,1695]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
</emphasis>
(vs. with 2 sts on basifemora IV, uncinate palpofemoral apophysis and spiculiform setae
<emphasis box="[946,993,1699,1731]" italics="true" pageId="6" pageNumber="527">
c
<subScript attach="right" box="[959,967,1717,1731]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
e
</emphasis>
<quantity box="[993,1027,1699,1731]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="527" unit="in" value="1.0">
<emphasis box="[993,1001,1717,1731]" italics="true" pageId="6" pageNumber="527">
<subScript attach="left" box="[993,1001,1717,1731]" fontSize="6" pageId="6" pageNumber="527">1</subScript>
</emphasis>
in
</quantity>
<taxonomicName baseAuthorityName="Chaudri" baseAuthorityYear="1980" box="[1034,1121,1699,1725]" class="Arachnida" family="Cunaxidae" genus="Cunaxa" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="doxa">
<emphasis box="[1034,1121,1699,1725]" italics="true" pageId="6" pageNumber="527">C. doxa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>