plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/99/87/039987ADFFCFE022D398FD71FB75FD08.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

316 lines
36 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="565AD6A727BA100A584318AFD245AD71" ID-DOI="10.11646/zootaxa.5352.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="8426638" ID-ZooBank="74DCF84E-60ED-49EA-B5E2-A794A60E4D06" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1696942088598" checkinUser="plazi" docAuthor="Matson, Tanner A." docDate="2023" docId="039987ADFFCFE022D398FD71FB75FD08" docLanguage="en" docName="zootaxa.5352.4.7.pdf" docOrigin="Zootaxa 5352 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5352.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achagua velata Matson 2023, n. sp." docType="treatment" docVersion="1" lastPageNumber="575" masterDocId="FFA0FFD5FFC7E028D30FFFA7FF89FFA1" masterDocTitle="The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae)" masterLastPageNumber="576" masterPageNumber="565" pageNumber="573" updateTime="1697115518778" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="C9588DC429F3E68A16F3F5DC61CA7267" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="730ECB94F3F31EA15F54BC34F5C4CE93">
<mods:title id="0D8D6666A3614E1F86D9AAE189D1015E">The known species of the genus Achagua Rindge, 1983, with the description of three new species from the Neotropics (Geometridae: Ennominae)</mods:title>
</mods:titleInfo>
<mods:name id="9121EEBF77F5CA986158CEF63692C9DD" type="personal">
<mods:role id="B0FCFD9B8702357FD8710642745F14CB">
<mods:roleTerm id="265D1D8015273F230898A25F411B8CF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D42B5B9D93A8EA86D63055443B38B32">Matson, Tanner A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2CED4532F4D3AD5272099B80E56A9783">text</mods:typeOfResource>
<mods:relatedItem id="C5C3A8BADFFD8BA9AE36A88E9E18A8D6" type="host">
<mods:titleInfo id="65571EA8DEF7B74E32A577BC5DD1958E">
<mods:title id="909E382ADA22A09C8C2EA1BC13C1B2EC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4BA8EEC299605F528F2444D0CF2475CA">
<mods:date id="E554A2AB75943DEC90353E1EDCA4035C">2023</mods:date>
<mods:detail id="F4DADD2D8C1BF3716B3BAD3B8F8804F8" type="pubDate">
<mods:number id="462950266146DBD19BA04D681B7BBDF7">2023-10-05</mods:number>
</mods:detail>
<mods:detail id="4969B240A842AF1CB5427A1600DB5747" type="volume">
<mods:number id="192FC59F98073024BFFF069F12D0903C">5352</mods:number>
</mods:detail>
<mods:detail id="521D0D02CE120B6C53586B440BA688B3" type="issue">
<mods:number id="C60E71A54128A75FD2D02BE6F91D249D">4</mods:number>
</mods:detail>
<mods:extent id="8C0762502F59F732D6F3C70D87AF9D73" unit="page">
<mods:start id="0D7BC27243F579F9A22FEE2759053248">565</mods:start>
<mods:end id="78B5562C6B3E6FC97BE8421C1F148B5E">576</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="6F6436A7EAFFF6F2EF7651A02A9A5277">
<mods:url id="DF9A4029605233A937E820DE78FFB55D">http://dx.doi.org/10.11646/zootaxa.5352.4.7</mods:url>
</mods:location>
<mods:classification id="B4CF59AD343FDCB094D2145C12BF043E">journal article</mods:classification>
<mods:identifier id="75A2EC04F1FF87CADF6CB348B8F261E2" type="DOI">10.11646/zootaxa.5352.4.7</mods:identifier>
<mods:identifier id="F272DCBDAB0BD91B7ACE5781759687E2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E009E7583E4648B8A51B8C029F9FF559" type="Zenodo-Dep">8426638</mods:identifier>
<mods:identifier id="98F83D88E8DE89FD42D18C9549087AB5" type="ZooBank">74DCF84E-60ED-49EA-B5E2-A794A60E4D06</mods:identifier>
</mods:mods>
<treatment id="039987ADFFCFE022D398FD71FB75FD08" LSID="urn:lsid:plazi:treatment:039987ADFFCFE022D398FD71FB75FD08" httpUri="http://treatment.plazi.org/id/039987ADFFCFE022D398FD71FB75FD08" lastPageId="10" lastPageNumber="575" pageId="8" pageNumber="573">
<subSubSection id="C32A6530FFCFE020D398FD71FD83FD51" box="[151,522,726,753]" pageId="8" pageNumber="573" type="nomenclature">
<paragraph id="8B8F36BBFFCFE020D398FD71FD83FD51" blockId="8.[151,750,726,825]" box="[151,522,726,753]" pageId="8" pageNumber="573">
<heading id="D0C781D7FFCFE020D398FD71FD83FD51" bold="true" box="[151,522,726,753]" fontSize="11" level="1" pageId="8" pageNumber="573" reason="1">
<emphasis id="B944EAA9FFCFE020D398FD71FD83FD51" bold="true" box="[151,522,726,753]" pageId="8" pageNumber="573">
<taxonomicName id="4C304D38FFCFE020D398FD71FE32FD51" authority="Matson" authorityName="Matson" authorityYear="2023" box="[151,443,726,753]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata" status="n. sp.">
<emphasis id="B944EAA9FFCFE020D398FD71FEDDFD50" bold="true" box="[151,340,726,753]" italics="true" pageId="8" pageNumber="573">Achagua velata</emphasis>
Matson
</taxonomicName>
,
<taxonomicNameLabel id="A27757D2FFCFE020D2C7FD71FD83FD51" box="[456,522,726,752]" pageId="8" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCFE020D398FD5DFD67FC98" pageId="8" pageNumber="573" type="description">
<paragraph id="8B8F36BBFFCFE020D398FD5DFEEFFCB5" blockId="8.[151,750,726,825]" box="[151,358,762,788]" pageId="8" pageNumber="573">
(
<figureCitation id="130B2A3EFFCFE020D390FD5DFF66FCB5" box="[159,239,762,788]" captionStart="FIGURES 1–4" captionStartId="3.[152,270,1870,1894]" captionTargetBox="[156,1432,179,1844]" captionTargetId="figure-16@3.[156,1432,179,1844]" captionTargetPageId="3" captionText="FIGURES 1–4. Achagua adult habitus. (a) dorsal view, (b) ventral view. (1) A. obsoleta Rindge, AMNH_IZC 00353009, paratype, (Colombia: Cundinamarca), AMNH; (2) A. magna n. sp., AMNH_IZC 00353021, holotype, (Peru: Cusco), AMNH; (3) A. cooperae n. sp., USNMENT01771270, holotype, (Costa Rica: Alajuela), USNM; (4) A. velata n. sp., holotype, (French Guiana: Régina), MNHN. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.8426640" httpUri="https://zenodo.org/record/8426640/files/figure.png" pageId="8" pageNumber="573">Figs. 4</figureCitation>
,
<figureCitation id="130B2A3EFFCFE020D3F4FD5DFE83FCB5" box="[251,266,762,788]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="8" pageNumber="573">8</figureCitation>
,
<figureCitation id="130B2A3EFFCFE020D218FD5DFEBDFCB5" box="[279,308,762,788]" captionStart="FIGURE 11" captionStartId="9.[152,255,1039,1063]" captionTargetBox="[151,1436,181,1015]" captionTargetId="figure-16@9.[151,1436,181,1015]" captionTargetPageId="9" captionText="FIGURE 11. Geographic distribution of Achagua: A. obsoleta Rindge (star), A. magna n. sp. (triangle), A. cooperae n. sp. (circle), and A. velata n. sp. (square). Single dots may represent&gt;1 individual. Scale bar = 600 km." figureDoi="http://doi.org/10.5281/zenodo.8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="8" pageNumber="573">11</figureCitation>
,
<figureCitation id="130B2A3EFFCFE020D24FFD5DFED4FCB5" box="[320,349,762,788]" captionStart="FIGURE 12" captionStartId="9.[152,255,1750,1774]" captionTargetBox="[172,1413,1136,1720]" captionTargetId="figure-60@9.[162,1426,1124,1726]" captionTargetPageId="9" captionText="FIGURE 12. COI neighbor-joining tree generated from all publicly available Achagua sequences. Taxon names followed by BOLD Process ID and country of origin." figureDoi="http://doi.org/10.5281/zenodo.8426648" httpUri="https://zenodo.org/record/8426648/files/figure.png" pageId="8" pageNumber="573">12</figureCitation>
)
</paragraph>
<paragraph id="8B8F36BBFFCFE020D398FCB9FD67FC98" blockId="8.[151,750,726,825]" box="[151,750,798,825]" pageId="8" pageNumber="573">
LSID:
<uuid id="FF960C6EFFCFE020D3EBFCB9FD67FC98" box="[228,750,798,825]" pageId="8" pageNumber="573">0E4A82F4-6BDF-43BE-86F8-572966809E46</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCFE020D398FCC1FD74FB65" pageId="8" pageNumber="573" type="diagnosis">
<paragraph id="8B8F36BBFFCFE020D398FCC1FD74FB65" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D398FCC1FE9BFC21" bold="true" box="[151,274,870,896]" pageId="8" pageNumber="573">Diagnosis.</emphasis>
The following diagnosis is based on information from a single individual of
<taxonomicName id="4C304D38FFCFE020D752FCC0FB36FC21" authorityName="Matson" authorityYear="2023" box="[1117,1215,870,897]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D752FCC0FB36FC21" box="[1117,1215,870,897]" italics="true" pageId="8" pageNumber="573">A. velata</emphasis>
</taxonomicName>
, and caution should be exercised when interpreting it. As of now,
<taxonomicName id="4C304D38FFCFE020D195FC2DFCC0FC05" authorityName="Matson" authorityYear="2023" box="[666,841,906,932]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D195FC2DFCC0FC05" box="[666,841,906,932]" italics="true" pageId="8" pageNumber="573">Achagua velata</emphasis>
</taxonomicName>
is currently the only member of the genus known to inhabit the Guianan moist forests of northeastern South America. This species bears a conspicuous antemedial black spot along the forewing inner margin and a weakly gray, discal, reniform spot that are absent in both
<taxonomicName id="4C304D38FFCFE020D621FC74FA15FC4D" authorityName="Matson" authorityYear="2023" box="[1326,1436,979,1005]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B944EAA9FFCFE020D621FC74FA15FC4D" box="[1326,1436,979,1005]" italics="true" pageId="8" pageNumber="573">A. magna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D38FFCFE020D3C8FC50FECCFBB1" box="[199,325,1014,1041]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="obsoleta">
<emphasis id="B944EAA9FFCFE020D3C8FC50FECCFBB1" box="[199,325,1014,1041]" italics="true" pageId="8" pageNumber="573">A. obsoleta</emphasis>
</taxonomicName>
. The terminal band on both wings is thicker and more complete than in other congeners. The male genitalia of
<taxonomicName id="4C304D38FFCFE020D211FBBCFE0BFB95" authorityName="Matson" authorityYear="2023" box="[286,386,1050,1077]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D211FBBCFE0BFB95" box="[286,386,1050,1077]" italics="true" pageId="8" pageNumber="573">A. velata</emphasis>
</taxonomicName>
lack a spatulate dorsal process of the uncus, a postmedial digitate protuberance on the costa, and cornuti on the vesica, all of which are found in
<taxonomicName id="4C304D38FFCFE020D1EAFB98FCD8FBF9" authorityName="Matson" authorityYear="2023" box="[741,849,1087,1113]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B944EAA9FFCFE020D1EAFB98FCD8FBF9" box="[741,849,1087,1113]" italics="true" pageId="8" pageNumber="573">A. magna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D38FFCFE020D085FB98FB80FBF9" box="[906,1033,1086,1113]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="obsoleta">
<emphasis id="B944EAA9FFCFE020D085FB98FB80FBF9" box="[906,1033,1086,1113]" italics="true" pageId="8" pageNumber="573">A. obsoleta</emphasis>
</taxonomicName>
. While male
<taxonomicName id="4C304D38FFCFE020D7ACFB98FA8EFBF9" authorityName="Matson" authorityYear="2023" box="[1187,1287,1086,1113]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D7ACFB98FA8EFBF9" box="[1187,1287,1086,1113]" italics="true" pageId="8" pageNumber="573">A. velata</emphasis>
</taxonomicName>
shares many genitalic similarities with
<taxonomicName id="4C304D38FFCFE020D2B2FBC4FDCCFBDD" authorityName="Matson" authorityYear="2023" box="[445,581,1123,1149]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="cooperae">
<emphasis id="B944EAA9FFCFE020D2B2FBC4FDCCFBDD" box="[445,581,1123,1149]" italics="true" pageId="8" pageNumber="573">A. cooperae</emphasis>
</taxonomicName>
, the dorsal process of the uncus appears knob-like in
<taxonomicName id="4C304D38FFCFE020D7ABFBC4FAA5FBDD" authorityName="Matson" authorityYear="2023" box="[1188,1324,1123,1149]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="cooperae">
<emphasis id="B944EAA9FFCFE020D7ABFBC4FAA5FBDD" box="[1188,1324,1123,1149]" italics="true" pageId="8" pageNumber="573">A. cooperae</emphasis>
</taxonomicName>
(
<figureCitation id="130B2A3EFFCFE020D633FBC5FA07FBDD" box="[1340,1422,1122,1148]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="8" pageNumber="573">Fig. 7a</figureCitation>
), whereas in
<taxonomicName id="4C304D38FFCFE020D218FB20FEF2FB01" authorityName="Matson" authorityYear="2023" box="[279,379,1158,1185]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D218FB20FEF2FB01" box="[279,379,1158,1185]" italics="true" pageId="8" pageNumber="573">A. velata</emphasis>
</taxonomicName>
(
<figureCitation id="130B2A3EFFCFE020D284FB21FE55FB01" box="[395,476,1158,1184]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="8" pageNumber="573">Fig. 8a</figureCitation>
), it is more uniform in shape.
<taxonomicName id="4C304D38FFCFE020D03FFB21FC56FB01" authorityName="Matson" authorityYear="2023" box="[816,991,1158,1184]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCFE020D03FFB21FC56FB01" box="[816,991,1158,1184]" italics="true" pageId="8" pageNumber="573">Achagua velata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D38FFCFE020D719FB20FB14FB01" authorityName="Matson" authorityYear="2023" box="[1046,1181,1159,1185]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="573" phylum="Arthropoda" rank="species" species="cooperae">
<emphasis id="B944EAA9FFCFE020D719FB20FB14FB01" box="[1046,1181,1159,1185]" italics="true" pageId="8" pageNumber="573">A. cooperae</emphasis>
</taxonomicName>
may also be separated by their COI barcode (see Molecular characterization).
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCFE020D3C8FB69FE4AF835" pageId="8" pageNumber="573" type="description">
<paragraph id="8B8F36BBFFCFE020D3C8FB69FCB8FB49" blockId="8.[151,1437,870,1940]" box="[199,817,1230,1257]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8FB69FE32FB49" bold="true" box="[199,443,1230,1256]" pageId="8" pageNumber="573">Description. MALE.</emphasis>
Forewing length,
<quantity id="4CC89B5EFFCFE020D187FB69FD5EFB48" box="[648,727,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="573" unit="mm" value="17.0">17 mm</quantity>
(n = 1).
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8FB55FB3DFAF4" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8FB55FE87FAAD" bold="true" box="[199,270,1266,1292]" pageId="8" pageNumber="573">Head.</emphasis>
Antenna mostly bipectinate, but with gradually diminishing rami that are absent in distal third of antenna; scales above white and light gray, rami dark gray to black. Vertex white; frons mostly light gray. Labial palpus short, decumbent, 1.5x diameter of eye, light gray and white. Chaetosemata in transverse row.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8FAF9FEADFA61" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8FAF9FEAEFAD9" bold="true" box="[199,295,1374,1400]" pageId="8" pageNumber="573">Thorax.</emphasis>
Patagium, tegula, and mesothorax admixture of gray and white scales. Legs mostly white and mottled with gray; epiphysis well-developed; hind tibia with hair pencil (not easily visualized in
<typeStatus id="548B8819FFCFE020D770FA25FB6FFA3D" box="[1151,1254,1410,1436]" pageId="8" pageNumber="573">holotype</typeStatus>
); tibial spur formula 0–2–4.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8FA6DFBF1F9F1" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8FA6DFEB6FA45" bold="true" box="[199,319,1482,1508]" pageId="8" pageNumber="573">Forewing.</emphasis>
Pearly-white; widely scattered with inconspicuous light gray scales. Basal and costal areas lightly maculated with gray to brown scales. Subtle, light gray, transverse antemedial line, and weakly gray, discal, reniform spot. Inner margin with antemedial black spot. Terminal area broadly maculated with brown scales. Underside patterned as in upperside, but darkened areas more diffuse and given more toward black.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8F9FDFF79F939" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8F9FDFECCF9D5" bold="true" box="[199,325,1626,1652]" pageId="8" pageNumber="573">Hindwing.</emphasis>
Pearly-white except for complete, broadly dark brown outer margin. Underside patterned as in upperside.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8F905FB56F91C" blockId="8.[151,1437,870,1940]" box="[199,1247,1698,1725]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8F905FECAF91D" bold="true" box="[199,323,1698,1724]" pageId="8" pageNumber="573">Abdomen.</emphasis>
Admixture of gray and white. Third sternite of male abdomen with comb of setae.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8F961FCA1F8D1" blockId="8.[151,1437,870,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8F961FEBCF941" bold="true" box="[199,309,1734,1760]" pageId="8" pageNumber="573">Genitalia</emphasis>
(
<figureCitation id="130B2A3EFFCFE020D249F961FE02F941" box="[326,395,1734,1760]" captionStart="FIGURES 5–8" captionStartId="4.[152,270,1869,1893]" captionTargetBox="[158,1432,190,1832]" captionTargetId="figure-25@4.[151,1436,181,1845]" captionTargetPageId="4" captionText="FIGURES 5–8. Achagua male genitalia. (a) genital capsule, (b) phallus. (5) A. obsoleta Rindge, TAM-2023-270, AMNH_IZC 00353009, paratype, AMNH; (6) A. magna n. sp., TAM-2023-268, AMNH_IZC 00353023, paratype. AMNH; (7) A. cooperae n. sp., TAM-2023-251,AMNH_IZC 00353028, paratype, AMNH; (8) A. velata n. sp., TAM-2023-289, holotype, MNHN. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8426642" httpUri="https://zenodo.org/record/8426642/files/figure.png" pageId="8" pageNumber="573">Fig. 8</figureCitation>
). Uncus abruptly widened at base; dorsal and ventral processes thumb-like. Base of gnathos subquadrangular, with upcurved, heavily sclerotized, pointed apical projection; projection lightly papillated. Valve elongate and large with heavily sclerotized costa; apex strongly falcate. Anellar processes large and triangulate, directed inward, and with apical recurved hooks. Juxta with medial, long cylindrical process with acuminate apex. Vesica with small medial sclerotized patch; cornuti absent.
</paragraph>
<paragraph id="8B8F36BBFFCFE020D3C8F8DDFE4AF835" blockId="8.[151,1437,870,1940]" box="[199,451,1914,1940]" pageId="8" pageNumber="573">
<emphasis id="B944EAA9FFCFE020D3C8F8DDFECFF835" bold="true" box="[199,326,1914,1940]" pageId="8" pageNumber="573">FEMALE.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF4F6633FFCEE021D397FBA8FB0CFBEA" ID-DOI="http://doi.org/10.5281/zenodo.8426646" ID-Zenodo-Dep="8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="9" pageNumber="574" startId="9.[152,255,1039,1063]" targetBox="[151,1436,181,1015]" targetPageId="9" targetType="figure">
<paragraph id="8B8F36BBFFCEE021D397FBA8FB0CFBEA" blockId="9.[151,1436,1039,1100]" pageId="9" pageNumber="574">
<emphasis id="B944EAA9FFCEE021D397FBA8FEAFFB86" bold="true" box="[152,294,1039,1063]" pageId="9" pageNumber="574">FIGURE 11.</emphasis>
Geographic distribution of
<taxonomicName id="4C304D38FFCEE021D148FBA8FD28FB86" authorityName="Rindge" authorityYear="1983" box="[583,673,1039,1063]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA9FFCEE021D148FBA8FD28FB86" box="[583,673,1039,1063]" italics="true" pageId="9" pageNumber="574">Achagua</emphasis>
</taxonomicName>
:
<taxonomicName id="4C304D38FFCEE021D1BFFBB7FCFCFB86" authority="Rindge" authorityName="Rindge" box="[688,885,1039,1063]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="species" species="obsoleta">
<emphasis id="B944EAA9FFCEE021D1BFFBB7FCADFB86" box="[688,804,1039,1063]" italics="true" pageId="9" pageNumber="574">A. obsoleta</emphasis>
Rindge
</taxonomicName>
(star),
<taxonomicName id="4C304D38FFCEE021D0CEFBB7FBACFB86" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[961,1061,1040,1063]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="species" species="magna" status="n. sp.">
<emphasis id="B944EAA9FFCEE021D0CEFBB7FBACFB86" box="[961,1061,1040,1063]" italics="true" pageId="9" pageNumber="574">A. magna</emphasis>
</taxonomicName>
<emphasis id="B944EAA9FFCEE021D722FBA8FBE1FB86" bold="true" box="[1069,1128,1039,1063]" pageId="9" pageNumber="574">
<taxonomicNameLabel id="A27757D2FFCEE021D722FBA8FBE1FB86" box="[1069,1128,1039,1063]" pageId="9" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(triangle),
<taxonomicName id="4C304D38FFCEE021D7D2FBB7FAD0FB86" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[1245,1369,1040,1063]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="species" species="cooperae" status="n. sp.">
<emphasis id="B944EAA9FFCEE021D7D2FBB7FAD0FB86" box="[1245,1369,1040,1063]" italics="true" pageId="9" pageNumber="574">A. cooperae</emphasis>
</taxonomicName>
<emphasis id="B944EAA9FFCEE021D66EFBA8FA15FB86" bold="true" box="[1377,1436,1039,1063]" pageId="9" pageNumber="574">
<taxonomicNameLabel id="A27757D2FFCEE021D66EFBA8FA15FB86" box="[1377,1436,1039,1063]" pageId="9" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(circle), and
<taxonomicName id="4C304D38FFCEE021D217FB93FEFBFBEA" authority="Matson, 2023" authorityName="Matson" authorityYear="2023" box="[280,370,1075,1099]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="species" species="velata" status="n. sp.">
<emphasis id="B944EAA9FFCEE021D217FB93FEFBFBEA" box="[280,370,1075,1099]" italics="true" pageId="9" pageNumber="574">A. velata</emphasis>
</taxonomicName>
<emphasis id="B944EAA9FFCEE021D277FB94FE38FBEA" bold="true" box="[376,433,1075,1099]" pageId="9" pageNumber="574">
<taxonomicNameLabel id="A27757D2FFCEE021D277FB94FE38FBEA" box="[376,433,1075,1099]" pageId="9" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(square). Single dots may represent&gt;1 individual. Scale bar = 600 km.
</paragraph>
</caption>
<caption id="DF4F6633FFCEE021D397F971FDBCF8B3" ID-DOI="http://doi.org/10.5281/zenodo.8426648" ID-Zenodo-Dep="8426648" httpUri="https://zenodo.org/record/8426648/files/figure.png" pageId="9" pageNumber="574" startId="9.[152,255,1750,1774]" targetBox="[172,1413,1136,1720]" targetPageId="9" targetType="figure">
<paragraph id="8B8F36BBFFCEE021D397F971FDBCF8B3" blockId="9.[151,1436,1750,1811]" pageId="9" pageNumber="574">
<emphasis id="B944EAA9FFCEE021D397F971FEAFF94F" bold="true" box="[152,294,1750,1774]" pageId="9" pageNumber="574">FIGURE 12.</emphasis>
COI neighbor-joining tree generated from all publicly available
<taxonomicName id="4C304D38FFCEE021D0B1F971FB91F94F" authorityName="Rindge" authorityYear="1983" box="[958,1048,1750,1774]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="574" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA9FFCEE021D0B1F971FB91F94F" box="[958,1048,1750,1774]" italics="true" pageId="9" pageNumber="574">Achagua</emphasis>
</taxonomicName>
sequences. Taxon names followed by BOLD Process ID and country of origin.
</paragraph>
</caption>
<subSubSection id="C32A6530FFCDE022D398FF30FAE7FEE0" pageId="10" pageNumber="575" type="materials_examined">
<paragraph id="8B8F36BBFFCDE022D398FF30FEC1FF11" blockId="10.[151,328,150,177]" box="[151,328,150,177]" pageId="10" pageNumber="575">
<heading id="D0C781D7FFCDE022D398FF30FEC1FF11" bold="true" box="[151,328,150,177]" fontSize="11" level="1" pageId="10" pageNumber="575" reason="1">
<emphasis id="B944EAA9FFCDE022D398FF30FEC1FF11" bold="true" box="[151,328,150,177]" pageId="10" pageNumber="575">
<typeStatus id="548B8819FFCDE022D398FF30FF5BFF10" box="[151,210,151,177]" pageId="10" pageNumber="575">Type</typeStatus>
Material.
</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CE6FFCDE022D398FF78FAE7FEE0" collectingDate="2010-07-09" collectionCode="MNHN" country="French Guiana" elevation="419" location="Nouragues Nature Reserve" municipality="Regina" pageId="10" pageNumber="575" specimenCode="LNOUD2050" specimenCount="1" typeStatus="holotype">
<paragraph id="8B8F36BBFFCDE022D398FF78FE8DFF58" blockId="10.[151,1437,223,321]" box="[151,260,223,249]" pageId="10" pageNumber="575">
<heading id="D0C781D7FFCDE022D398FF78FE8DFF58" bold="true" box="[151,260,223,249]" fontSize="11" level="1" pageId="10" pageNumber="575" reason="1">
<emphasis id="B944EAA9FFCDE022D398FF78FE8DFF58" bold="true" box="[151,260,223,249]" pageId="10" pageNumber="575">
<typeStatus id="548B8819FFCDE022D398FF78FE8DFF58" box="[151,260,223,249]" pageId="10" pageNumber="575">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36BBFFCDE022D398FEA4FAE7FEE0" blockId="10.[151,1437,223,321]" pageId="10" pageNumber="575">
<collectingCountry id="F327762BFFCDE022D398FEA4FEF2FEBC" box="[151,379,259,285]" name="French Guiana" pageId="10" pageNumber="575">FRENCH GUIANA</collectingCountry>
[
<collectingCountry id="F327762BFFCDE022D286FEA4FE77FEBC" box="[393,510,259,285]" name="France" pageId="10" pageNumber="575">FRANCE</collectingCountry>
] • ♁;
<taxonomicName id="4C304D38FFCDE022D14DFEA4FC93FEBC" authority=", Nouragues" authorityName="Nouragues" box="[578,794,259,285]" class="Trilobita" family="Ptychopariidae" genus="Regina" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="575" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="6BEBACC1FFCDE022D14DFEA4FD1DFEBC" box="[578,660,259,285]" pageId="10" pageNumber="575">Régina</collectingMunicipality>
,
<location id="8EEF6060FFCDE022D1AEFEA4FC5DFEBC" LSID="urn:lsid:plazi:treatment:039987ADFFCFE022D398FD71FB75FD08:8EEF6060FFCDE022D1AEFEA4FC5DFEBC" box="[673,980,259,285]" country="French Guiana" municipality="Regina" name="Nouragues Nature Reserve" pageId="10" pageNumber="575">Nouragues Nature Reserve</location>
</taxonomicName>
; (4.096°, -52.683°); elev.
<quantity id="4CC89B5EFFCDE022D7F6FEA4FACDFEBC" box="[1273,1348,259,285]" metricMagnitude="2" metricUnit="m" metricValue="4.1899999999999995" pageId="10" pageNumber="575" unit="m" value="419.0">
<elevation id="001DD188FFCDE022D7F6FEA4FACDFEBC" box="[1273,1348,259,285]" metricMagnitude="2" metricUnit="m" metricValue="4.1899999999999995" pageId="10" pageNumber="575" unit="m" value="419.0">419 m</elevation>
</quantity>
;
<date id="FF8E107BFFCDE022D65EFEA4FF5AFEE0" pageId="10" pageNumber="575" value="2010-07-09">
<collectingDate id="EFCAE993FFCDE022D65EFEA4FF5AFEE0" pageId="10" pageNumber="575" value="2010-07-09">09 Jul. 2010</collectingDate>
</date>
;
<location id="8EEF6060FFCDE022D3D1FE80FE7AFEE0" LSID="urn:lsid:plazi:treatment:039987ADFFCFE022D398FD71FB75FD08:8EEF6060FFCDE022D3D1FE80FE7AFEE0" box="[222,499,295,321]" country="French Guiana" municipality="Regina" name="Carlos Lopez-Vaamonde" pageId="10" pageNumber="575">Carlos Lopez-Vaamonde</location>
leg.; Genitalia: TAM-2023-289; BOLD Process ID:
<specimenCode id="DB969EC0FFCDE022D74CFE81FB6DFEE0" box="[1091,1252,294,321]" pageId="10" pageNumber="575">LNOUD2050</specimenCode>
-12;
<collectionCode id="ED21AE7EFFCDE022D61EFE80FAE3FEE0" box="[1297,1386,295,321]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="10" pageNumber="575">MNHN</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C32A6530FFCDE022D3C8FEC9FEE8FE0C" pageId="10" pageNumber="575" type="distribution">
<paragraph id="8B8F36BBFFCDE022D3C8FEC9FEE8FE0C" blockId="10.[151,1437,366,681]" pageId="10" pageNumber="575">
<emphasis id="B944EAA9FFCDE022D3C8FEC9FED0FE29" bold="true" box="[199,345,366,392]" pageId="10" pageNumber="575">Distribution</emphasis>
(
<figureCitation id="130B2A3EFFCDE022D265FEC8FE32FE28" box="[362,443,367,393]" captionStart="FIGURE 11" captionStartId="9.[152,255,1039,1063]" captionTargetBox="[151,1436,181,1015]" captionTargetId="figure-16@9.[151,1436,181,1015]" captionTargetPageId="9" captionText="FIGURE 11. Geographic distribution of Achagua: A. obsoleta Rindge (star), A. magna n. sp. (triangle), A. cooperae n. sp. (circle), and A. velata n. sp. (square). Single dots may represent&gt;1 individual. Scale bar = 600 km." figureDoi="http://doi.org/10.5281/zenodo.8426646" httpUri="https://zenodo.org/record/8426646/files/figure.png" pageId="10" pageNumber="575">Fig. 11</figureCitation>
). Presently,
<taxonomicName id="4C304D38FFCDE022D146FEC8FD24FE28" authorityName="Matson" authorityYear="2023" box="[585,685,367,393]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="575" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCDE022D146FEC8FD24FE28" box="[585,685,367,393]" italics="true" pageId="10" pageNumber="575">A. velata</emphasis>
</taxonomicName>
is only known from the Guianan moist forests at the
<typeStatus id="548B8819FFCDE022D61FFEC8FAC9FE28" box="[1296,1344,367,393]" pageId="10" pageNumber="575">type</typeStatus>
locality in
<collectingCountry id="F327762BFFCDE022D3BBFE34FED7FE0C" box="[180,350,403,429]" name="French Guiana" pageId="10" pageNumber="575">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCDE022D3C8FE11FF78FE54" pageId="10" pageNumber="575" type="biology_ecology">
<paragraph id="8B8F36BBFFCDE022D3C8FE11FF78FE54" blockId="10.[151,1437,366,681]" pageId="10" pageNumber="575">
<emphasis id="B944EAA9FFCDE022D3C8FE11FEAEFE71" bold="true" box="[199,295,438,464]" pageId="10" pageNumber="575">Biology.</emphasis>
Immature stages and host associations remain unknown. Adults are known to fly in July at the
<typeStatus id="548B8819FFCDE022D663FE10FA15FE70" box="[1388,1436,439,465]" pageId="10" pageNumber="575">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCDE022D3C8FE58FDCEFDC0" pageId="10" pageNumber="575" type="etymology">
<paragraph id="8B8F36BBFFCDE022D3C8FE58FDCEFDC0" blockId="10.[151,1437,366,681]" pageId="10" pageNumber="575">
<emphasis id="B944EAA9FFCDE022D3C8FE58FEC7FDB8" bold="true" box="[199,334,511,537]" pageId="10" pageNumber="575">Etymology.</emphasis>
The specific epithet
<emphasis id="B944EAA9FFCDE022D132FE58FD08FDB8" box="[573,641,511,537]" italics="true" pageId="10" pageNumber="575">velata</emphasis>
is derived from the Latin “velatus,” meaning “veiled.” The name was chosen because of the rarely seen nature of this species—thus far only known from a single individual—and for its white, wedding-veil-like ground color.
</paragraph>
</subSubSection>
<subSubSection id="C32A6530FFCDE022D3C8FDCCFB75FD08" pageId="10" pageNumber="575" type="description">
<paragraph id="8B8F36BBFFCDE022D3C8FDCCFB75FD08" blockId="10.[151,1437,366,681]" pageId="10" pageNumber="575">
<emphasis id="B944EAA9FFCDE022D3C8FDCCFD9CFD25" bold="true" box="[199,533,618,645]" pageId="10" pageNumber="575">Molecular characterization.</emphasis>
<taxonomicName id="4C304D38FFCDE022D113FDCCFD45FD24" authorityName="Matson" authorityYear="2023" box="[540,716,619,645]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="575" phylum="Arthropoda" rank="species" species="velata">
<emphasis id="B944EAA9FFCDE022D113FDCCFD45FD24" box="[540,716,619,645]" italics="true" pageId="10" pageNumber="575">Achagua velata</emphasis>
</taxonomicName>
is represented in BOLD by the BIN: BOLD:ABV2454 (n = 1). The pairwise distance to the nearest neighbor,
<taxonomicName id="4C304D38FFCDE022D190FD28FCFBFD09" authorityName="Matson" authorityYear="2023" box="[671,882,655,681]" class="Insecta" family="Geometridae" genus="Achagua" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="575" phylum="Arthropoda" rank="species" species="cooperae">
<emphasis id="B944EAA9FFCDE022D190FD28FCFBFD09" box="[671,882,655,681]" italics="true" pageId="10" pageNumber="575">Achagua cooperae</emphasis>
</taxonomicName>
(n = 5,
<collectingCountry id="F327762BFFCDE022D0C5FD28FBC1FD08" box="[970,1096,655,681]" name="Costa Rica" pageId="10" pageNumber="575">Costa Rica</collectingCountry>
), is about 3.8%.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink