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<document id="C4C6B78A334FFA4E7007C7EAB193A841" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A138BFFF602B22FEE5FE3FFA91" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium fraternum Darby 2020, sp. nov." docType="treatment" docUuid="619FAB89-D53D-4C06-A2B0-3023099107E3" docUuidSource="ZooBank" docVersion="7" lastPageNumber="175" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="174" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D1B0A1C4F50A4CCAB13A1FDFC8A95A27" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="686C69B6EAF45EF90CB7F5B55609B6F2">
<mods:title id="15FAD51DD257A3037E3D4DC26A1099AF">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="5EE221BCB814BE4B10CD3E535F71152A" type="personal">
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<mods:roleTerm id="335275A1597C41C4B05A1B2F187C6740">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD156A9B4D8C3A3B1389726A4295FD2E">Darby, Michael</mods:namePart>
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<mods:title id="35C4B0BD865558FE822186CE6A84BD9A">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BADED201D86F878E19A8D2FEF0D19F1F">
<mods:date id="600FDB33B48E5A526B7C044E05D45CF9">2020</mods:date>
<mods:detail id="B11388F1785EC3C1E5C5FCC92BE99DFA" type="pubDate">
<mods:number id="B4E83BBB19EF0B4654FBBFE93FB90A6A">2020-04-02</mods:number>
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<mods:number id="9EFE4EDC269380C53789EE34D1F769E5">622</mods:number>
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<mods:classification id="F834171D1479E59C2693D3F392CD1684">journal article</mods:classification>
<mods:identifier id="FF0B91D6D6394B65507CD6A2A16A4A58" type="CLB-Dataset">22458</mods:identifier>
<mods:identifier id="6F493A34D0E28562818B4F6D72A8650C" type="DOI">10.5852/ejt.2020.622</mods:identifier>
<mods:identifier id="1DC8EFCCBE37F6CC0E9BEB0B16A17907" type="GBIF-Dataset">9cb3ea96-0ee4-40b4-8006-19b5fed80b46</mods:identifier>
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<treatment id="039987A138BFFF602B22FEE5FE3FFA91" ID-DOI="http://doi.org/10.5281/zenodo.3795742" ID-GBIF-Taxon="163877773" ID-Zenodo-Dep="3795742" LSID="urn:lsid:zoobank.org:act:619FAB89-D53D-4C06-A2B0-3023099107E3" httpUri="http://treatment.plazi.org/id/039987A138BFFF602B22FEE5FE3FFA91" lastPageId="174" lastPageNumber="175" pageId="173" pageNumber="174">
<subSubSection id="C32A653C38BFFF632B22FEE5FB5CFED1" pageId="173" pageNumber="174" type="nomenclature">
<paragraph id="8B8F36B738BFFF632B22FEE5FC55FEBE" blockId="173.[371,1216,267,364]" box="[619,969,267,294]" pageId="173" pageNumber="174">
<heading id="D0C781DB38BFFF632B22FEE5FC55FEBE" box="[619,969,267,294]" centered="true" fontSize="11" level="2" pageId="173" pageNumber="174" reason="2">
<taxonomicName id="4C304D3438BFFF632B22FEE5FCFBFEBD" ID-CoL="8XXQ4" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[619,871,267,294]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
<emphasis id="B944EAA538BFFF632B22FEE5FCFBFEBD" bold="true" box="[619,871,267,294]" italics="true" pageId="173" pageNumber="174">Cissidium fraternum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BFFF632A27FEE2FC55FEBE" box="[878,969,268,294]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B738BFFF63283AFEC0FB5CFED1" blockId="173.[371,1216,267,364]" box="[371,1216,302,329]" pageId="173" pageNumber="174">
<uri id="FFA13AB538BFFF63283AFEC0FB5CFED1" box="[371,1216,302,329]" pageId="173" pageNumber="174">
urn:lsid:zoobank.org:act:
<uuid id="FF960C6238BFFF632BD8FEC0FB5CFED1" box="[657,1216,302,329]" pageId="173" pageNumber="174">619FAB89-D53D-4C06-A2B0-3023099107E3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C38BFFF632B87FEBCFCFAFEF4" box="[718,870,338,364]" pageId="173" pageNumber="174" type="description">
<paragraph id="8B8F36B738BFFF632B87FEBCFCFAFEF4" blockId="173.[371,1216,267,364]" box="[718,870,338,364]" pageId="173" pageNumber="174">
<figureCitation id="130B2A3238BFFF632B87FEBCFCB1FEF4" box="[718,813,338,364]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="173" pageNumber="174">Figs 6H</figureCitation>
,
<figureCitation id="130B2A3238BFFF632A73FEBCFCFAFEF4" box="[826,870,338,364]" captionStart="Fig" captionStartId="173.[189,232,1964,1990]" captionTargetBox="[227,1376,1103,1920]" captionTargetId="figure@173.[220,1376,1098,1920]" captionTargetPageId="173" captionText="Fig. 109. Cissidium fraternum sp. nov. A. Habitus. B. Pronotum, × 415. C. Mesoventrite showing median process of collar, mid-keel and keel, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777502" httpUri="https://zenodo.org/record/3777502/files/figure.png" pageId="173" pageNumber="174">109</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C38BFFF6329F4FE7DFC1DFDBA" pageId="173" pageNumber="174" type="etymology">
<paragraph id="8B8F36B738BFFF6329F4FE7DFED8FE35" blockId="173.[189,324,403,429]" box="[189,324,403,429]" pageId="173" pageNumber="174">
<heading id="D0C781DB38BFFF6329F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" fontSize="11" level="3" pageId="173" pageNumber="174" reason="3">
<emphasis id="B944EAA538BFFF6329F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" pageId="173" pageNumber="174">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B738BFFF6329F4FE2CFC1DFDBA" blockId="173.[189,1398,450,547]" pageId="173" pageNumber="174">
Johnson ms name, derivation unclear but presumably from the Latin adjective ‘
<emphasis id="B944EAA538BFFF632DC3FE2CFB6AFE44" box="[1162,1270,450,476]" italics="true" pageId="173" pageNumber="174">fraternus</emphasis>
’, meaning ‘brotherly’ and possibly referring to the similarity of this species to
<taxonomicName id="4C304D3438BFFF632D5CFE0BFB4EFE67" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1045,1234,485,511]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="brachypterm" status="sp. nov.">
<emphasis id="B944EAA538BFFF632D5CFE0BFB4EFE67" box="[1045,1234,485,511]" italics="true" pageId="173" pageNumber="174">C. brachypterm</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BFFF632D96FE0BFADCFE67" box="[1247,1344,485,511]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C304D3438BFFF6329F4FDE7FEFBFDBA" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[189,359,520,547]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="greensladei" status="sp. nov.">
<emphasis id="B944EAA538BFFF6329F4FDE7FEFBFDBA" box="[189,359,520,547]" italics="true" pageId="173" pageNumber="174">C. greensladei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BFFF632827FDE7FE57FDBB" box="[366,459,521,547]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
, both also from the
<collectingCountry id="F327762738BFFF632BFCFDE6FCE1FDBA" box="[693,893,520,546]" name="Solomon Islands" pageId="173" pageNumber="174">Solomon Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C38BFFF6329F4FDA7FB72FCF9" pageId="173" pageNumber="174" type="materials_examined">
<paragraph id="8B8F36B738BFFF6329F4FDA7FE36FDFB" blockId="173.[189,426,585,611]" box="[189,426,585,611]" pageId="173" pageNumber="174">
<heading id="D0C781DB38BFFF6329F4FDA7FE36FDFB" bold="true" box="[189,426,585,611]" fontSize="11" level="3" pageId="173" pageNumber="174" reason="3">
<emphasis id="B944EAA538BFFF6329F4FDA7FE36FDFB" bold="true" box="[189,426,585,611]" pageId="173" pageNumber="174">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA38BFFF6329A3FD96FC99FD41" ID-GBIF-Occurrence="2609493671" collectingDate="1965-07-08" collectingMethod="thicket litter, matted roots" collectionCode="BMNH" collectorName="P. N. Lawrence" country="Solomon Islands" location="Umosani River" municipality="Monitor Creek" pageId="173" pageNumber="174" specimenCount="1" stateProvince="Guadalcanal" typeStatus="holotype">
<paragraph id="8B8F36B738BFFF6329A3FD96FEC0FD0A" blockId="173.[189,1398,632,729]" box="[234,348,632,658]" pageId="173" pageNumber="174">
<emphasis id="B944EAA538BFFF6329A3FD96FEC0FD0A" bold="true" box="[234,348,632,658]" pageId="173" pageNumber="174">
<typeStatus id="548B881538BFFF6329A3FD96FEC0FD0A" box="[234,348,632,658]" pageId="173" pageNumber="174" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B738BFFF6329F4FD75FC99FD41" blockId="173.[189,1398,632,729]" pageId="173" pageNumber="174">
<collectingCountry id="F327762738BFFF6329F4FD75FE48FD2D" box="[189,468,667,693]" name="Solomon Islands" pageId="173" pageNumber="174">SOLOMON ISLANDS</collectingCountry>
•
<specimenCount id="9D36FD3E38BFFF6328BBFD72FD96FD2E" box="[498,522,668,694]" pageId="173" pageNumber="174" type="male">♂</specimenCount>
;
<collectingRegion id="49F4F85538BFFF632B53FD75FD2CFD2D" box="[538,688,667,693]" country="Solomon Islands" name="Guadalcanal" pageId="173" pageNumber="174">Guadalcanal</collectingRegion>
,
<collectingMunicipality id="6BEBACCD38BFFF632BF7FD75FCEDFD2D" box="[702,881,667,693]" pageId="173" pageNumber="174">Monitor Creek</collectingMunicipality>
,
<location id="8EEF606C38BFFF632A36FD75FBA4FD2D" LSID="urn:lsid:plazi:treatment:039987A138BFFF602B22FEE5FE3FFA91:8EEF606C38BFFF632A36FD75FBA4FD2D" box="[895,1080,667,693]" country="Solomon Islands" municipality="Monitor Creek" name="Umosani River" pageId="173" pageNumber="174" stateProvince="Guadalcanal">Umosani River</location>
;
<date id="FF8E107738BFFF632D0FFD75FB4CFD2E" box="[1094,1232,667,694]" pageId="173" pageNumber="174" value="1965-07-08">
<collectingDate id="EFCAE99F38BFFF632D0FFD75FB4CFD2E" box="[1094,1232,667,694]" pageId="173" pageNumber="174" value="1965-07-08">8 Jul. 1965</collectingDate>
</date>
;
<collectingMethod id="52714EA038BFFF632D96FD75FECEFD41" pageId="173" pageNumber="174">thicket litter, matted roots</collectingMethod>
; no. 85;
<collectorName id="26C5536138BFFF6328FEFD51FDFFFD41" box="[439,611,703,729]" pageId="173" pageNumber="174">P.N. Lawrence</collectorName>
leg.;
<collectionCode id="ED21AE7238BFFF632BECFD51FC9DFD41" box="[677,769,703,729]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="173" pageNumber="174">BMNH</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B738BFFF6329A3FCEEFEFBFC82" blockId="173.[189,1398,768,865]" box="[234,359,768,794]" pageId="173" pageNumber="174">
<materialsCitation id="3B583CEA38BFFF6329A3FCEEFEFBFC82" ID-GBIF-Occurrence="2609493662" box="[234,359,768,794]" collectedFrom="gully bottom litter" collectingDate="1965-05-31" collectionCode="MMUE" collectorName="P. Naturaga" country="Solomon Islands" elevation="305" location="Nuhu" pageId="173" pageNumber="174" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B944EAA538BFFF6329A3FCEEFEFBFC82" bold="true" box="[234,359,768,794]" pageId="173" pageNumber="174">
<typeStatus id="548B881538BFFF6329A3FCEEFEFBFC82" box="[234,359,768,794]" pageId="173" pageNumber="174" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B738BFFF6329F4FCCDFB72FCF9" blockId="173.[189,1398,768,865]" pageId="173" pageNumber="174">
<collectingCountry id="F327762738BFFF6329F4FCCDFE4EFCA5" box="[189,466,803,829]" name="Solomon Islands" pageId="173" pageNumber="174">SOLOMON ISLANDS</collectingCountry>
•
<specimenCount id="9D36FD3E38BFFF6328A5FCCDFD86FCA6" box="[492,538,803,830]" pageId="173" pageNumber="174" type="female">1 ♀</specimenCount>
;
<location id="8EEF606C38BFFF632B60FCCDFDF2FCA5" LSID="urn:lsid:plazi:treatment:039987A138BFFF602B22FEE5FE3FFA91:8EEF606C38BFFF632B60FCCDFDF2FCA5" box="[553,622,803,829]" country="Solomon Islands" name="Nuhu" pageId="173" pageNumber="174">Nuhu</location>
;
<elevation id="001DD18438BFFF632B32FCCDFC8CFCA5" box="[635,784,803,829]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="173" pageNumber="174" unit="ft" value="1000.0">
<quantity id="4CC89B5238BFFF632B32FCCDFD53FCA5" box="[635,719,803,829]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="173" pageNumber="174" unit="ft" value="1000.0">1000 ft</quantity>
a.s.l.
</elevation>
;
<date id="FF8E107738BFFF632A57FCCDFC23FCA6" box="[798,959,803,830]" pageId="173" pageNumber="174" value="1965-05-31">
<collectingDate id="EFCAE99F38BFFF632A57FCCDFC23FCA6" box="[798,959,803,830]" pageId="173" pageNumber="174" value="1965-05-31">31 May 1965</collectingDate>
</date>
;
<collectedFrom id="382C754A38BFFF632A85FCCDFB3BFCA5" box="[972,1191,803,829]" pageId="173" pageNumber="174">gully bottom litter</collectedFrom>
;
<collectorName id="26C5536138BFFF632DFAFCCAFAA0FCA6" box="[1203,1340,804,830]" pageId="173" pageNumber="174">P. Naturaga</collectorName>
leg.;
<collectionCode id="ED21AE7238BFFF6329F4FCA9FE84FCF9" box="[189,280,839,865]" collectionName="United Kingdom, Manchester, The University, Manchester Museum" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="173" pageNumber="174">MMUE</collectionCode>
•
<materialsCitation id="3B583CEA38BFFF632878FCA8FB76FCF9" ID-GBIF-Occurrence="2609493660" box="[305,1258,838,865]" collectingDate="1966-03" collectionCode="BMNH" collectorName="P. Greenslade" country="Solomon Islands" location="Mt. Austen" pageId="173" pageNumber="174" specimenCount="1" specimenCount-female="1" stateProvince="Guadalcanal" typeStatus="paratype">
<specimenCount id="9D36FD3E38BFFF632878FCA8FEC2FCF9" box="[305,350,838,865]" pageId="173" pageNumber="174" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49F4F85538BFFF632824FCA8FD9FFCF8" box="[365,515,838,864]" country="Solomon Islands" name="Guadalcanal" pageId="173" pageNumber="174">Guadalcanal</collectingRegion>
,
<location id="8EEF606C38BFFF632B47FCA9FD08FCF9" LSID="urn:lsid:plazi:treatment:039987A138BFFF602B22FEE5FE3FFA91:8EEF606C38BFFF632B47FCA9FD08FCF9" box="[526,660,839,865]" country="Solomon Islands" name="Mt. Austen" pageId="173" pageNumber="174" stateProvince="Guadalcanal">Mt. Austen</location>
;
<date id="FF8E107738BFFF632BE9FCA9FC80FCF8" box="[672,796,838,865]" pageId="173" pageNumber="174" value="1966-03">
<collectingDate id="EFCAE99F38BFFF632BE9FCA9FC80FCF8" box="[672,796,838,865]" pageId="173" pageNumber="174" value="1966-03">Mar. 1966</collectingDate>
</date>
; no. 21336;
<collectorName id="26C5536138BFFF632AE4FCA9FBD0FCF8" box="[941,1100,838,865]" pageId="173" pageNumber="174">P. Greenslade</collectorName>
leg.;
<collectionCode id="ED21AE7238BFFF632DC7FCA9FB76FCF9" box="[1166,1258,839,865]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="173" pageNumber="174">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C38BFFF6029F4FC66FBCEFBB1" lastPageId="174" lastPageNumber="175" pageId="173" pageNumber="174" type="description">
<paragraph id="8B8F36B738BFFF6329F4FC66FED2FC3A" blockId="173.[189,334,904,930]" box="[189,334,904,930]" pageId="173" pageNumber="174">
<heading id="D0C781DB38BFFF6329F4FC66FED2FC3A" bold="true" box="[189,334,904,930]" fontSize="11" level="3" pageId="173" pageNumber="174" reason="3">
<emphasis id="B944EAA538BFFF6329F4FC66FED2FC3A" bold="true" box="[189,334,904,930]" pageId="173" pageNumber="174">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B738BFFF6329F4FC58FD24FC49" blockId="173.[189,696,950,977]" box="[189,696,950,977]" pageId="173" pageNumber="174">
SIZE. Habitus (
<figureCitation id="130B2A3238BFFF63282EFC58FE7FFC48" box="[359,483,950,976]" captionStart="Fig" captionStartId="173.[189,232,1964,1990]" captionTargetBox="[227,1376,1103,1920]" captionTargetId="figure@173.[220,1376,1098,1920]" captionTargetPageId="173" captionText="Fig. 109. Cissidium fraternum sp. nov. A. Habitus. B. Pronotum, × 415. C. Mesoventrite showing median process of collar, mid-keel and keel, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777502" httpUri="https://zenodo.org/record/3777502/files/figure.png" pageId="173" pageNumber="174">Fig. 109A</figureCitation>
), length
<quantity id="4CC89B5238BFFF632B01FC58FD28FC49" box="[584,692,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="173" pageNumber="174" unit="mm" value="0.6">0.60 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B738BFFF6329F4FC16FC77FB8A" blockId="173.[189,1003,1016,1042]" box="[189,1003,1016,1042]" pageId="173" pageNumber="174">COLOUR. Dusky yellow brown, antennae, legs and pubescence lighter.</paragraph>
<caption id="DF4F663F38BFFF6329F4F842FC8AF872" ID-DOI="http://doi.org/10.5281/zenodo.3777502" ID-Zenodo-Dep="3777502" httpUri="https://zenodo.org/record/3777502/files/figure.png" pageId="173" pageNumber="174" startId="173.[189,232,1964,1990]" targetBox="[227,1376,1103,1920]" targetPageId="173">
<paragraph id="8B8F36B738BFFF6329F4F842FC8AF872" blockId="173.[189,1398,1964,2026]" pageId="173" pageNumber="174">
<emphasis id="B944EAA538BFFF6329F4F842FEB1F85E" bold="true" box="[189,301,1964,1990]" pageId="173" pageNumber="174">Fig. 109.</emphasis>
<taxonomicName id="4C304D3438BFFF632871F842FDB2F85F" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[312,558,1964,1991]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
<emphasis id="B944EAA538BFFF632871F842FDB2F85F" box="[312,558,1964,1991]" italics="true" pageId="173" pageNumber="174">Cissidium fraternum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BFFF632B73F843FD05F85F" box="[570,665,1965,1991]" pageId="173" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B944EAA538BFFF632BECF842FD26F85E" bold="true" box="[677,698,1964,1990]" pageId="173" pageNumber="174">A</emphasis>
. Habitus.
<emphasis id="B944EAA538BFFF632A72F843FCD3F85F" bold="true" box="[827,847,1965,1991]" pageId="173" pageNumber="174">B</emphasis>
. Pronotum, × 415.
<emphasis id="B944EAA538BFFF632D0DF842FBC5F85E" bold="true" box="[1092,1113,1964,1990]" pageId="173" pageNumber="174">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel and keel, × 590.
</paragraph>
</caption>
<paragraph id="8B8F36B738BCFF6029F4FEE2FDCDFED2" blockId="174.[189,1399,267,331]" pageId="174" pageNumber="175">
HEAD. Width across eyes
<quantity id="4CC89B5238BCFF6028A6FEE5FDC1FEBE" box="[495,605,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="174" pageNumber="175" unit="mm" value="0.2">0.20 mm</quantity>
; antennomeres III–XI
<quantity id="4CC89B5238BCFF602A24FEE5FC4BFEBE" box="[877,983,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="174" pageNumber="175" unit="mm" value="0.3">0.30 mm</quantity>
long, III–IX
<quantity id="4CC89B5238BCFF602D33FEE5FB79FEBE" box="[1146,1253,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="174" pageNumber="175" unit="mm" value="0.16">0.16 mm</quantity>
long, X–XI
<quantity id="4CC89B5238BCFF6029F4FEDEFEBAFED3" box="[189,294,304,331]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="174" pageNumber="175" unit="mm" value="0.14">0.14 mm</quantity>
long; mentum as
<figureCitation id="130B2A3238BCFF6028B1FEDEFDCDFED2" box="[504,593,304,330]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="174" pageNumber="175">Fig 4A.</figureCitation>
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FE9AFC8EFDB9" blockId="174.[189,1399,371,545]" pageId="174" pageNumber="175">
PRONOTUM. Length
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, width
<quantity id="4CC89B5238BCFF602B26FE9DFD42FE16" box="[623,734,371,398]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="174" pageNumber="175" unit="mm" value="0.26">0.26 mm</quantity>
; base with a series of four or five foveae on each side separated by distinct space opposite scutellum and a few scattered smaller foveae anterior to these; lateral margins sharply angled, concave to slightly acute hind angles, sinuous to obtuse front angles, bordered, the border along the posterior margin interrupted between the hind angles and the median emargination in front of the scutellum (
<figureCitation id="130B2A3238BCFF602BC3FDE8FC98FDB9" box="[650,772,518,545]" captionStart="Fig" captionStartId="173.[189,232,1964,1990]" captionTargetBox="[227,1376,1103,1920]" captionTargetId="figure@173.[220,1376,1098,1920]" captionTargetPageId="173" captionText="Fig. 109. Cissidium fraternum sp. nov. A. Habitus. B. Pronotum, × 415. C. Mesoventrite showing median process of collar, mid-keel and keel, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777502" httpUri="https://zenodo.org/record/3777502/files/figure.png" pageId="174" pageNumber="175">Fig. 109B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FDA4FABDFDFB" blockId="174.[189,1313,585,612]" box="[189,1313,585,612]" pageId="174" pageNumber="175">
ELYTRA. Width
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; length.
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, basal half rather sparsely covered with large foveolae.
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FD63FED1FCA2" blockId="174.[189,1399,652,826]" pageId="174" pageNumber="175">
MESOVENTRITE. Collar barely extended medially onto mid-keel; mid-keel wider anteriorly than posteriorly, the posterior angles clearly joining the mesocoxal anterior border as two carinae, sharply raised medially to junction with keel; keel with ± two setae, parallel-sided to blunt termination posterior to the interruption point of the mesocoxal margins; mesoventral lateral margins serrate in posterior half (
<figureCitation id="130B2A3238BCFF60298FFCCEFEDCFCA2" box="[198,320,800,826]" captionStart="Fig" captionStartId="173.[189,232,1964,1990]" captionTargetBox="[227,1376,1103,1920]" captionTargetId="figure@173.[220,1376,1098,1920]" captionTargetPageId="173" captionText="Fig. 109. Cissidium fraternum sp. nov. A. Habitus. B. Pronotum, × 415. C. Mesoventrite showing median process of collar, mid-keel and keel, × 590." figureDoi="http://doi.org/10.5281/zenodo.3777502" httpUri="https://zenodo.org/record/3777502/files/figure.png" pageId="174" pageNumber="175">Fig. 109C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FC8DFBCDFC3A" blockId="174.[189,1399,867,930]" pageId="174" pageNumber="175">
METAVENTRITE. Sparsely pubescent, length
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, width across spines
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, with a triangularly impressed area medially and a small medial fovea posterior to the mesocoxae.
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FC25FE5EFC7D" blockId="174.[189,450,971,997]" box="[189,450,971,997]" pageId="174" pageNumber="175">WINGS. Macropterous.</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FBE0FBCEFBB1" blockId="174.[189,1106,1038,1065]" box="[189,1106,1038,1065]" pageId="174" pageNumber="175">
GENITALIA. Female spermatheca globular. Male aedeagus distinctive (
<figureCitation id="130B2A3238BCFF602AAFFBE0FBD8FBB1" box="[998,1092,1038,1065]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="174" pageNumber="175">Fig. 6H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A653C38BCFF6029F4FBBFFE3FFA91" pageId="174" pageNumber="175" type="discussion">
<paragraph id="8B8F36B738BCFF6029F4FBBFFEB3FBF3" blockId="174.[189,303,1105,1131]" box="[189,303,1105,1131]" pageId="174" pageNumber="175">
<heading id="D0C781DB38BCFF6029F4FBBFFEB3FBF3" bold="true" box="[189,303,1105,1131]" fontSize="11" level="3" pageId="174" pageNumber="175" reason="3">
<emphasis id="B944EAA538BCFF6029F4FBBFFEB3FBF3" bold="true" box="[189,303,1105,1131]" pageId="174" pageNumber="175">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B738BCFF6029F4FB6FFE3FFA91" blockId="174.[189,1399,1153,1289]" pageId="174" pageNumber="175">
Compared with the other five
<collectingCountry id="F327762738BCFF602B5CFB6FFD44FB03" box="[533,728,1153,1179]" name="Solomon Islands" pageId="174" pageNumber="175">Solomon Islands</collectingCountry>
species in this group,
<taxonomicName id="4C304D3438BCFF602A9EFB6FFBF1FB03" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[983,1133,1153,1179]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="fraternum" status="sp. nov.">
<emphasis id="B944EAA538BCFF602A9EFB6FFBF1FB03" box="[983,1133,1153,1179]" italics="true" pageId="174" pageNumber="175">C. fraternum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BCFF602D3AFB6FFB50FB03" box="[1139,1228,1153,1179]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
is very similar in size to
<taxonomicName id="4C304D3438BCFF60287EFB48FD9DFB58" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[311,513,1190,1216]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="brachypterum" status="sp. nov.">
<emphasis id="B944EAA538BCFF60287EFB48FD9DFB58" box="[311,513,1190,1216]" italics="true" pageId="174" pageNumber="175">C. brachypterum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BCFF602B43FB48FDF6FB58" box="[522,618,1190,1216]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
, but with longer antennae. Compared to
<taxonomicName id="4C304D3438BCFF602D12FB48FA94FB58" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1115,1288,1190,1216]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="174" pageNumber="175" phylum="Arthropoda" rank="species" species="greensladei" status="sp. nov.">
<emphasis id="B944EAA538BCFF602D12FB48FA94FB58" box="[1115,1288,1190,1216]" italics="true" pageId="174" pageNumber="175">C. greensladei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE38BCFF602C5BFB48FAEEFB58" box="[1298,1394,1190,1216]" pageId="174" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
, which it also much resembles, it is smaller and with shorter antennae. The aedeagus of the male provides decisive characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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