294 lines
36 KiB
XML
294 lines
36 KiB
XML
<document id="457C3CF321FD0270C698E20B4C1E5311" ID-DOI="10.1016/j.jcz.2022.11.003" ID-ISSN="1873-2674" ID-Zenodo-Dep="10375255" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1702544566555" checkinUser="felipe" docAuthor="Nakahama, Naoyuki, Yamasaki, Takeshi, Komazawa, Masaki & Nakano, Takafumi" docDate="2023" docId="039987A7FFF0FFA3FCB0FA64FBC360F5" docLanguage="en" docName="ZoolAnz.302.17-27.pdf" docOrigin="Zoologischer Anzeiger (Zool. Anz.) 302" docSource="http://dx.doi.org/10.1016/j.jcz.2022.11.003" docStyle="DocumentStyle:32483D6D135DE2FF927BA3F1400E0FFD.1:ZoolAnz.2022-.journal_article" docStyleId="32483D6D135DE2FF927BA3F1400E0FFD" docStyleName="ZoolAnz.2022-.journal_article" docStyleVersion="1" docTitle="Arctigalloisiana Nakahama & Yamasaki & Komazawa & Nakano 2023, gen. nov." docType="treatment" docVersion="1" lastPageNumber="22" masterDocId="FFA0FFDFFFF3FFA6FF82FF9DFFA5651C" masterDocTitle="Integrative approach clarifies the distinct taxonomic account of gryloblattids endemic to Hokkaido, Japan, with a description of two new species (Insecta, Grylloblattodea)" masterLastPageNumber="27" masterPageNumber="17" pageNumber="20" updateTime="1730164259555" updateUser="diego">
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<mods:title id="9DC898EBCD5250319D423E3D89622A21">Integrative approach clarifies the distinct taxonomic account of gryloblattids endemic to Hokkaido, Japan, with a description of two new species (Insecta, Grylloblattodea)</mods:title>
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<mods:namePart id="8FC52CCE18E9CDB22485B8CDC48B890A">Nakahama, Naoyuki</mods:namePart>
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<mods:affiliation id="3E16CE2C8AA78F715820409AEE2B01F2">*, &, Masaki Komazawa & Institute of Natural and Environmental Sciences, University of Hyogo, 6 Yayoigaoka, Sanda City, Hyogo, 669 - 1546, Japan & *, &, Masaki Komazawa & The Museum of Nature and Human Activities, Hyogo, 6 Yayoigaoka, Sanda City, Hyogo, 669 - 1546, Japan &, Takeshi Yamasaki &, Masaki Komazawa</mods:affiliation>
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<mods:roleTerm id="CC60B23E9E65153AD488A7E359F23F5F">Author</mods:roleTerm>
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<mods:namePart id="28278BDAB79B7BF48B2555A247B99441">Yamasaki, Takeshi</mods:namePart>
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<mods:affiliation id="31BB8C17F52740ED44F2CE2CE27076F9">*, &, Masaki Komazawa & Institute of Natural and Environmental Sciences, University of Hyogo, 6 Yayoigaoka, Sanda City, Hyogo, 669 - 1546, Japan & *, &, Masaki Komazawa & The Museum of Nature and Human Activities, Hyogo, 6 Yayoigaoka, Sanda City, Hyogo, 669 - 1546, Japan &, Takeshi Yamasaki &, Masaki Komazawa</mods:affiliation>
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<mods:roleTerm id="EDCB875734D7F4A971B416D8C7054B46">Author</mods:roleTerm>
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<mods:namePart id="1F432D78351FB3E3C3CB0A40E944C4A0">Komazawa, Masaki</mods:namePart>
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<mods:affiliation id="54271ACD5F1FC2619C49465003986688">, Takafumi Nakano & Nishiki-mati, Asahikawa-shi, Hokkaido, Japan</mods:affiliation>
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<mods:namePart id="92501A534DB701FB9B95D69ED63B871E">Nakano, Takafumi</mods:namePart>
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<mods:affiliation id="025ED6EEFE296FF1BFAA8990103C5F64">Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawa-oiwakecho, Sakyo, Kyoto, 606 - 8502, Japan</mods:affiliation>
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<mods:typeOfResource id="40D124CA200670EC490DD50A86DF0176">text</mods:typeOfResource>
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<mods:title id="63E59FEEFD16109AF0B9EC016DF853FD">Zoologischer Anzeiger</mods:title>
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<mods:part id="57BDC3DA1F2931AF8EBFCDD4468FA19B">
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<mods:date id="1062A8F60862E33E7437D49554861630">2023</mods:date>
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<mods:title id="9E61B75A381D649D77555EF0726A4512">Zool. Anz.</mods:title>
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<mods:number id="B4E18F6C625B399BBDF3EA72A81CB68B">2023-01-31</mods:number>
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<mods:number id="541933CAC1CEEF45E5D7C4E2555EEDFF">302</mods:number>
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<mods:url id="3A8FC8957762A44C02D548BF7818B53F">http://dx.doi.org/10.1016/j.jcz.2022.11.003</mods:url>
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<mods:classification id="ED877736B295B43648079A08BD5FFE12">journal article</mods:classification>
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<mods:identifier id="483217CB7E63D635B9DB74549A3D75B0" type="DOI">10.1016/j.jcz.2022.11.003</mods:identifier>
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<mods:identifier id="01326A0A0794A6583A094C5D22C43932" type="ISSN">1873-2674</mods:identifier>
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<treatment id="039987A7FFF0FFA3FCB0FA64FBC360F5" LSID="urn:lsid:plazi:treatment:039987A7FFF0FFA3FCB0FA64FBC360F5" httpUri="http://treatment.plazi.org/id/039987A7FFF0FFA3FCB0FA64FBC360F5" lastPageId="5" lastPageNumber="22" pageId="3" pageNumber="20">
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<subSubSection id="C32A653AFFF0FFA5FCB0FA64FBF26311" box="[818,1111,1529,1549]" pageId="3" pageNumber="20" type="nomenclature">
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<paragraph id="8B8F36B1FFF0FFA5FCB0FA64FBF26311" blockId="3.[818,1488,1529,1967]" box="[818,1111,1529,1549]" pageId="3" pageNumber="20">
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<emphasis id="B944EAA3FFF0FFA5FCB0FA64FBF26311" bold="true" box="[818,1111,1529,1549]" italics="true" pageId="3" pageNumber="20">
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3.3.1.
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<taxonomicName id="4C304D32FFF0FFA5FCF6FA64FC5A6310" ID-CoL="BXLMK" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[884,1023,1529,1548]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">Arctigalloisiana</taxonomicName>
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<taxonomicNameLabel id="A27757D8FFF0FFA5FB84FA67FBF26311" box="[1030,1111,1530,1549]" pageId="3" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C32A653AFFF0FFA5FCD3F988FC626358" pageId="3" pageNumber="20" type="description">
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<paragraph id="8B8F36B1FFF0FFA5FCD3F988FC626358" blockId="3.[818,1488,1529,1967]" pageId="3" pageNumber="20">
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<uri id="FFA13AB3FFF0FFA5FCD3F988FC626358" pageId="3" pageNumber="20">
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urn:lsid:zoobank.org:act:
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<uuid id="FF960C64FFF0FFA5FBB3F988FC626358" pageId="3" pageNumber="20">E0C52FF1-641A-4544-B7FB-939D13AF72C1</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A653AFFF0FFA5FCD3F9D0FA88637C" box="[849,1325,1613,1632]" pageId="3" pageNumber="20" type="type_taxon">
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<paragraph id="8B8F36B1FFF0FFA5FCD3F9D0FA88637C" blockId="3.[818,1488,1529,1967]" box="[849,1325,1613,1632]" pageId="3" pageNumber="20">
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<emphasis id="B944EAA3FFF0FFA5FCD3F9D0FC74637C" bold="true" box="[849,977,1613,1632]" pageId="3" pageNumber="20">
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<typeStatus id="548B8813FFF0FFA5FCD3F9D0FC27637C" box="[849,898,1613,1632]" pageId="3" pageNumber="20">Type</typeStatus>
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species
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||
</emphasis>
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.
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<taxonomicName id="4C304D32FFF0FFA5FC5FF9D0FA8D637C" authority="Asahina, 1961" authorityName="Asahina" authorityYear="1961" box="[989,1320,1613,1632]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="yezoensis">
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<emphasis id="B944EAA3FFF0FFA5FC5FF9D0FB3F637C" bold="true" box="[989,1178,1613,1632]" italics="true" pageId="3" pageNumber="20">Galloisiana yezoensis</emphasis>
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<bibRefCitation id="EFA14B40FFF0FFA5FB22F9D0FA8D637C" author="Asahina, S." box="[1184,1320,1613,1632]" pageId="3" pageNumber="20" pagination="85 - 87" refId="ref7106" refString="Asahina, S., 1961. A new Galloisiana from Hokkaido (Grylloblattoidea). Kontyu 29, 85 - 87." type="journal article" year="1961">Asahina, 1961</bibRefCitation>
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</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32A653AFFF0FFA5FCD3F9F4FC4463F0" pageId="3" pageNumber="20" type="diagnosis">
|
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<paragraph id="8B8F36B1FFF0FFA5FCD3F9F4FC4463F0" blockId="3.[818,1488,1529,1967]" pageId="3" pageNumber="20">
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||
<emphasis id="B944EAA3FFF0FFA5FCD3F9F4FC166360" bold="true" box="[849,947,1641,1660]" pageId="3" pageNumber="20">Diagnosis</emphasis>
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.
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<taxonomicName id="4C304D32FFF0FFA5FC3DF9F4FBEF6360" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[959,1098,1641,1660]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B944EAA3FFF0FFA5FC3DF9F4FBEF6360" bold="true" box="[959,1098,1641,1660]" italics="true" pageId="3" pageNumber="20">Arctigalloisiana</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF0FFA5FBD2F9F4FB0D6360" box="[1104,1192,1641,1660]" pageId="3" pageNumber="20" rank="genus">gen. nov.</taxonomicNameLabel>
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||
is very close to
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<taxonomicName id="4C304D32FFF0FFA5FAC2F9F4FA016360" authorityName="Caudell and King" authorityYear="1924" box="[1344,1444,1641,1660]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA3FFF0FFA5FAC2F9F4FA016360" bold="true" box="[1344,1444,1641,1660]" italics="true" pageId="3" pageNumber="20">Galloisiana</emphasis>
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||
</taxonomicName>
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, but can be distinguished from the latter by a macrotrichia-less margin of the cervical sclerite (
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<figureCitation id="130B2A34FFF0FFA5FC5BF93CFB8F63A8" box="[985,1066,1697,1716]" captionStart="Fig" captionStartId="3.[100,130,1390,1407]" captionTargetBox="[340,1247,149,1360]" captionTargetId="figure-361@3.[338,1248,148,1363]" captionTargetPageId="3" captionText="Fig. 4. Arctigalloisiana yezoensis, female (B2-407738). (A) Head and Pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) tarsal pulvillus of anterior leg, ventral view. (G) ovipositor, dorsal view. (H) same, lateral view. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10375267" httpUri="https://zenodo.org/record/10375267/files/figure.png" pageId="3" pageNumber="20">Figs. 4D</figureCitation>
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,
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<figureCitation id="130B2A34FFF0FFA5FBB5F93CFBF763A8" box="[1079,1106,1697,1716]" captionStart="Fig" captionStartId="4.[100,130,1086,1103]" captionTargetBox="[340,1247,149,1056]" captionTargetId="figure-61@4.[338,1248,148,1058]" captionTargetPageId="4" captionText="Fig. 5. Arctigalloisiana yezoensis, male (B2-407735). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. Scales: A–D, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375269" httpUri="https://zenodo.org/record/10375269/files/figure.png" pageId="3" pageNumber="20">5D</figureCitation>
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and
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<figureCitation id="130B2A34FFF0FFA5FB05F93CFB0163A8" box="[1159,1188,1697,1716]" captionStart="Fig" captionStartId="5.[100,130,1382,1399]" captionTargetBox="[340,1247,149,1354]" captionTargetId="figure-292@5.[338,1248,148,1355]" captionTargetPageId="5" captionText="Fig. 7. Arctigalloisiana poropnetopa sp. nov., holotype female (B2-407746). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) ovipositor, dorsal view. (G) same, lateral view. Scales: A–D, F–G, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375276" httpUri="https://zenodo.org/record/10375276/files/figure.png" pageId="3" pageNumber="20">7D</figureCitation>
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,
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<figureCitation id="130B2A34FFF0FFA5FB33F93CFB6A63A8" box="[1201,1231,1697,1716]" captionStart="Fig" captionStartId="6.[100,130,1383,1400]" captionTargetBox="[340,1247,150,1355]" captionTargetId="figure-376@6.[338,1248,148,1356]" captionTargetPageId="6" captionText="Fig. 8. Arctigalloisiana yubariensis sp. nov., holotype female (B2-407748). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) ovipositor, dorsal view. (G) same, lateral view. Scales: A–D, F–G, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375280" httpUri="https://zenodo.org/record/10375280/files/figure.png" pageId="3" pageNumber="20">8D</figureCitation>
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), one pair of setae on the postmentum (
|
||
<figureCitation id="130B2A34FFF0FFA5FC3AF920FBAC63CC" box="[952,1033,1725,1744]" captionStart="Fig" captionStartId="3.[100,130,1390,1407]" captionTargetBox="[340,1247,149,1360]" captionTargetId="figure-361@3.[338,1248,148,1363]" captionTargetPageId="3" captionText="Fig. 4. Arctigalloisiana yezoensis, female (B2-407738). (A) Head and Pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) tarsal pulvillus of anterior leg, ventral view. (G) ovipositor, dorsal view. (H) same, lateral view. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10375267" httpUri="https://zenodo.org/record/10375267/files/figure.png" pageId="3" pageNumber="20">Figs. 4C</figureCitation>
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,
|
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<figureCitation id="130B2A34FFF0FFA5FB94F920FB8A63CC" box="[1046,1071,1725,1744]" captionStart="Fig" captionStartId="4.[100,130,1086,1103]" captionTargetBox="[340,1247,149,1056]" captionTargetId="figure-61@4.[338,1248,148,1058]" captionTargetPageId="4" captionText="Fig. 5. Arctigalloisiana yezoensis, male (B2-407735). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. Scales: A–D, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375269" httpUri="https://zenodo.org/record/10375269/files/figure.png" pageId="3" pageNumber="20">5C</figureCitation>
|
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and
|
||
<figureCitation id="130B2A34FFF0FFA5FBE7F920FB2463CC" box="[1125,1153,1725,1744]" captionStart="Fig" captionStartId="5.[100,130,1382,1399]" captionTargetBox="[340,1247,149,1354]" captionTargetId="figure-292@5.[338,1248,148,1355]" captionTargetPageId="5" captionText="Fig. 7. Arctigalloisiana poropnetopa sp. nov., holotype female (B2-407746). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) ovipositor, dorsal view. (G) same, lateral view. Scales: A–D, F–G, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375276" httpUri="https://zenodo.org/record/10375276/files/figure.png" pageId="3" pageNumber="20">7C</figureCitation>
|
||
,
|
||
<figureCitation id="130B2A34FFF0FFA5FB0CF920FB0E63CC" box="[1166,1195,1725,1744]" captionStart="Fig" captionStartId="6.[100,130,1383,1400]" captionTargetBox="[340,1247,150,1355]" captionTargetId="figure-376@6.[338,1248,148,1356]" captionTargetPageId="6" captionText="Fig. 8. Arctigalloisiana yubariensis sp. nov., holotype female (B2-407748). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) ovipositor, dorsal view. (G) same, lateral view. Scales: A–D, F–G, 1 mm; E, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10375280" httpUri="https://zenodo.org/record/10375280/files/figure.png" pageId="3" pageNumber="20">8C</figureCitation>
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||
), and poorly developed tarsal pulvillus (
|
||
<figureCitation id="130B2A34FFF0FFA5FC13F944FC7363F0" box="[913,982,1753,1772]" captionStart="Fig" captionStartId="3.[100,130,1390,1407]" captionTargetBox="[340,1247,149,1360]" captionTargetId="figure-361@3.[338,1248,148,1363]" captionTargetPageId="3" captionText="Fig. 4. Arctigalloisiana yezoensis, female (B2-407738). (A) Head and Pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) tarsal pulvillus of anterior leg, ventral view. (G) ovipositor, dorsal view. (H) same, lateral view. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10375267" httpUri="https://zenodo.org/record/10375267/files/figure.png" pageId="3" pageNumber="20">Fig. 4F</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C32A653AFFF0FFA2FCD3F969FE9E6271" lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20" type="description">
|
||
<paragraph id="8B8F36B1FFF0FFA2FCD3F969FE9E6271" blockId="3.[818,1488,1529,1967]" lastBlockId="4.[100,770,1881,1958]" lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20">
|
||
<emphasis id="B944EAA3FFF0FFA5FCD3F969FC63621B" bold="true" box="[849,966,1780,1799]" pageId="3" pageNumber="20">Description</emphasis>
|
||
. Antennomere number varying from 28 to 34. Third antennomere relatively short (Ant3L/Ant2L 1.2–1.7). Compound eye black. Lacinia with two teeth. Postmentum with a pair of setae. Outer margin of cervical sclerite without macrotrichiae. Pronotum with convex posterior margin; transverse sulcus at anterior margin convex or sometime straight. Cercal nine or ten segmented; length of each segment not stable among individuals. Ovipositor long, from 1.1 to 1.5 times longer than pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F6639FFF7FFA2FFE6FBA3FD066174" ID-DOI="http://doi.org/10.5281/zenodo.10375269" ID-Zenodo-Dep="10375269" httpUri="https://zenodo.org/record/10375269/files/figure.png" pageId="4" pageNumber="21" startId="4.[100,130,1086,1103]" targetBox="[340,1247,149,1056]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8F36B1FFF7FFA2FFE6FBA3FD066174" blockId="4.[100,1487,1086,1129]" pageId="4" pageNumber="21">
|
||
<emphasis id="B944EAA3FFF7FFA2FFE6FBA3FED76153" bold="true" box="[100,370,1086,1103]" pageId="4" pageNumber="21">
|
||
Fig. 5.
|
||
<taxonomicName id="4C304D32FFF7FFA2FF24FBA3FED76153" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" baseAuthorityName="Asahina" baseAuthorityYear="1961" box="[166,370,1086,1103]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="yezoensis">
|
||
<emphasis id="B944EAA3FFF7FFA2FF24FBA3FED76153" bold="true" box="[166,370,1086,1103]" italics="true" pageId="4" pageNumber="21">Arctigalloisiana yezoensis</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
, male (B2-407735). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. Scales: A–D, 1 mm; E, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6639FFF7FFA2FFEDF882FA63622C" ID-DOI="http://doi.org/10.5281/zenodo.10375273" ID-Zenodo-Dep="10375273" box="[111,1478,1823,1840]" httpUri="https://zenodo.org/record/10375273/files/figure.png" pageId="4" pageNumber="21" startId="4.[111,141,1823,1840]" targetBox="[340,1247,1189,1793]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8F36B1FFF7FFA2FFEDF882FA63622C" blockId="4.[111,1478,1823,1840]" box="[111,1478,1823,1840]" pageId="4" pageNumber="21">
|
||
<emphasis id="B944EAA3FFF7FFA2FFEDF882FF0C622C" bold="true" box="[111,169,1823,1840]" pageId="4" pageNumber="21">Fig. 6.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="4C304D32FFF7FFA2FE9EF882FE4C622C" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" baseAuthorityName="Asahina" baseAuthorityYear="1961" box="[284,489,1823,1840]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="yezoensis">
|
||
<emphasis id="B944EAA3FFF7FFA2FE9EF882FE4C622C" bold="true" box="[284,489,1823,1840]" italics="true" pageId="4" pageNumber="21">Arctigalloisiana yezoensis</emphasis>
|
||
</taxonomicName>
|
||
, male (B2-407736). (A) ventral view. (B) posterior view. (C) left-lateral view. (D) right-lateral view. Scales: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A653AFFF7FFA2FF06F8E8FC0F62B8" pageId="4" pageNumber="21" type="etymology">
|
||
<paragraph id="8B8F36B1FFF7FFA2FF06F8E8FC0F62B8" blockId="4.[100,770,1881,1958]" lastBlockId="4.[818,1487,1881,1956]" pageId="4" pageNumber="21">
|
||
<emphasis id="B944EAA3FFF7FFA2FF06F8E8FF556294" bold="true" box="[132,240,1909,1928]" pageId="4" pageNumber="21">Etymology</emphasis>
|
||
. The generic name is a compound noun derived from the Latin word,
|
||
<taxonomicName id="4C304D32FFF7FFA2FF60F80CFEBE62B8" box="[226,283,1937,1956]" class="Malacostraca" family="Scyllaridae" genus="Arctus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA3FFF7FFA2FF60F80CFEBE62B8" bold="true" box="[226,283,1937,1956]" italics="true" pageId="4" pageNumber="21">Arctus</emphasis>
|
||
</taxonomicName>
|
||
(Ursa Major/Minor = north; from αρκτος), and
|
||
<taxonomicName id="4C304D32FFF7FFA2FCB0F8C4FC336270" authorityName="Caudell and King" authorityYear="1924" box="[818,918,1881,1900]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="4" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA3FFF7FFA2FCB0F8C4FC336270" bold="true" box="[818,918,1881,1900]" italics="true" pageId="4" pageNumber="21">Galloisiana</emphasis>
|
||
</taxonomicName>
|
||
(feminine), referring to the fact that this genus is endemic to
|
||
<collectingRegion id="49F4F853FFF7FFA2FCB0F8E8FC2A6294" box="[818,911,1909,1928]" country="Japan" name="Hokkaido" pageId="4" pageNumber="21">Hokkaido</collectingRegion>
|
||
, which is the northernmost main island in the Japanese Archipelago.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F6639FFF6FFA3FFE6FAFBFAFE608C" ID-DOI="http://doi.org/10.5281/zenodo.10375276" ID-Zenodo-Dep="10375276" httpUri="https://zenodo.org/record/10375276/files/figure.png" pageId="5" pageNumber="22" startId="5.[100,130,1382,1399]" targetBox="[340,1247,149,1354]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8F36B1FFF6FFA3FFE6FAFBFAFE608C" blockId="5.[100,1487,1382,1425]" pageId="5" pageNumber="22">
|
||
<emphasis id="B944EAA3FFF6FFA3FFE6FAFBFE7C606A" bold="true" box="[100,473,1382,1399]" pageId="5" pageNumber="22">
|
||
Fig. 7.
|
||
<taxonomicName id="4C304D32FFF6FFA3FF24FAFBFE29606A" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[166,396,1382,1399]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="poropnetopa" status="sp. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FF24FAFBFE29606A" bold="true" box="[166,396,1382,1399]" italics="true" pageId="5" pageNumber="22">Arctigalloisiana poropnetopa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FE13FAFBFE7C606A" box="[401,473,1382,1398]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female (B2-407746). (A) head and pronotum, dorsal view. (B) same, lateral view. (C) head, ventral view. (D) cervical sclerite, ventral view. (E) left antenna, dorsal view. (F) ovipositor, dorsal view. (G) same, lateral view. Scales: A–D, F–G, 1 mm; E, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A653AFFF6FFA3FF06FA24FD8760F5" pageId="5" pageNumber="22" type="taxon_list">
|
||
<paragraph id="8B8F36B1FFF6FFA3FF06FA24FD8760F5" blockId="5.[100,770,1465,1708]" pageId="5" pageNumber="22">
|
||
<emphasis id="B944EAA3FFF6FFA3FF06FA24FEE660D0" bold="true" box="[132,323,1465,1484]" pageId="5" pageNumber="22">Species included</emphasis>
|
||
.
|
||
<taxonomicName id="4C304D32FFF6FFA3FEE4FA24FDC560D0" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" baseAuthorityName="Asahina" baseAuthorityYear="1961" box="[358,608,1465,1484]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yezoensis" status="comb. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FEE4FA24FDC560D0" bold="true" box="[358,608,1465,1484]" italics="true" pageId="5" pageNumber="22">Arctigalloisiana yezoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FDFFFA24FD5A60D1" box="[637,767,1465,1485]" pageId="5" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B944EAA3FFF6FFA3FFE6FA48FEE760F5" bold="true" box="[100,322,1493,1513]" pageId="5" pageNumber="22">
|
||
<taxonomicName id="4C304D32FFF6FFA3FFE6FA48FF4860F5" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[100,237,1493,1513]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="poropnetopa" status="sp. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FFE6FA48FF4860F5" bold="true" box="[100,237,1493,1513]" italics="true" pageId="5" pageNumber="22">A. poropnetopa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FF71FA4BFEE760F5" box="[243,322,1494,1513]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C304D32FFF6FFA3FECDFA48FE6860F4" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[335,461,1493,1512]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yubariensis" status="sp. nov.">A. yubariensis</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FE51FA4BFD8760F5" box="[467,546,1494,1513]" pageId="5" pageNumber="22" rank="species">
|
||
<emphasis id="B944EAA3FFF6FFA3FE51FA4BFDB960F5" bold="true" box="[467,540,1494,1513]" pageId="5" pageNumber="22">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A653AFFF6FFA3FF06FA6CFEFC638C" pageId="5" pageNumber="22" type="discussion">
|
||
<paragraph id="8B8F36B1FFF6FFA3FF06FA6CFEFC638C" blockId="5.[100,770,1465,1708]" pageId="5" pageNumber="22">
|
||
<emphasis id="B944EAA3FFF6FFA3FF06FA6CFF796318" bold="true" box="[132,220,1521,1540]" pageId="5" pageNumber="22">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation id="EFA14B40FFF6FFA3FF65FA6CFE116318" author="Schoville, S. D. & Uchifune, T. & Machida, R." box="[231,436,1521,1541]" pageId="5" pageNumber="22" pagination="915 - 927" refId="ref7938" refString="Schoville, S. D., Uchifune, T., Machida, R., 2013. Colliding fragment islands transport independent lineages of endemic rock-crawlers (Grylloblattodea: Grylloblattidae) in the Japanese archipelago. Mol. Phylogenet. Evol. 66, 915 - 927. https: // doi. org / 10.1016 / j. ympev. 2012.11.022." type="journal article" year="2013">Schoville et al. (2013)</bibRefCitation>
|
||
shows a result in which the clade of “
|
||
<taxonomicName id="4C304D32FFF6FFA3FFECF990FE9E633C" authorityName="Asahina" authorityYear="1961" box="[110,315,1549,1568]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yezoensis">
|
||
<emphasis id="B944EAA3FFF6FFA3FFECF990FF77633C" bold="true" box="[110,210,1549,1568]" italics="true" pageId="5" pageNumber="22">Galloisiana</emphasis>
|
||
”
|
||
<emphasis id="B944EAA3FFF6FFA3FF6BF990FE9E633C" bold="true" box="[233,315,1549,1568]" italics="true" pageId="5" pageNumber="22">yezoensis</emphasis>
|
||
</taxonomicName>
|
||
is separated from another
|
||
<taxonomicName id="4C304D32FFF6FFA3FDDCF990FD67633C" authorityName="Caudell and King" authorityYear="1924" box="[606,706,1549,1568]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAA3FFF6FFA3FDDCF990FD67633C" bold="true" box="[606,706,1549,1568]" italics="true" pageId="5" pageNumber="22">Galloisiana</emphasis>
|
||
</taxonomicName>
|
||
clade including the
|
||
<typeStatus id="548B8813FFF6FFA3FF65F9B7FEB56321" box="[231,272,1578,1597]" pageId="5" pageNumber="22">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C304D32FFF6FFA3FEE1F9B4FD8C6320" baseAuthorityName="Caudell & King" baseAuthorityYear="1924" box="[355,553,1577,1596]" class="Insecta" family="Grylloblattidae" genus="Galloisina" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B944EAA3FFF6FFA3FEE1F9B4FD8C6320" bold="true" box="[355,553,1577,1596]" italics="true" pageId="5" pageNumber="22">Galloisina nipponensis</emphasis>
|
||
</taxonomicName>
|
||
. Our phylogenetic tree also supports the independence of “
|
||
<taxonomicName id="4C304D32FFF6FFA3FE52F9D8FD396344" authorityName="Asahina" authorityYear="1961" box="[464,668,1605,1624]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yezoensis">
|
||
<emphasis id="B944EAA3FFF6FFA3FE52F9D8FD916344" bold="true" box="[464,564,1605,1624]" italics="true" pageId="5" pageNumber="22">Galloisiana</emphasis>
|
||
”
|
||
<emphasis id="B944EAA3FFF6FFA3FDC8F9D8FD396344" bold="true" box="[586,668,1605,1624]" italics="true" pageId="5" pageNumber="22">yezoensis</emphasis>
|
||
</taxonomicName>
|
||
. Here we establish a new genus,
|
||
<taxonomicName id="4C304D32FFF6FFA3FED8F9FCFE406368" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[346,485,1633,1652]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FED8F9FCFE406368" bold="true" box="[346,485,1633,1652]" italics="true" pageId="5" pageNumber="22">Arctigalloisiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FE70F9FCFDF76368" box="[498,594,1633,1652]" pageId="5" pageNumber="22" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, for “
|
||
<taxonomicName id="4C304D32FFF6FFA3FD16F9FCFF13638C" authorityName="Asahina" authorityYear="1961" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yezoensis">
|
||
<emphasis id="B944EAA3FFF6FFA3FD16F9FCFD5D6368" bold="true" box="[660,760,1633,1652]" italics="true" pageId="5" pageNumber="22">Galloisiana</emphasis>
|
||
”
|
||
<emphasis id="B944EAA3FFF6FFA3FFE6F9E0FF13638C" bold="true" box="[100,182,1661,1680]" italics="true" pageId="5" pageNumber="22">yezoensis</emphasis>
|
||
</taxonomicName>
|
||
and its relatives.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A653AFFF6FFA3FF06F904FECE63B0" box="[132,363,1689,1708]" pageId="5" pageNumber="22" type="multiple">
|
||
<paragraph id="8B8F36B1FFF6FFA3FF06F904FECE63B0" blockId="5.[100,770,1465,1708]" box="[132,363,1689,1708]" pageId="5" pageNumber="22">Keys to species (Female)</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A653AFFF6FFA3FF06F94CFBC360F5" pageId="5" pageNumber="22" type="key">
|
||
<key id="FFA16180FFF6FFA3FF06F94CFBC360F5" pageId="5" pageNumber="22">
|
||
<keyStep id="30C42E14FFF6FFA3FF06F94CFE3F6273" pageId="5" pageNumber="22">
|
||
<paragraph id="8B8F36B1FFF6FFA3FF06F94CFEC46200" blockId="5.[132,770,1744,1959]" pageId="5" pageNumber="22">
|
||
<keyLead id="30C19584FFF6FFA3FF06F94CFEC46200" pageId="5" pageNumber="22">
|
||
1.DV/PL less than 1.20 (the ovipositor short) ——————————————————————————————
|
||
<taxonomicName id="4C304D32FFF6FFA3FF06F895FF546207" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[132,241,1800,1819]" class="Insecta" family="Grylloblattidae" genus="Galloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yezoensis" status="comb. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FF06F895FF546207" bold="true" box="[132,241,1800,1819]" italics="true" pageId="5" pageNumber="22">A. yezoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FF75F895FEC46200" box="[247,353,1800,1820]" pageId="5" pageNumber="22" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8F36B1FFF6FFA3FF26F8B8FE3F6273" blockId="5.[132,770,1744,1959]" pageId="5" pageNumber="22">
|
||
<keyLead id="30C19584FFF6FFA3FF26F8B8FE3F6273" pageId="5" pageNumber="22">-. DV/PL more than 1.20 (the ovipositor long) ————————————————————————————————————————2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30C42E14FFF6FFA3FF06F8E5FBC360F5" pageId="5" pageNumber="22">
|
||
<paragraph id="8B8F36B1FFF6FFA3FF06F8E5FE3C62BB" blockId="5.[132,770,1744,1959]" pageId="5" pageNumber="22">
|
||
<keyLead id="30C19584FFF6FFA3FF06F8E5FE3C62BB" pageId="5" pageNumber="22">
|
||
2. Ant3L/Ant2L more than 1.50 (the antennomere 3rd long) ——
|
||
<emphasis id="B944EAA3FFF6FFA3FF2DF80EFE3C62BB" bold="true" box="[175,409,1939,1959]" pageId="5" pageNumber="22">
|
||
<taxonomicName id="4C304D32FFF6FFA3FF2DF80EFEE162BB" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[175,324,1939,1959]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="poropnetopa" status="sp. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FF2DF80EFEE162BB" bold="true" box="[175,324,1939,1959]" italics="true" pageId="5" pageNumber="22">A. poropnetopa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FEC8F809FE3C62BB" box="[330,409,1940,1959]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8F36B1FFF6FFA3FCF3FA27FBC360F5" blockId="5.[881,1487,1465,1513]" pageId="5" pageNumber="22">
|
||
<keyLead id="30C19584FFF6FFA3FCF3FA27FBC360F5" pageId="5" pageNumber="22">
|
||
-. Ant3L/Ant2L less than 1.50 (the antennomere 3rd short) —
|
||
<emphasis id="B944EAA3FFF6FFA3FC05FA49FBC360F5" bold="true" box="[903,1126,1492,1513]" pageId="5" pageNumber="22">
|
||
<taxonomicName id="4C304D32FFF6FFA3FC05FA49FBB560F4" authority="Nakahama & Yamasaki & Komazawa & Nakano, 2023" authorityName="Nakahama & Yamasaki & Komazawa & Nakano" authorityYear="2023" box="[903,1040,1492,1512]" class="Insecta" family="Grylloblattidae" genus="Arctigalloisiana" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="yubariensis" status="sp. nov.">
|
||
<emphasis id="B944EAA3FFF6FFA3FC05FA49FBB560F4" bold="true" box="[903,1040,1492,1512]" italics="true" pageId="5" pageNumber="22">A. yubariensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757D8FFF6FFA3FB95FA4BFBC360F5" box="[1047,1126,1494,1513]" pageId="5" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |