253 lines
34 KiB
XML
253 lines
34 KiB
XML
<document id="161455688813378C74D893F6867FCE08" ID-DOI="10.11646/zootaxa.4006.3.3" ID-GBIF-Dataset="3370151c-4fdf-4e5e-a759-a563046055a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="241009" ID-ZooBank="D601264E-A134-4341-A678-263678C68F32" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461212766545" checkinUser="plazi" docAuthor="Lopes-Andrade, Cristiano & Grebennikov, Vasily V." docDate="2015" docId="039E0263CA5FFFA074BD4D1CFA86D3E1" docLanguage="en" docName="zt04006p480.pdf" docOrigin="Zootaxa 4006 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syncosmetus medusa Lopes-Andrade et Grebennikov, sp. nov." docType="treatment" docVersion="8" lastPageNumber="471" masterDocId="FFA77A1BCA58FFA8742A4C3BFFE3D400" masterDocTitle="First record and five new species of Xylographellini (Coleoptera: Ciidae) from China, with online DNA barcode library of the family" masterLastPageNumber="480" masterPageNumber="463" pageNumber="470" updateTime="1698602272098" updateUser="plazi">
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<mods:titleInfo id="E4F9D32DA9254FB4B45BC1610BCB3565">
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<mods:title id="B47EC0DA8C02873988ED00FCCC6FAEBF">First record and five new species of Xylographellini (Coleoptera: Ciidae) from China, with online DNA barcode library of the family</mods:title>
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</mods:titleInfo>
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<mods:name id="0D3F67308459CC4A3343C700598E9A10" type="personal">
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<mods:roleTerm id="59BBA5CE358332395A068B58D6821B7C">Author</mods:roleTerm>
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<mods:namePart id="99F35B543C4C92561B2E9503F50B55B2">Lopes-Andrade, Cristiano</mods:namePart>
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<mods:name id="ABC36EC78A379A72E0279A90859B7AAD" type="personal">
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<mods:roleTerm id="61E34E0ECF279250F044E94473D336FB">Author</mods:roleTerm>
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<mods:namePart id="0772EB0E5C3A55C3EDA8D492803B8C59">Grebennikov, Vasily V.</mods:namePart>
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<mods:typeOfResource id="FFF6F2FEFE2512FAB0DC2FFB7BD4AE23">text</mods:typeOfResource>
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<mods:title id="C07BF4DAE754829773AEA45DB81A8238">Zootaxa</mods:title>
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<mods:part id="E0D6AC41C2DCA3B2A66474655ED77B09">
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<mods:date id="361881CD25AD8974B62574DBFD70DC40">2015</mods:date>
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<mods:detail id="87608EA6578529BFC875DAD989F0E65A" type="volume">
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<mods:number id="62C062D4D2F9E9935EBC022E8981014E">4006</mods:number>
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<mods:number id="7FD2ACECB4A736F2DFFB5024B80C509E">3</mods:number>
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<mods:classification id="2E9A2EB998ED5B5D1448F42F20904832">journal article</mods:classification>
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<mods:identifier id="7C198F804707578B5285B51EFD838363" type="DOI">10.11646/zootaxa.4006.3.3</mods:identifier>
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<mods:identifier id="1D8FC6DC3B001E1EB01A1A7A1AE23CEE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5AFF02DC9E36164F178DF94BDF11DF47" type="Zenodo-Dep">241009</mods:identifier>
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<treatment id="039E0263CA5FFFA074BD4D1CFA86D3E1" ID-DOI="http://doi.org/10.5281/zenodo.6103170" ID-GBIF-Taxon="119630541" ID-Zenodo-Dep="6103170" LSID="urn:lsid:plazi:treatment:039E0263CA5FFFA074BD4D1CFA86D3E1" httpUri="http://treatment.plazi.org/id/039E0263CA5FFFA074BD4D1CFA86D3E1" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470">
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<subSubSection id="C32DE0FECA5FFFAF74BD4D1CFE8CD563" pageId="7" pageNumber="470" type="nomenclature">
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<paragraph id="8B88B375CA5FFFAF74BD4D1CFC75D542" blockId="7.[151,918,295,355]" box="[151,918,295,322]" pageId="7" pageNumber="470">
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<heading id="D0C00419CA5FFFAF74BD4D1CFC75D542" bold="true" box="[151,918,295,322]" fontSize="11" level="1" pageId="7" pageNumber="470" reason="1">
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<emphasis id="B9436F67CA5FFFAF74BD4D1CFC75D542" bold="true" box="[151,918,295,322]" pageId="7" pageNumber="470">
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<taxonomicName id="4C37C8F6CA5FFFAF74BD4D1CFCC8D542" authority="Lopes-Andrade et Grebennikov" authorityName="Lopes-Andrade et Grebennikov" box="[151,811,295,322]" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="medusa" status="sp. nov.">
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<emphasis id="B9436F67CA5FFFAF74BD4D1CFE74D541" bold="true" box="[151,407,295,321]" italics="true" pageId="7" pageNumber="470">Syncosmetus medusa</emphasis>
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Lopes-Andrade et Grebennikov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A270D21CCA5FFFAF77124D13FC75D542" box="[824,918,296,322]" pageId="7" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B88B375CA5FFFAF74BD4D71FE8CD563" blockId="7.[151,918,295,355]" box="[151,367,330,355]" pageId="7" pageNumber="470">
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<figureCitation id="130CAFF0CA5FFFAF74BD4D71FEFED563" box="[151,285,330,355]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Figs. 15–21</figureCitation>
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,
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<figureCitation id="130CAFF0CA5FFFAF75034D71FEA4D563" box="[297,327,330,355]" captionStart="FIGURE 35" captionStartId="13.[151,249,857,879]" captionTargetBox="[151,1436,193,835]" captionTargetId="figure@13.[151,1436,193,836]" captionTargetPageId="13" captionText="FIGURE 35. Known distribution of Xylographellini beetles in the Palaearctic Region outside of Japan." httpUri="https://zenodo.org/record/241015/files/figure.png" pageId="7" pageNumber="470">35</figureCitation>
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,
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<figureCitation id="130CAFF0CA5FFFAF75794D70FE8CD563" box="[339,367,331,355]" captionStart="FIGURE 37" captionStartId="15.[151,249,1365,1386]" captionTargetBox="[151,1436,197,1331]" captionTargetId="figure@15.[151,1436,197,1331]" captionTargetPageId="15" captionText="FIGURE 37. Lower part of the Maximum Likelihood inference phylogram, upper part of which is depicted on Fig. 36. HT: holotype." httpUri="https://zenodo.org/record/241017/files/figure.png" pageId="7" pageNumber="470">37</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32DE0FECA5FFFAF74BD4DA9FD03D63B" pageId="7" pageNumber="470" type="diagnosis">
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<paragraph id="8B88B375CA5FFFAF74BD4DA9FD03D63B" blockId="7.[151,1437,402,2011]" pageId="7" pageNumber="470">
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<emphasis id="B9436F67CA5FFFAF74BD4DA9FEF2D5AB" bold="true" box="[151,273,402,427]" pageId="7" pageNumber="470">Diagnosis.</emphasis>
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Prothorax with sides explanate and visible for most of their lengths from above; edge of lateral carina minutely crenulated. Pronotum with longitudinal midline in low relief, mostly glabrous and unpunctate, surface microreticulate; anterior portion with surface between punctures somewhat in high relief; border along the posterior edge unpunctate, glabrous. Aedeagus with basal portion of tegmen homogeneously sclerotized, and apical portion with somewhat a spoon-like shape (
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<figureCitation id="130CAFF0CA5FFFAF76AA4E19FD30D63B" box="[640,723,546,571]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 19</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32DE0FECA5FFFA074ED4E7DFACCD293" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470" type="description">
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<paragraph id="8B88B375CA5FFFAF74ED4E7DFE16D3DB" blockId="7.[151,1437,402,2011]" pageId="7" pageNumber="470">
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<emphasis id="B9436F67CA5FFFAF74ED4E7DFDFDD65F" bold="true" box="[199,542,582,607]" pageId="7" pageNumber="470">
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Description,
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<typeStatus id="548C0DD7CA5FFFAF75414E7DFE31D65F" box="[363,466,582,607]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
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male
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</emphasis>
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(
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<figureCitation id="130CAFF0CA5FFFAF76134E7DFD28D65F" box="[569,715,582,607]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Figs. 15–21</figureCitation>
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). GenBank accession:
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<accessionNumber id="94642E96CA5FFFAF77DA4E7CFB93D65F" box="[1008,1136,583,607]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HQ987127" pageId="7" pageNumber="470">HQ987127</accessionNumber>
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. Fully pigmented adult. Measurements in mm: TL 1.53, PL 0.63, PW 0.83, EL 0.90, EW 0.98, GD 0.78. Ratios: PL/PW 0.76, EL/EW 0.92, EL/PL 1.44, GD/EW 0.79, TL/EW 1.56.
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<emphasis id="B9436F67CA5FFFAF76534EB5FD54D6A7" bold="true" box="[633,695,654,679]" pageId="7" pageNumber="470">Body</emphasis>
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rounded, extremely convex, mostly dark brown; venter slightly lighter than dorsum, with antennal funicle and palpi yellowish brown; dorsal vestiture of minute setae, conspicuously shorter than a puncture-width; ventral vestiture of slender decumbent setae, as long as a puncturewidth or less.
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<emphasis id="B9436F67CA5FFFAF751C4EC1FE95D713" bold="true" box="[310,374,762,787]" pageId="7" pageNumber="470">Head</emphasis>
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with coarse punctures, separated from each other by about a puncture-width; surface between punctures microreticulate; vertex with a median knob.
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<emphasis id="B9436F67CA5FFFAF773C4F25FC6AD737" bold="true" box="[790,905,798,823]" pageId="7" pageNumber="470">Antennae</emphasis>
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bearing ten antennomeres with the following lengths (in mm, left antenna measured): 0.09, 0.04, 0.03, 0.02, 0.02, 0.02, 0.02, 0.03, 0.04, 0.08 (FL
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<quantity id="4CCF1E90CA5FFFAF712D4F78FA8FD75A" box="[1287,1388,835,859]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="470" unit="mm" value="0.11">0.11 mm</quantity>
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, CL
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<quantity id="4CCF1E90CA5FFFAF74BD4F5DFF1FD77E" box="[151,252,870,895]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="470" unit="mm" value="0.15">0.15 mm</quantity>
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, CL/FL 1.36).
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<emphasis id="B9436F67CA5FFFAF75884F5DFE39D77F" bold="true" box="[418,474,870,895]" pageId="7" pageNumber="470">Eyes</emphasis>
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coarsely facetted and with about 25 ommatidia; GW
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<quantity id="4CCF1E90CA5FFFAF70074F5CFB70D77E" box="[1069,1171,871,895]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="470" unit="mm" value="0.11">0.11 mm</quantity>
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.
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<emphasis id="B9436F67CA5FFFAF70B74F5DFAF5D77F" bold="true" box="[1181,1302,870,895]" pageId="7" pageNumber="470">Pronotum</emphasis>
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with coarse punctures, irregular and separated from each other by about a puncture-width; in the anterior portion, the surface in between punctures is somewhat microstriated and tumid, forming lines in high relief; in the posterior portion, the surface is microreticulate and usually not tumid; longitudinal midline in low relief, mostly unpunctate, glabrous and smooth, forming a sulcus broadest and most conspicuous at disc, with surface microreticulate; anterior edge broadly rounded; anterior angles slightly projected forward; sides broad, explanate, weakly crenulate, with microreticulate and unpunctate surfaces, their posterior portion visible from above; posterior edge broadly rounded, the surface bordering it unpunctate, glabrous and somewhat granulate.
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<emphasis id="B9436F67CA5FFFAF70694859FB58D07B" bold="true" box="[1091,1211,1122,1147]" pageId="7" pageNumber="470">Scutellum</emphasis>
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very small, almost indiscernible, confused with the beginning of the median keel of elytra; BW
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<quantity id="4CCF1E90CA5FFFAF77C148BCFBB2D09E" box="[1003,1105,1159,1183]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="470" unit="mm" value="0.02">0.02 mm</quantity>
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.
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<emphasis id="B9436F67CA5FFFAF707148BDFB44D09F" bold="true" box="[1115,1191,1158,1183]" pageId="7" pageNumber="470">Elytra</emphasis>
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tightly locked but not fused, with punctures deep and coarse, separated from each other by at least a puncture-width and with smooth surface in between them; pseudoepipleura unpunctate and glabrous, their surface smooth and dull; each elytron with first longitudinal keel extending to about the middle of elytral edge; first two keels diverging close to elytral base (
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<figureCitation id="130CAFF0CA5FFFAF74F3492DFECED12F" box="[217,301,1302,1327]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 16</figureCitation>
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, arrow); second and third keels not united at their apices.
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<emphasis id="B9436F67CA5FFFAF77EF492DFBB0D12F" bold="true" box="[965,1107,1302,1327]" pageId="7" pageNumber="470">Prosternum</emphasis>
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in front of coxae biconcave, extremely tumid at middle and bearing a longitudinal carina; lateral concavities deep, with posterior margin carinate; punctation shallower at concavities than at disc and prosternal process, with surface in between punctures microreticulate.
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<emphasis id="B9436F67CA5FFFAF756549B9FE2CD19B" bold="true" box="[335,463,1410,1435]" pageId="7" pageNumber="470">Hypomera</emphasis>
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with shallow coarse punctures, the surface in between them microreticulate.
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<emphasis id="B9436F67CA5FFFAF711049B9FA7FD19B" bold="true" box="[1338,1436,1410,1435]" pageId="7" pageNumber="470">Protibia</emphasis>
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not expanded, three times as long as broad; inner facet with a conspicuous tuft of long bristles at the apical half; apical angle broadly rounded, its edge bearing a row of spines; outer edge apparently devoid of spines (a few spines visible at higher magnification).
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<emphasis id="B9436F67CA5FFFAF763C49D5FCF6D207" bold="true" box="[534,789,1518,1543]" pageId="7" pageNumber="470">Meso- and metatibia</emphasis>
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not expanded, four times as long as broad; outer edge apparently devoid of spines; outer apical angle rounded, not produced; apical edge with a row of spines.
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<emphasis id="B9436F67CA5FFFAF74BD4A0DFED0D24F" bold="true" box="[151,307,1590,1615]" pageId="7" pageNumber="470">Metaventrite</emphasis>
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without discrimen; punctures very coarse and separated from each other by a puncture-width or less; surface in between punctures microreticulate.
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<emphasis id="B9436F67CA5FFFAF76824A61FC43D273" bold="true" box="[680,928,1626,1651]" pageId="7" pageNumber="470">Abdominal ventrites</emphasis>
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with very coarse punctures, concentrated at the anterior portion of each ventrite and separated by approximately a puncture-width; surface in between punctures and elsewhere microreticulate; length of the ventrites (in mm, from base to apex at the longitudinal midline) as follows: 0.22, 0.04, 0.04, 0.04, 0.13; first ventrite bearing a small circular, foveate sex mark, located anterad of center and with a transverse diameter of
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<quantity id="4CCF1E90CA5FFFAF76C94AD0FCA8D302" box="[739,843,1771,1795]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="470" unit="mm" value="0.02">0.02 mm</quantity>
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.
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<emphasis id="B9436F67CA5FFFAF777D4AD1FABDD303" bold="true" box="[855,1374,1770,1795]" pageId="7" pageNumber="470">
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Male abdominal terminalia in the
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<typeStatus id="548C0DD7CA5FFFAF70DD4AD1FABDD303" box="[1271,1374,1770,1795]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="130CAFF0CA5FFFAF71444AD1FF3DD327" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 18–21</figureCitation>
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) with
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<emphasis id="B9436F67CA5FFFAF750C4B35FE5FD327" bold="true" box="[294,444,1806,1831]" pageId="7" pageNumber="470">sternite VIII</emphasis>
|
||
(
|
||
<figureCitation id="130CAFF0CA5FFFAF75E14B35FDFFD327" box="[459,540,1806,1831]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 18</figureCitation>
|
||
) subtrapezoidal; posterior edge narrow, curved inward; posterior angles bearing a few long setae.
|
||
<emphasis id="B9436F67CA5FFFAF75704B09FE56D34B" bold="true" box="[346,437,1842,1867]" pageId="7" pageNumber="470">Tegmen</emphasis>
|
||
(
|
||
<figureCitation id="130CAFF0CA5FFFAF75ED4B09FDFAD34B" box="[455,537,1842,1867]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 19</figureCitation>
|
||
) with basal edge mostly straight, with a short subtriangular extension at middle; lateral edges subparallel at basal two-thirds; apex enlarged and broadly rounded, most sclerotized laterally and bearing a broad concavity, making the tegmen similar to a spoon.
|
||
<emphasis id="B9436F67CA5FFFAF77B64B41FBC6D393" bold="true" box="[924,1061,1914,1939]" pageId="7" pageNumber="470">Basal piece</emphasis>
|
||
(
|
||
<figureCitation id="130CAFF0CA5FFFAF70124B41FB6FD393" box="[1080,1164,1914,1939]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 20</figureCitation>
|
||
) subquadrate, the sides slightly convergent to the anterior portion.
|
||
<emphasis id="B9436F67CA5FFFAF76514BA5FD58D3B7" bold="true" box="[635,699,1950,1975]" pageId="7" pageNumber="470">Penis</emphasis>
|
||
(
|
||
<figureCitation id="130CAFF0CA5FFFAF76E04BA5FCFFD3B7" box="[714,796,1950,1975]" captionStart="FIGURES 15 – 21" captionStartId="8.[151,263,1252,1275]" captionTargetBox="[167,1409,192,1219]" captionTargetId="figure@8.[166,1418,179,1226]" captionTargetPageId="8" captionText="FIGURES 15 – 21. Syncosmetus medusa sp. nov., male holotype from Emei Shan (Figs. 15 – 17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18 – 21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15 – 17); 0.1 mm (Figs. 18 – 21)." httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="7" pageNumber="470">Fig. 21</figureCitation>
|
||
) very elongate; lateral struts linked at base; lateral edges subparallel; apex membranous.
|
||
</paragraph>
|
||
<caption id="DF48E3FDCA50FFA074BD48DFFEB7D182" httpUri="https://zenodo.org/record/241012/files/figure.png" pageId="8" pageNumber="471" targetBox="[167,1409,192,1219]" targetPageId="8">
|
||
<paragraph id="8B88B375CA50FFA074BD48DFFEB7D182" blockId="8.[151,1436,1252,1410]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074BD48DFFEB7D0FA" bold="true" box="[151,340,1252,1275]" pageId="8" pageNumber="471">FIGURES 15–21.</emphasis>
|
||
<taxonomicName id="4C37C8F6CA50FFA0757148DDFDD3D0FB" box="[347,560,1253,1275]" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="medusa" status="sp. nov.">
|
||
<emphasis id="B9436F67CA50FFA0757148DDFDD3D0FB" box="[347,560,1253,1275]" italics="true" pageId="8" pageNumber="471">Syncosmetus medusa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270D21CCA50FFA0761248DDFD6AD0FB" box="[568,649,1254,1275]" pageId="8" pageNumber="471" rank="species">
|
||
<emphasis id="B9436F67CA50FFA0761248DDFD6AD0FB" bold="true" box="[568,649,1254,1275]" pageId="8" pageNumber="471">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype from Emei Shan (Figs. 15–17): (15) dorsal view; (16) lateral view, with arrow showing first two elytral keels diverging close to elytral base; (17) ventral view. Pregenital segment and aedeagus of a paratype from the type locality (Figs. 18–21): (18) sternite VIII; (19) tegmen; (20) basal piece (= phallobase); (21) penis. The apices of sclerites of male abdominal terminalia are oriented to the left. Scale bars: 0.5 mm (Figs. 15–17); 0.1 mm (Figs. 18–21).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B88B375CA50FFA074ED4989FCF9D20E" blockId="8.[151,1437,1458,2017]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074ED4989FED3D1CB" bold="true" box="[199,304,1458,1483]" pageId="8" pageNumber="471">Females.</emphasis>
|
||
Similar to male, but devoid of abdominal sex patch.
|
||
<emphasis id="B9436F67CA50FFA077CE4989FABAD1CB" bold="true" box="[996,1369,1458,1483]" pageId="8" pageNumber="471">Female abdominal terminalia</emphasis>
|
||
with conspicuous gonostyli; gonocoxites transversely divided into two parts; paraprocts shorter than gonocoxites together; opening of proctiger close to basal gonocoxites.
|
||
</paragraph>
|
||
<paragraph id="8B88B375CA50FFA074ED4A2DFACCD293" blockId="8.[151,1437,1458,2017]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074ED4A2DFED8D22F" bold="true" box="[199,315,1558,1583]" pageId="8" pageNumber="471">Variation.</emphasis>
|
||
No male other than the
|
||
<typeStatus id="548C0DD7CA50FFA0767D4A2DFD5AD22F" box="[599,697,1558,1583]" pageId="8" pageNumber="471" type="holotype">holotype</typeStatus>
|
||
is known. Females, measurements in mm (n = 2): TL 1.68–1.80 (1.74 ± 0.09), PL 0.70–0.75 (0.73 ± 0.04), PW 0.83–0.88(0.85 ± 0.04), EL 0.93–1.00 (0.96 ± 0.05), EW 1.08–1.13 (1.10 ± 0.04), GD 0.85–0.90 (0.88 ± 0.04). Ratios: PL/PW 0.80–0.91 (0.85 ± 0.08), EL/EW 0.86–0.89 (0.87 ± 0.02), EL/PL 1.32–1.33 (1.33 ± 0.01), GD/EW 0.79–0.80 (0.80 ± 0.01), TL/EW 1.56–1.60 (1.58 ± 0.03).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DE0FECA50FFA074ED4AA7FBB6D37C" pageId="8" pageNumber="471" type="materials_examined">
|
||
<paragraph id="8B88B375CA50FFA074ED4AA7FBB6D37C" blockId="8.[151,1437,1458,2017]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074ED4AA7FE0DD2B5" bold="true" box="[199,494,1692,1717]" pageId="8" pageNumber="471">
|
||
<typeStatus id="548C0DD7CA50FFA074ED4AA7FEE1D2B5" box="[199,258,1692,1717]" pageId="8" pageNumber="471">Type</typeStatus>
|
||
series.
|
||
<typeStatus id="548C0DD7CA50FFA0755D4AA7FE0BD2B5" box="[375,488,1692,1717]" pageId="8" pageNumber="471" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
male (dissected,
|
||
<collectionCode id="ED262BB0CA50FFA076F74AA7FCD3D2B4" box="[733,816,1692,1716]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="8" pageNumber="471">IZCAS</collectionCode>
|
||
) “P.R.
|
||
<collectingCountry id="F320F3E5CA50FFA077B74AA7FC00D2B5" box="[925,995,1692,1717]" name="China" pageId="8" pageNumber="471">China</collectingCountry>
|
||
, Sichuan, Emeishan,
|
||
<geoCoordinate id="EE03D5B2CA50FFA0712A4AA7FA7FD2B5" box="[1280,1436,1692,1717]" direction="north" orientation="latitude" pageId="8" pageNumber="471" precision="1" value="29.560083">N29º33’36.3”</geoCoordinate>
|
||
<geoCoordinate id="EE03D5B2CA50FFA074BD4A85FEDDD2D6" box="[151,318,1726,1750]" direction="east" orientation="longitude" pageId="8" pageNumber="471" precision="1" value="103.343895">E103º20’38.0”</geoCoordinate>
|
||
<date id="FF8995B5CA50FFA075604A86FE27D2D6" box="[330,452,1725,1750]" pageId="8" pageNumber="471" value="2010-06-22">22.vi.2010</date>
|
||
, 1947m sifting39, V. Grebennikov \ CNCCOLVG00000915 \
|
||
<taxonomicName id="4C37C8F6CA50FFA070804A85FDE5D2F7" authority="Lopes-Andrade & Grebennikov" authorityName="Lopes-Andrade & Grebennikov" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="medusa">
|
||
<emphasis id="B9436F67CA50FFA070804A85FA78D2D6" box="[1194,1435,1725,1750]" italics="true" pageId="8" pageNumber="471">Syncosmetus medusa</emphasis>
|
||
Lopes-Andrade & Grebennikov
|
||
</taxonomicName>
|
||
<typeStatus id="548C0DD7CA50FFA076384AE4FD5BD2F7" box="[530,696,1759,1783]" pageId="8" pageNumber="471" type="holotype">HOLOTYPUS</typeStatus>
|
||
[handwritten, red paper]”.
|
||
<emphasis id="B9436F67CA50FFA077D54AE5FB63D2F7" bold="true" box="[1023,1152,1758,1783]" pageId="8" pageNumber="471">
|
||
<typeStatus id="548C0DD7CA50FFA077D54AE5FB98D2F7" box="[1023,1147,1758,1783]" pageId="8" pageNumber="471" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D3178FCCA50FFA070A74AE4FB17D2F7" box="[1165,1268,1758,1783]" pageId="8" pageNumber="471" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED262BB0CA50FFA071224AE4FAADD2F7" box="[1288,1358,1759,1783]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="8" pageNumber="471">CNCI</collectionCode>
|
||
) “P.R.
|
||
<collectingCountry id="F320F3E5CA50FFA074BD4B3BFF3ED319" box="[151,221,1792,1817]" name="China" pageId="8" pageNumber="471">China</collectingCountry>
|
||
, Sichuan, Emeishan,
|
||
<geoCoordinate id="EE03D5B2CA50FFA075C44B3BFD69D318" box="[494,650,1792,1816]" direction="north" orientation="latitude" pageId="8" pageNumber="471" precision="1" value="29.55275">N29º33’09.9”</geoCoordinate>
|
||
E103º20.39.1”
|
||
<date id="FF8995B5CA50FFA077784B3BFC2FD319" box="[850,972,1792,1817]" pageId="8" pageNumber="471" value="2010-06-16">16.vi.2010</date>
|
||
, 2266m, sifting34, V. Grebennikov \ CNCCOLVG00000916”;
|
||
<specimenCount id="9D3178FCCA50FFA075EE4B19FDCDD33A" box="[452,558,1825,1850]" pageId="8" pageNumber="471" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED262BB0CA50FFA0766D4B19FD71D33A" box="[583,658,1826,1850]" pageId="8" pageNumber="471">LAPC</collectionCode>
|
||
) “P.R.
|
||
<collectingCountry id="F320F3E5CA50FFA076DF4B1AFCD8D33A" box="[757,827,1825,1850]" name="China" pageId="8" pageNumber="471">China</collectingCountry>
|
||
, Sichuan, Emeishan,
|
||
<geoCoordinate id="EE03D5B2CA50FFA070634B19FB07D33A" box="[1097,1252,1826,1850]" direction="north" orientation="latitude" pageId="8" pageNumber="471" precision="1" value="29.546778">N29º32’48.4”</geoCoordinate>
|
||
<geoCoordinate id="EE03D5B2CA50FFA070DC4B1AFA7FD33A" box="[1270,1436,1825,1850]" direction="east" orientation="longitude" pageId="8" pageNumber="471" precision="1" value="103.33508">E103º20’06.3”</geoCoordinate>
|
||
<date id="FF8995B5CA50FFA074BD4B79FEF2D35B" box="[151,273,1858,1883]" pageId="8" pageNumber="471" value="2010-06-17">17.vi.2010</date>
|
||
, 2342m, sifting36, V. Grebennikov \ CNCCOLVG00000917”. All
|
||
<typeStatus id="548C0DD7CA50FFA0701C4B78FB42D35B" box="[1078,1185,1859,1883]" pageId="8" pageNumber="471" type="paratype">paratypes</typeStatus>
|
||
additionally labelled \
|
||
<taxonomicName id="4C37C8F6CA50FFA074B44B5EFD16D37D" authority="Lopes-Andrade & Grebennikov" authorityName="Lopes-Andrade & Grebennikov" box="[158,757,1892,1917]" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="medusa">
|
||
<emphasis id="B9436F67CA50FFA074B44B5EFE69D37D" box="[158,394,1892,1917]" italics="true" pageId="8" pageNumber="471">Syncosmetus medusa</emphasis>
|
||
Lopes-Andrade & Grebennikov
|
||
</taxonomicName>
|
||
<typeStatus id="548C0DD7CA50FFA076D64B5FFC79D37C" box="[764,922,1892,1916]" pageId="8" pageNumber="471" type="paratype">PARATYPUS</typeStatus>
|
||
[yellow paper]\.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DE0FECA50FFA074ED4BBEFCFED3BF" pageId="8" pageNumber="471" type="etymology">
|
||
<paragraph id="8B88B375CA50FFA074ED4BBEFCFED3BF" blockId="8.[151,1437,1458,2017]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074ED4BBEFEADD39E" bold="true" box="[199,334,1925,1950]" pageId="8" pageNumber="471">Etymology.</emphasis>
|
||
The species epithet is a Latinized Greek mythical name of the youngest of the three Gorgon sisters, beheaded by the hero
|
||
<taxonomicName id="4C37C8F6CA50FFA075CA4B93FDDAD3BF" box="[480,569,1960,1983]" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="476" phylum="Arthropoda" rank="species" species="perseus" status="sp. nov.">Perseus</taxonomicName>
|
||
; noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DE0FECA50FFA074ED4BF3FA86D3E1" box="[199,1381,1992,2017]" pageId="8" pageNumber="471" type="discussion">
|
||
<paragraph id="8B88B375CA50FFA074ED4BF3FA86D3E1" blockId="8.[151,1437,1458,2017]" box="[199,1381,1992,2017]" pageId="8" pageNumber="471">
|
||
<emphasis id="B9436F67CA50FFA074ED4BF3FEACD3E1" bold="true" box="[199,335,1992,2017]" pageId="8" pageNumber="471">Comments.</emphasis>
|
||
The species closely resembles
|
||
<emphasis id="B9436F67CA50FFA076874BF2FCA7D3E1" box="[685,836,1992,2017]" italics="true" pageId="8" pageNumber="471">
|
||
Sync.
|
||
<taxonomicName id="4C37C8F6CA50FFA076DA4BF3FCA7D3E1" box="[752,836,1992,2017]" class="Insecta" family="Ciidae" genus="Syncosmetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="euryale" status="sp. nov.">euryale</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A270D21CCA50FFA077614BF3FC46D3E1" box="[843,933,1992,2017]" pageId="8" pageNumber="471" rank="species">
|
||
<emphasis id="B9436F67CA50FFA077614BF3FC46D3E1" bold="true" box="[843,933,1992,2017]" pageId="8" pageNumber="471">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, which is known from Gaoligong Shan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |