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<mods:title id="C6421E4559FAD2DD627ECF7FCEE07694">Taxonomy in the phylogenomic era: species boundaries and phylogenetic relationships among North American ants of the Crematogaster scutellaris group (Formicidae: Hymenoptera)</mods:title>
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<mods:namePart id="35C2414048B910DB14C5963AF5CFFDFD">Ward, Philip S.</mods:namePart>
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<mods:date id="FE59F014A08EDA2CDF955894F911A516">2022</mods:date>
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<subSubSection id="C32A657BFFE5FFC9FC3E78F2FAEE638A" box="[925,1344,1426,1451]" pageId="25" pageNumber="918" type="nomenclature">
<paragraph id="8B8F36F0FFE5FFC9FC3E78F2FAEE638A" blockId="25.[925,1344,1426,1451]" box="[925,1344,1426,1451]" pageId="25" pageNumber="918">
<heading id="D0C7819CFFE5FFC9FC3E78F2FAEE638A" box="[925,1344,1426,1451]" centered="true" fontSize="9" level="2" pageId="25" pageNumber="918" reason="2">
<taxonomicName id="4C304D73FFE5FFC9FC3E78F2FAEE638A" ID-CoL="ZBT7" authority="MAYR, 1870" authorityName="Mayr" authorityYear="1870" box="[925,1344,1426,1451]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE5FFC9FC3E78F2FB1F638B" box="[925,1201,1426,1450]" italics="true" pageId="25" pageNumber="918">CREMATOGASTER OPACA</emphasis>
<bibRefCitation id="EFA14B01FFE5FFC9FB1B78F2FAEE638A" author="Mayr G" box="[1208,1344,1426,1451]" pageId="25" pageNumber="918" pagination="939 - 996" refId="ref36481" refString="Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 20: 939 - 996." type="journal article" year="1870">MAYR, 1870</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A657BFFE5FFCAFC9878DBFEB366DB" lastPageId="26" lastPageNumber="919" pageId="25" pageNumber="918" type="reference_group">
<paragraph id="8B8F36F0FFE5FFC9FC9878DBFB456048" blockId="25.[827,1443,1467,1856]" pageId="25" pageNumber="918">
<treatmentCitationGroup id="AB2011DEFFE5FFC9FC9878DBFB456048" pageId="25" pageNumber="918">
<treatmentCitation id="0A9110E1FFE5FFC9FC9878DBFA8563F1" author="Mayr G" box="[827,1323,1467,1488]" page="992" pageId="25" pageNumber="918" year="1870">
<taxonomicName id="4C304D73FFE5FFC9FC9878DBFA8563F1" ID-CoL="ZBT7" authority="Mayr, 1870: 992" authorityName="Mayr" authorityPageNumber="992" authorityYear="1870" box="[827,1323,1467,1488]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE5FFC9FC9878DBFBE763F1" box="[827,1097,1467,1488]" italics="true" pageId="25" pageNumber="918">Crematogaster opaca</emphasis>
<bibRefCitation id="EFA14B01FFE5FFC9FBF578DBFA8563F1" author="Mayr G" box="[1110,1323,1467,1488]" pageId="25" pageNumber="918" pagination="939 - 996" refId="ref36481" refString="Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 20: 939 - 996." type="journal article" year="1870">Mayr, 1870: 992</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="548B8852FFE5FFC9FA9F78DBFA0F63F1" box="[1340,1441,1467,1488]" pageId="25" pageNumber="918" type="syntype">Syntype</typeStatus>
<materialsCitation id="3B583CADFFE5FFC9FCF078B9FC5E600D" collectionCode="NHMW" country="Mexico" location="Tabukaya" pageId="25" pageNumber="918" specimenCount="1" typeStatus="syntype">
worker,
<location id="8EEF602BFFE5FFC9FC0E78B9FB8E63CE" LSID="urn:lsid:plazi:treatment:039987E6FFE5FFCAFC3E78F2FB216755:8EEF602BFFE5FFC9FC0E78B9FB8E63CE" box="[941,1056,1497,1519]" country="Mexico" name="Tabukaya" pageId="25" pageNumber="918">Tabukaya</location>
,
<collectingCountry id="F3277660FFE5FFC9FB8F78B9FBD063CE" box="[1068,1150,1497,1519]" name="Mexico" pageId="25" pageNumber="918">Mexico</collectingCountry>
, 1865 (Bilimek) (
<collectionCode id="ED21AE35FFE5FFC9FAE278B9FA3263CE" box="[1345,1436,1497,1519]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="25" pageNumber="918" type="Museum">NHMW</collectionCode>
) (examined via AntWeb image: CASENT0919705) and
<typeStatus id="548B8852FFE5FFC9FC2E7B77FC5E600D" box="[909,1008,1559,1580]" pageId="25" pageNumber="918" type="syntype">syntype</typeStatus>
</materialsCitation>
<materialsCitation id="3B583CADFFE5FFC9FC5E7B76FB4A6048" country="Mexico" location="Mexico" pageId="25" pageNumber="918" specimenCount="1" typeStatus="lectotype">
worker,
<collectingCountry id="F3277660FFE5FFC9FBC47B76FB11600D" box="[1127,1215,1558,1580]" name="Mexico" pageId="25" pageNumber="918">Mexico</collectingCountry>
(Sichel) (BMNH) (examined via AntWeb image: CASENT0902152); latter
<emphasis id="B944EAE2FFE5FFC9FC3A7B34FB4A6048" bold="true" box="[921,1252,1620,1642]" pageId="25" pageNumber="918">
here designated
<typeStatus id="548B8852FFE5FFC9FBCD7B34FB4A6048" box="[1134,1252,1620,1641]" pageId="25" pageNumber="918" type="lectotype">lectotype</typeStatus>
</emphasis>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F36F0FFE5FFC9FC987B13FC696087" blockId="25.[827,1443,1467,1856]" pageId="25" pageNumber="918">
<taxonomicName id="4C304D73FFE5FFC9FC987B13FAEA60A9" ID-CoL="ZBT7" authority="Mayr" authorityName="Mayr" authorityYear="1870" box="[827,1348,1650,1672]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="subSpecies" species="lineolata" subSpecies="opaca">
<emphasis id="B944EAE2FFE5FFC9FC987B13FBF460A6" box="[827,1114,1650,1672]" italics="true" pageId="25" pageNumber="918">Crematogaster lineolata</emphasis>
subsp.
<emphasis id="B944EAE2FFE5FFC9FB157B13FB5460A9" box="[1206,1274,1651,1672]" italics="true" pageId="25" pageNumber="918">opaca</emphasis>
Mayr
</taxonomicName>
; Emery, 1895: 283.
</paragraph>
<paragraph id="8B8F36F0FFE5FFC9FC987BD0FAF860E4" blockId="25.[827,1443,1467,1856]" box="[827,1366,1712,1734]" pageId="25" pageNumber="918">
<treatmentCitationGroup id="AB2011DEFFE5FFC9FC987BD0FAF860E4" box="[827,1366,1712,1734]" pageId="25" pageNumber="918">
<taxonomicName id="4C304D73FFE5FFC9FC987BD0FBC160E4" ID-CoL="ZBT7" authority="Mayr" authorityName="Mayr" authorityYear="1870" box="[827,1135,1712,1733]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE5FFC9FC987BD0FB8760E4" box="[827,1065,1712,1733]" italics="true" pageId="25" pageNumber="918">Crematogaster opaca</emphasis>
Mayr
</taxonomicName>
;
<treatmentCitation id="0A9110E1FFE5FFC9FBDA7BD0FAFC60E4" author="Wheeler WM" box="[1145,1362,1712,1734]" page="111" pageId="25" pageNumber="918" year="1919">
<bibRefCitation id="EFA14B01FFE5FFC9FBDA7BD0FAFC60E4" author="Wheeler WM" box="[1145,1362,1712,1734]" pageId="25" pageNumber="918" pagination="107 - 114" refId="ref37693" refString="Wheeler WM. 1919. A new paper-making Crematogaster from the south-eastern United States. Psyche (Cambridge) 26: 107 - 114." type="journal article" year="1919">Wheeler, 1919: 111</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F36F0FFE5FFC9FC987BAFFBC86123" blockId="25.[827,1443,1467,1856]" pageId="25" pageNumber="918">
<treatmentCitationGroup id="AB2011DEFFE5FFC9FC987BAFFBC86123" pageId="25" pageNumber="918">
<taxonomicName id="4C304D73FFE5FFC9FC987BAFFC3C6122" ID-CoL="ZBT7" authority="Mayr" authorityName="Mayr" authorityYear="1870" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="subSpecies" species="lineolata" subGenus="Acrocoelia" subSpecies="opaca">
<emphasis id="B944EAE2FFE5FFC9FC987BAFFC4260C5" box="[827,1004,1743,1764]" italics="true" pageId="25" pageNumber="918">Crematogaster</emphasis>
(
<emphasis id="B944EAE2FFE5FFC9FBA37BAEFBD360C2" box="[1024,1149,1742,1763]" italics="true" pageId="25" pageNumber="918">Acrocoelia</emphasis>
)
<emphasis id="B944EAE2FFE5FFC9FB337BAEFB5260C2" box="[1168,1276,1742,1763]" italics="true" pageId="25" pageNumber="918">lineolata</emphasis>
subsp.
<emphasis id="B944EAE2FFE5FFC9FAFE7BAFFA0D60C5" box="[1373,1443,1743,1764]" italics="true" pageId="25" pageNumber="918">opaca</emphasis>
Mayr
</taxonomicName>
;
<treatmentCitation id="0A9110E1FFE5FFC9FC3F7B8EFBCC6123" author="Emery C" box="[924,1122,1773,1795]" page="141" pageId="25" pageNumber="918" year="1922">
<bibRefCitation id="EFA14B01FFE5FFC9FC3F7B8EFBCC6123" author="Emery C" box="[924,1122,1773,1795]" pageId="25" pageNumber="918" pagination="95 - 206" refId="ref35295" refString="Emery C. 1922. Hymenoptera, fam. Formicidae, subfam. Myrmicinae. Genera Insectorum 174 B: 95 - 206." type="journal article" year="1922">Emery, 1922: 141</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F36F0FFE5FFC9FC987A6CFC7A6161" blockId="25.[827,1443,1467,1856]" pageId="25" pageNumber="918">
<treatmentCitationGroup id="AB2011DEFFE5FFC9FC987A6CFC7A6161" pageId="25" pageNumber="918">
<taxonomicName id="4C304D73FFE5FFC9FC987A6CFA846100" ID-CoL="ZBT7" authority="Mayr" authorityName="Mayr" authorityYear="1870" box="[827,1322,1804,1825]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="918" phylum="Arthropoda" rank="species" species="opaca" subGenus="Acrocoelia">
<emphasis id="B944EAE2FFE5FFC9FC987A6CFC406100" box="[827,1006,1804,1825]" italics="true" pageId="25" pageNumber="918">Crematogaster</emphasis>
(
<emphasis id="B944EAE2FFE5FFC9FBA17A6CFB2F6100" box="[1026,1153,1804,1825]" italics="true" pageId="25" pageNumber="918">Acrocoelia</emphasis>
)
<emphasis id="B944EAE2FFE5FFC9FB367A6CFB756100" box="[1173,1243,1804,1825]" italics="true" pageId="25" pageNumber="918">opaca</emphasis>
Mayr
</taxonomicName>
;
<treatmentCitation id="0A9110E1FFE5FFC9FA9B7A6CFC7E6161" author="Wheeler WM" page="139" pageId="25" pageNumber="918" year="1934">
<bibRefCitation id="EFA14B01FFE5FFC9FA9B7A6CFC7E6161" author="Wheeler WM" pageId="25" pageNumber="918" pagination="132 - 144" refId="ref37791" refString="Wheeler WM. 1934 a. Ants from the islands off the west coast of Lower California and Mexico. Pan-Pacific Entomologist 10: 132 - 144." type="journal article" year="1934">Wheeler, 1934a: 139</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8F36F0FFE6FFCAFF327DA6FEB366DB" blockId="26.[145,760,197,250]" pageId="26" pageNumber="919">
<treatmentCitationGroup id="AB2011DEFFE6FFCAFF327DA6FEB366DB" pageId="26" pageNumber="919">
<taxonomicName id="4C304D73FFE6FFCAFF327DA6FD3366FA" ID-CoL="ZBT7" authority="Mayr" authorityName="Mayr" authorityYear="1870" box="[145,669,197,219]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca" subGenus="Crematogaster">
<emphasis id="B944EAE2FFE6FFCAFF327DA6FE9066FA" box="[145,318,198,219]" italics="true" pageId="26" pageNumber="919">Crematogaster</emphasis>
(
<emphasis id="B944EAE2FFE6FFCAFEF37DA6FE5366FA" box="[336,509,198,219]" italics="true" pageId="26" pageNumber="919">Crematogaster</emphasis>
)
<emphasis id="B944EAE2FFE6FFCAFDAD7DA6FDFC66FA" box="[526,594,198,219]" italics="true" pageId="26" pageNumber="919">opaca</emphasis>
Mayr
</taxonomicName>
;
<treatmentCitation id="0A9110E1FFE6FFCAFD097DA5FEB666DB" author="Smith MR" page="126" pageId="26" pageNumber="919" year="1958">
<bibRefCitation id="EFA14B01FFE6FFCAFD097DA5FEB666DB" author="Smith MR" pageId="26" pageNumber="919" pagination="1 - 305" refId="ref37230" refString="Smith MR. 1958. Family Formicidae. Pp. 108 - 162 in: Krombein KV, ed. Hymenoptera of America north of Mexico. Synoptic catalogue. First supplement. United States Department of Agriculture. Agriculture Monograph 2 (suppl. 1): 1 - 305." type="journal article" year="1958">Smith, 1958: 126</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32A657BFFE6FFCAFF327C43FB216755" pageId="26" pageNumber="919" type="discussion">
<paragraph id="8B8F36F0FFE6FFCAFF327C43FEFB607E" blockId="26.[145,762,290,1907]" pageId="26" pageNumber="919">
<emphasis id="B944EAE2FFE6FFCAFF327C43FEA16719" box="[145,271,291,312]" italics="true" pageId="26" pageNumber="919">Discussion:</emphasis>
<bibRefCitation id="EFA14B01FFE6FFCAFEB67C43FD9E6718" author="Morgan C &amp; Mackay W" box="[277,560,290,313]" pageId="26" pageNumber="919" refId="ref36528" refString="Morgan C, Mackay W. 2017. The North America acrobat ants of the hyperdiverse genus Crematogaster. Balti, Moldova: Lambert Academic Publishing." type="book" year="2017">Morgan &amp; Mackay (2017)</bibRefCitation>
refer to a
<typeStatus id="548B8852FFE6FFCAFD027C43FD576719" box="[673,761,291,312]" pageId="26" pageNumber="919" type="syntype">syntype</typeStatus>
worker of
<taxonomicName id="4C304D73FFE6FFCAFEAA7C22FEC56776" authorityName="Mayr" authorityYear="1870" box="[265,363,322,343]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE6FFCAFEAA7C22FEC56776" box="[265,363,322,343]" italics="true" pageId="26" pageNumber="919">C. opaca</emphasis>
</taxonomicName>
in NHMW collected by Professor Bilimek and Dr Sichel in México. This statement is misleading because there is no single
<typeStatus id="548B8852FFE6FFCAFD997C1FFD3C67B5" box="[570,658,383,404]" pageId="26" pageNumber="919" type="syntype">syntype</typeStatus>
collected by Bilimek and Sichel; their collections were separate. In the BMNH there is a
<typeStatus id="548B8852FFE6FFCAFE257CDCFE7067F0" box="[390,478,444,465]" pageId="26" pageNumber="919" type="syntype">syntype</typeStatus>
worker (here designated
<typeStatus id="548B8852FFE6FFCAFF327CBAFF5567D1" box="[145,251,474,496]" pageId="26" pageNumber="919" type="lectotype">lectotype</typeStatus>
) (CASENT0902152) labelled Mex.|Sichel, while the NHMW
<typeStatus id="548B8852FFE6FFCAFEDB7C9AFE76642E" box="[376,472,506,527]" pageId="26" pageNumber="919" type="syntype">syntype</typeStatus>
(now
<typeStatus id="548B8852FFE6FFCAFD8B7C99FD7F642E" box="[552,721,505,527]" pageId="26" pageNumber="919" type="paralectotype">paralectotype</typeStatus>
) is labelled
<collectingCountry id="F3277660FFE6FFCAFF5A7F78FEF7640F" box="[249,345,536,558]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
|Takubaya|Bilim. 65. However, the important point is that these
<specimenCount id="9D36FD79FFE6FFCAFE797F57FDD2646D" box="[474,636,566,588]" pageId="26" pageNumber="919" type="generic">two specimens</specimenCount>
are almost certainly not conspecific. The Sichel specimen has the head, mesosoma, petiole and postpetiole densely reticulate-foveolate and opaque, and the gaster finely reticulate and sublucid; the promesonotum is evenly convex in profile and broad in dorsal view (pronotum width/head width ~0.62); the propodeal spine distance is relatively large (SPTD/HW ~0.49); and standing pilosity appears to be sparse (not visibly present on the mesosoma dorsum). The Bilimek specimen is more weakly sculptured with a correspondingly shinier integument, especially on the petiole, postpetiole and gaster; the promesonotum is somewhat flattened in profile and narrower in dorsal view (pronotum width/ head width ~0.53); the propodeal spine tips are less distant (SPTD/HW ~0.44); and standing pilosity is more conspicuous, being present on the mesosoma dorsum on both the pronotal humeri and the mesonotal declivity. Although in this study we are concerned primarily with the
<taxonomicName id="4C304D73FFE6FFCAFF1F791DFEF162B3" box="[188,351,1149,1170]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAE2FFE6FFCAFF1F791DFEF162B3" box="[188,351,1149,1170]" italics="true" pageId="26" pageNumber="919">Crematogaster</emphasis>
</taxonomicName>
of America north of
<collectingCountry id="F3277660FFE6FFCAFDE0791DFD3D62B2" box="[579,659,1149,1171]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
, we have decided to select a
<typeStatus id="548B8852FFE6FFCAFEC079FCFE666293" box="[355,456,1180,1202]" pageId="26" pageNumber="919" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C304D73FFE6FFCAFE4979FCFDE66290" authorityName="Mayr" authorityYear="1870" box="[490,584,1180,1201]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE6FFCAFE4979FCFDE66290" box="[490,584,1180,1201]" italics="true" pageId="26" pageNumber="919">C. opaca</emphasis>
</taxonomicName>
to assist future work. We designate the BMNH specimen as
<typeStatus id="548B8852FFE6FFCAFD3279DAFD5662F1" box="[657,760,1210,1232]" pageId="26" pageNumber="919" type="lectotype">lectotype</typeStatus>
because it better matches the original description, which emphasizes the densely reticulate-punctate sculpture and the scarcity of standing pilosity (
<bibRefCitation id="EFA14B01FFE6FFCAFDC17877FD47630A" author="Mayr G" box="[610,745,1302,1324]" pageId="26" pageNumber="919" pagination="939 - 996" refId="ref36481" refString="Mayr G. 1870. Neue Formiciden. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 20: 939 - 996." type="journal article" year="1870">Mayr, 1870</bibRefCitation>
). Unfortunately the provenance of this specimen other than
<collectingCountry id="F3277660FFE6FFCAFF6C7834FE8E634B" box="[207,288,1364,1386]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
is unknown. Sichel evidently supplied the specimen to Mayr, but there is no evidence that he visited
<collectingCountry id="F3277660FFE6FFCAFF4178F1FE9F6386" box="[226,305,1425,1447]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
or anywhere else outside Europe during his lifetime (
<bibRefCitation id="EFA14B01FFE6FFCAFE8978D0FE7A63E7" author="Mulsant E" box="[298,468,1456,1478]" pageId="26" pageNumber="919" pagination="383 - 410" refId="ref36555" refString="Mulsant E. 1869. Notice sur le Dr. Jules Sichel. Annales de la Societe Linneenne de Lyon 17: 383 - 410." type="journal article" year="1869">Mulsant, 1869</bibRefCitation>
), so the specimen would have been acquired from another collector, possibly de Saussure. The non-conspecific
<typeStatus id="548B8852FFE6FFCAFE5E788DFD336022" box="[509,669,1517,1539]" pageId="26" pageNumber="919" type="paralectotype">paralectotype</typeStatus>
is from Takubaya (= Tacubaya), a district of
<collectingRegion id="49F4F812FFE6FFCAFD907B6CFD176003" box="[563,697,1548,1570]" country="Mexico" name="Distrito Federal" pageId="26" pageNumber="919">Mexico City</collectingRegion>
, near Bilimeks place of residence during his stay in
<collectingCountry id="F3277660FFE6FFCAFD0A7B4AFD576061" box="[681,761,1578,1600]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
(
<bibRefCitation id="EFA14B01FFE6FFCAFF3A7B29FEE7607E" author="Roth PHJ" box="[153,329,1609,1631]" pageId="26" pageNumber="919" pagination="338 - 354" refId="ref36921" refString="Roth PHJ. 1965. Dominik Bilimek Leben und Werk eines osterreichischen Naturforschers: Zur mexikanischen Expedition der Osterreicher vor hundert Jahren. Sudhoffs Archiv fur Geschichte der Medizin und der Naturwissenschaften 49: 338 - 354." type="journal article" year="1965">Roth, 1965: 346</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36F0FFE6FFCAFF0A7B08FB216755" blockId="26.[145,762,290,1907]" lastBlockId="26.[809,1425,197,372]" pageId="26" pageNumber="919">
There is a confusing array of
<taxonomicName id="4C304D73FFE6FFCAFE467B08FDEE605C" authorityName="Mayr" authorityYear="1870" box="[485,576,1640,1661]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE6FFCAFE467B08FDEE605C" box="[485,576,1640,1661]" italics="true" pageId="26" pageNumber="919">C. opaca</emphasis>
</taxonomicName>
-like forms found in
<collectingCountry id="F3277660FFE6FFCAFF0C7BE6FF5160BD" box="[175,255,1670,1692]" name="Mexico" pageId="26" pageNumber="919">Mexico</collectingCountry>
and Central America, showing considerable variation in sculpture, pilosity, configuration of the promesonotum, size of propodeal spines and petiole shape. More comprehensive sampling and careful study is necessary to determine how many species are present in this complex. These ants just barely enter the
<collectingCountry id="F3277660FFE6FFCAFD0F7A40FF706172" name="United States of America" pageId="26" pageNumber="919">United States</collectingCountry>
, in southern
<collectingRegion id="49F4F812FFE6FFCAFE207A5EFE4A6175" box="[387,484,1854,1876]" country="United States of America" name="Arizona" pageId="26" pageNumber="919">Arizona</collectingRegion>
. Because of taxonomic uncertainty, we refer to the
<collectingRegion id="49F4F812FFE6FFCAFE477A3DFDEF6152" box="[484,577,1885,1907]" country="United States of America" name="Arizona" pageId="26" pageNumber="919">Arizona</collectingRegion>
populations as
<taxonomicName id="4C304D73FFE6FFCAFC8A7DA5FC4066FB" box="[809,1006,197,218]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B944EAE2FFE6FFCAFC8A7DA5FC6766FB" box="[809,969,197,218]" italics="true" pageId="26" pageNumber="919">Crematogaster</emphasis>
sp.
</taxonomicName>
<taxonomicName id="4C304D73FFE6FFCAFC517DA5FBE166FA" authorityName="Mayr" authorityYear="1870" box="[1010,1103,197,219]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca">
cf.
<emphasis id="B944EAE2FFE6FFCAFBAC7DA6FBE166FA" box="[1039,1103,198,219]" italics="true" pageId="26" pageNumber="919">opaca</emphasis>
</taxonomicName>
. These specimens are similar to the
<taxonomicName id="4C304D73FFE6FFCAFCD57D84FC7666D8" authorityName="Mayr" authorityYear="1870" box="[886,984,228,249]" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="919" phylum="Arthropoda" rank="species" species="opaca">
<emphasis id="B944EAE2FFE6FFCAFCD57D84FC7666D8" box="[886,984,228,249]" italics="true" pageId="26" pageNumber="919">C. opaca</emphasis>
</taxonomicName>
<typeStatus id="548B8852FFE6FFCAFC427D84FBE466DB" box="[993,1098,228,250]" pageId="26" pageNumber="919" type="lectotype">lectotype</typeStatus>
with respect to integument sculpture, promesonotal shape (pronotum width/head width 0.590.62) and propodeal spine distance (SPTD/ HW 0.450.51), but they have more abundant standing pilosity (MSC 25, A4SC 1018).
</paragraph>
</subSubSection>
</treatment>
</document>