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<document id="19B76C84A4ABB030A3107EBB3C083840" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13858FF852B0EFE7FFB6CFB18" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium amboroense Darby 2015" docType="treatment" docVersion="7" lastPageNumber="76" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="75" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="87BEE0EEB3457A2B5A27BBD764A7B402" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="01D3CAD36EF41FBC4DD5C3E1E0E569CD">
<mods:title id="F5B491D8A65A7FBEF814611866652C3E">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
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<mods:role id="50724C5C17CDBEB90887FAF3D535451E">
<mods:roleTerm id="7A14F650A8B82710AAB0729B8ECD3D16">Author</mods:roleTerm>
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<mods:namePart id="98E4189C9F55FC48818271323BCA846D">Darby, Michael</mods:namePart>
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<mods:title id="F340366A3F7BA45EA45A294F598BDD93">European Journal of Taxonomy</mods:title>
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<mods:part id="5AFE28A670E86E704B47C135CB2FB508">
<mods:date id="E1CB8CDC83E3D2D26AAA01E44B702B55">2020</mods:date>
<mods:detail id="DAA4B4C118E790073167E72EC667E3A9" type="pubDate">
<mods:number id="F5759C4D9549D9A4258FCC362867560E">2020-04-02</mods:number>
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<mods:number id="BE6FEBEBEFB7AF61CEE7C0F3840C19EE">622</mods:number>
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<mods:identifier id="511CC3870B0716302FCEE7AA69D74BC1" type="CLB-Dataset">22458</mods:identifier>
<mods:identifier id="2EAC50FDF01E0595817D206518255C54" type="DOI">10.5852/ejt.2020.622</mods:identifier>
<mods:identifier id="08E32FC78ACC2C49B87A428E0B69FE11" type="GBIF-Dataset">9cb3ea96-0ee4-40b4-8006-19b5fed80b46</mods:identifier>
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<treatment id="039987A13858FF852B0EFE7FFB6CFB18" ID-DOI="http://doi.org/10.5281/zenodo.3795616" ID-GBIF-Taxon="163877824" ID-Zenodo-Dep="3795616" LSID="urn:lsid:plazi:treatment:039987A13858FF852B0EFE7FFB6CFB18" httpUri="http://treatment.plazi.org/id/039987A13858FF852B0EFE7FFB6CFB18" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75">
<subSubSection id="C32A653C3858FF842B0EFE7FFC71FE34" box="[583,1005,401,428]" pageId="74" pageNumber="75" type="nomenclature">
<paragraph id="8B8F36B73858FF842B0EFE7FFC71FE34" blockId="74.[583,1005,401,463]" box="[583,1005,401,428]" pageId="74" pageNumber="75">
<heading id="D0C781DB3858FF842B0EFE7FFC71FE34" box="[583,1005,401,428]" centered="true" fontSize="11" level="2" pageId="74" pageNumber="75" reason="2">
<taxonomicName id="4C304D343858FF842B0EFE7FFC71FE34" ID-CoL="VK84" authority="Darby 2015" authorityName="Darby" authorityYear="2015" box="[583,1005,401,428]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="amboroense">
<emphasis id="B944EAA53858FF842B0EFE7FFCC6FE34" bold="true" box="[583,858,401,428]" italics="true" pageId="74" pageNumber="75">Cissidium amboroense</emphasis>
<bibRefCitation id="EFA14B463858FF842A28FE7FFC71FE34" author="Darby M." box="[865,1005,401,428]" pageId="74" pageNumber="75" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby 2015</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3858FF842BEDFE5BFC13FE57" box="[676,911,437,463]" pageId="74" pageNumber="75" type="description">
<paragraph id="8B8F36B73858FF842BEDFE5BFC13FE57" blockId="74.[583,1005,401,463]" box="[676,911,437,463]" pageId="74" pageNumber="75">
<figureCitation id="130B2A323858FF842BEDFE5BFC9FFE57" box="[676,771,437,463]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="74" pageNumber="75">Figs 4C</figureCitation>
,
<figureCitation id="130B2A323858FF842A46FE5BFCAFFE57" box="[783,819,437,463]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="74" pageNumber="75">6B</figureCitation>
,
<figureCitation id="130B2A323858FF842A09FE5BFCF9FE57" box="[832,869,437,463]" captionStart="Fig" captionStartId="8.[189,232,1861,1887]" captionTargetBox="[190,1393,524,1810]" captionTargetId="figure@8.[189,1398,523,1818]" captionTargetPageId="8" captionText="Fig. 7. Metaventral and male ventrite examples. A–B. Metaventrite posterior margin to show spines. A. Cissidium riparium sp. nov., × 610. B. Cissidium plaumanni sp. nov., × 970. C–D. Metaventral discs. C. Cissidium greensladei sp. nov., × 455. D. Cissidium insulare sp. nov., × 595. E. Cissidium obscenum sp. nov., × 840. F–G. Terminal male ventrites. F. Cissidium petri Darby, 2015, × 1580. G. Cissidium amboroense Darby, 2015, × 1380." figureDoi="http://doi.org/10.5281/zenodo.3777296" httpUri="https://zenodo.org/record/3777296/files/figure.png" pageId="74" pageNumber="75">7G</figureCitation>
,
<figureCitation id="130B2A323858FF842A3BFE5BFC13FE57" box="[882,911,437,463]" captionStart="Fig" captionStartId="74.[189,232,1964,1990]" captionTargetBox="[218,1368,1027,1927]" captionTargetId="figure@74.[210,1368,1017,1928]" captionTargetPageId="74" captionText="Fig. 47. Cissidium amboroense Darby, 2015. A. Habitus. B. Pronotum. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1220." figureDoi="http://doi.org/10.5281/zenodo.3777378" httpUri="https://zenodo.org/record/3777378/files/figure.png" pageId="74" pageNumber="75">47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3858FF8529F4FE18FC6AFC74" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="8B8F36B73858FF8429F4FE18FE36FD88" blockId="74.[189,426,502,528]" box="[189,426,502,528]" pageId="74" pageNumber="75">
<heading id="D0C781DB3858FF8429F4FE18FE36FD88" bold="true" box="[189,426,502,528]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B944EAA53858FF8429F4FE18FE36FD88" bold="true" box="[189,426,502,528]" pageId="74" pageNumber="75">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA3858FF8429A3FDCBFBF1FD1D" ID-GBIF-Occurrence="2609493572" collectingDate="2013-11-28" collectingMethod="sifting litter; Winkler app. extr." collectionCode="UASC" collectorName="P. Banar" country="Bolivia" elevation="2450" latitude="-17.836668" location="Siberia Pass" longLatPrecision="21" longitude="64.70222" pageId="74" pageNumber="75" specimenCount="1" specimenCount-female="1" stateProvince="Santa Cruz" typeStatus="holotype">
<paragraph id="8B8F36B73858FF8429A3FDCBFEC0FDA7" blockId="74.[189,1399,549,646]" box="[234,348,549,575]" pageId="74" pageNumber="75">
<emphasis id="B944EAA53858FF8429A3FDCBFEC0FDA7" bold="true" box="[234,348,549,575]" pageId="74" pageNumber="75">
<typeStatus id="548B88153858FF8429A3FDCBFEC0FDA7" box="[234,348,549,575]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73858FF8429F4FDA6FBF1FD1D" blockId="74.[189,1399,549,646]" pageId="74" pageNumber="75">
<collectingCountry id="F32776273858FF8429F4FDA6FEA8FDFA" box="[189,308,584,610]" name="Bolivia" pageId="74" pageNumber="75">BOLIVIA</collectingCountry>
•
<specimenCount id="9D36FD3E3858FF84281EFDA6FEF2FDFA" box="[343,366,584,610]" pageId="74" pageNumber="75" type="female">♀</specimenCount>
;
<collectingRegion id="49F4F8553858FF8428CBFDA6FD95FDFA" box="[386,521,584,610]" country="Bolivia" name="Santa Cruz" pageId="74" pageNumber="75">Santa Cruz</collectingRegion>
dep., W. Amboro NP,
<location id="8EEF606C3858FF842A63FDA6FC22FDFA" LSID="urn:lsid:plazi:treatment:039987A13858FF852B0EFE7FFB6CFB18:8EEF606C3858FF842A63FDA6FC22FDFA" box="[810,958,584,610]" country="Bolivia" latitude="-17.836668" longLatPrecision="21" longitude="64.70222" name="Siberia Pass" pageId="74" pageNumber="75" stateProvince="Santa Cruz">Siberia Pass</location>
env.;
<elevation id="001DD1843858FF842D46FDA6FB2EFDFA" box="[1039,1202,584,611]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="74" pageNumber="75" unit="m" value="2450.0">
<quantity id="4CC89B523858FF842D46FDA6FBF1FDFA" box="[1039,1133,584,611]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="74" pageNumber="75" unit="m" value="2450.0">2450 m</quantity>
a.s.l.
</elevation>
;
<date id="FF8E10773858FF842D8CFDA6FAEEFDFA" box="[1221,1394,584,610]" pageId="74" pageNumber="75" value="2013-11-28">
<collectingDate id="EFCAE99F3858FF842D8CFDA6FAEEFDFA" box="[1221,1394,584,610]" pageId="74" pageNumber="75" value="2013-11-28">28 Nov. 2013</collectingDate>
</date>
;
<geoCoordinate id="EE0450703858FF8429F4FD87FED1FD1D" box="[189,333,617,645]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="74" pageNumber="75" precision="15" seconds="12" value="-17.836668">17°50′12″ S</geoCoordinate>
,
<geoCoordinate id="EE0450703858FF842813FD87FE70FD1E" box="[346,492,617,646]" degrees="64" direction="east" minutes="42" orientation="longitude" pageId="74" pageNumber="75" precision="15" seconds="08" value="64.70222">64°42′08″ E</geoCoordinate>
;
<collectingMethod id="52714EA03858FF8428B0FD85FCF4FD1E" box="[505,872,619,646]" pageId="74" pageNumber="75">sifting litter; Winkler app. extr.</collectingMethod>
;
<collectorName id="26C553613858FF842A3DFD82FC49FD1D" box="[884,981,619,646]" pageId="74" pageNumber="75">P. Baňař</collectorName>
leg.;
<collectionCode id="ED21AE723858FF842D5EFD85FBF5FD1D" box="[1047,1129,619,645]" country="Bolivia" httpUri="http://grbio.org/cool/kme8-7yd8" name="Museo de Historia Natural &amp;quot;Noel Kempff Mercado&amp;quot;" pageId="74" pageNumber="75">UASC</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B73858FF8429A3FD43FEFBFD5F" blockId="74.[189,1399,685,817]" box="[234,359,685,711]" pageId="74" pageNumber="75">
<materialsCitation id="3B583CEA3858FF8429A3FD43FEFBFD5F" ID-GBIF-Occurrence="2609493540" box="[234,359,685,711]" collectingDate="2013-11-28" collectingMethod="sifting litter; Winkler app. extr." collectionCode="MMBC, BMNH" collectorName="P. Banar" country="Bolivia" elevation="2450" latitude="-17.836668" location="BOLIVIA" longLatPrecision="21" longitude="64.70222" pageId="74" pageNumber="75" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="paratype">
<emphasis id="B944EAA53858FF8429A3FD43FEFBFD5F" bold="true" box="[234,359,685,711]" pageId="74" pageNumber="75">
<typeStatus id="548B88153858FF8429A3FD43FEFBFD5F" box="[234,359,685,711]" pageId="74" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B73858FF8429F4FD3EFB1DFCA9" blockId="74.[189,1399,685,817]" pageId="74" pageNumber="75">
<collectingCountry id="F32776273858FF8429F4FD3EFEA8FD72" box="[189,308,720,746]" name="Bolivia" pageId="74" pageNumber="75">BOLIVIA</collectingCountry>
•
<specimenCount id="9D36FD3E3858FF842806FD3EFE0FFD72" box="[335,403,720,746]" pageId="74" pageNumber="75" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D36FD3E3858FF8428E8FD3EFE4CFD72" box="[417,464,720,746]" pageId="74" pageNumber="75" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED21AE723858FF842AD2FD3EFC6AFD72" box="[923,1014,720,746]" collectionName="Czech Republic, Brno, Moravske Muzeum [Moravian Museum]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="74" pageNumber="75">MMBC</collectionCode>
/
<collectionCode id="ED21AE723858FF842AB7FD3EFBCAFD72" box="[1022,1110,720,746]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="74" pageNumber="75">BMNH</collectionCode>
•
<materialsCitation id="3B583CEA3858FF842D38FD3EFBE1FCA9" ID-GBIF-Occurrence="2609493695" collectedFrom="sifting litter" collectingDate="2013-11" collectingMethod="Winkler app. extr." collectionCode="BMNH" collectorName="P. Banar" country="Bolivia" elevation="2481" latitude="-17.840279" location="Siberia Pass" longLatPrecision="21" longitude="64.69945" pageId="74" pageNumber="75" specimenCode="GPS 31" specimenCount="1" stateProvince="Santa Cruz" typeStatus="paratype">
<specimenCount id="9D36FD3E3858FF842D38FD3EFB32FD72" box="[1137,1198,720,746]" pageId="74" pageNumber="75" type="generic">1 ex.</specimenCount>
; “
<specimenCode id="DB969ECC3858FF842D8FFD3EFABBFD72" box="[1222,1319,720,746]" pageId="74" pageNumber="75">GPS 31</specimenCode>
, BOL/
<collectingDate id="EFCAE99F3858FF8429F4FD1DFEFCFC95" box="[189,352,755,781]" pageId="74" pageNumber="75">
<date id="FF8E10773858FF8429F4FD1DFEA5FC95" box="[189,313,755,781]" pageId="74" pageNumber="75" value="2013-11">Nov 2013</date>
/08
</collectingDate>
;
<collectingCountry id="F32776273858FF84283BFD1DFE75FC95" box="[370,489,755,781]" name="Bolivia" pageId="74" pageNumber="75">BOLIVIA</collectingCountry>
2013
<collectingRegion id="49F4F8553858FF842B74FD1DFD59FC95" box="[573,709,755,781]" country="Bolivia" name="Santa Cruz" pageId="74" pageNumber="75">Santa Cruz</collectingRegion>
dep.”, W. Amboro NP,
<location id="8EEF606C3858FF842ABCFD1DFB15FC95" LSID="urn:lsid:plazi:treatment:039987A13858FF852B0EFE7FFB6CFB18:8EEF606C3858FF842ABCFD1DFB15FC95" box="[1013,1161,755,781]" country="Bolivia" latitude="-17.840279" longLatPrecision="21" longitude="64.69945" name="Siberia Pass" pageId="74" pageNumber="75" stateProvince="Santa Cruz">Siberia Pass</location>
env.;
<geoCoordinate id="EE0450703858FF842D92FD1FFAEDFC95" box="[1243,1393,753,781]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="74" pageNumber="75" precision="15" seconds="25" value="-17.840279">17°50′25″ S</geoCoordinate>
,
<geoCoordinate id="EE0450703858FF8429F4FCFAFECCFCA9" box="[189,336,788,817]" degrees="64" direction="east" minutes="41" orientation="longitude" pageId="74" pageNumber="75" precision="15" seconds="58" value="64.69945">64°41′58″ E</geoCoordinate>
;
<elevation id="001DD1843858FF842815FCF8FE69FCA8" box="[348,501,790,817]" metricMagnitude="3" metricUnit="m" metricValue="2.481" pageId="74" pageNumber="75" unit="m" value="2481.0">
<quantity id="4CC89B523858FF842815FCF8FE29FCA9" box="[348,437,790,817]" metricMagnitude="3" metricUnit="m" metricValue="2.481" pageId="74" pageNumber="75" unit="m" value="2481.0">2481 m</quantity>
a.s.l.
</elevation>
;
<collectedFrom id="382C754A3858FF842B4BFCF8FD0CFCA8" box="[514,656,790,816]" pageId="74" pageNumber="75">sifting litter</collectedFrom>
;
<collectingMethod id="52714EA03858FF842BD2FCF8FCEDFCA9" box="[667,881,790,817]" pageId="74" pageNumber="75">Winkler app. extr.</collectingMethod>
;
<collectorName id="26C553613858FF842A34FCF9FC43FCA8" box="[893,991,790,817]" pageId="74" pageNumber="75">P. Baňař</collectorName>
leg.;
<collectionCode id="ED21AE723858FF842D68FCF9FBE1FCA9" box="[1057,1149,791,817]" pageId="74" pageNumber="75">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F36B73858FF8429F4FCB9FD8DFCE9" blockId="74.[189,529,855,881]" box="[189,529,855,881]" pageId="74" pageNumber="75">
<heading id="D0C781DB3858FF8429F4FCB9FD8DFCE9" bold="true" box="[189,529,855,881]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B944EAA53858FF8429F4FCB9FD8DFCE9" bold="true" box="[189,529,855,881]" pageId="74" pageNumber="75">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73858FF8429F4FC68FB76FC5C" blockId="74.[189,1398,902,964]" pageId="74" pageNumber="75">
The following description amplifies the
<typeStatus id="548B88153858FF842BDCFC69FD54FC39" box="[661,712,903,929]" pageId="74" pageNumber="75">type</typeStatus>
description of
<bibRefCitation id="EFA14B463858FF842A30FC68FB87FC39" author="Darby M." box="[889,1051,902,929]" pageId="74" pageNumber="75" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby (2015)</bibRefCitation>
, in particular by adding more details of the meso- and metaventral characters, in order to conform with the present paper.
</paragraph>
<caption id="DF4F663F3858FF8429F4F842FD5BF872" ID-DOI="http://doi.org/10.5281/zenodo.3777378" ID-Zenodo-Dep="3777378" httpUri="https://zenodo.org/record/3777378/files/figure.png" pageId="74" pageNumber="75" startId="74.[189,232,1964,1990]" targetBox="[218,1368,1027,1927]" targetPageId="74">
<paragraph id="8B8F36B73858FF8429F4F842FD5BF872" blockId="74.[189,1398,1964,2026]" pageId="74" pageNumber="75">
<emphasis id="B944EAA53858FF8429F4F842FE8BF85F" bold="true" box="[189,279,1964,1991]" pageId="74" pageNumber="75">Fig. 47.</emphasis>
<taxonomicName id="4C304D343858FF842854F842FD23F85F" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[285,703,1964,1991]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="amboroense">
<emphasis id="B944EAA53858FF842854F842FDBAF85E" box="[285,550,1964,1990]" italics="true" pageId="74" pageNumber="75">Cissidium amboroense</emphasis>
<bibRefCitation id="EFA14B463858FF842B65F842FD23F85F" author="Darby M." box="[556,703,1964,1991]" pageId="74" pageNumber="75" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
.
<emphasis id="B944EAA53858FF842B81F842FD41F85E" bold="true" box="[712,733,1964,1990]" pageId="74" pageNumber="75">A</emphasis>
. Habitus.
<emphasis id="B944EAA53858FF842A1AF843FCFBF85F" bold="true" box="[851,871,1965,1991]" pageId="74" pageNumber="75">B</emphasis>
. Pronotum.
<emphasis id="B944EAA53858FF842ABDF842FB95F85E" bold="true" box="[1012,1033,1964,1990]" pageId="74" pageNumber="75">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel and keel, × 1220.
</paragraph>
</caption>
<paragraph id="8B8F36B73859FF8529F4FEE5FD36FEBE" blockId="75.[189,682,267,294]" box="[189,682,267,294]" pageId="75" pageNumber="76">
SIZE. Habitus (
<figureCitation id="130B2A323859FF85282EFEE5FE48FEBD" box="[359,468,267,293]" captionStart="Fig" captionStartId="74.[189,232,1964,1990]" captionTargetBox="[218,1368,1027,1927]" captionTargetId="figure@74.[210,1368,1017,1928]" captionTargetPageId="74" captionText="Fig. 47. Cissidium amboroense Darby, 2015. A. Habitus. B. Pronotum. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1220." figureDoi="http://doi.org/10.5281/zenodo.3777378" httpUri="https://zenodo.org/record/3777378/files/figure.png" pageId="75" pageNumber="76">Fig. 47A</figureCitation>
), length
<quantity id="4CC89B523859FF852B73FEE5FD39FEBE" box="[570,677,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="75" pageNumber="76" unit="mm" value="0.78">0.78 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FEA3FCDDFEFF" blockId="75.[189,833,333,360]" box="[189,833,333,360]" pageId="75" pageNumber="76">COLOUR. Dusky yellow, legs and antennae a little paler.</paragraph>
<paragraph id="8B8F36B73859FF8529F4FE7EFCC1FE55" blockId="75.[189,1399,399,461]" pageId="75" pageNumber="76">
HEAD. Antennomeres III–XI length
<quantity id="4CC89B523859FF852B24FE61FD40FE32" box="[621,732,399,426]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="75" pageNumber="76" unit="mm" value="0.38">0.38 mm</quantity>
, III–IX length
<quantity id="4CC89B523859FF852ADCFE61FB98FE32" box="[917,1028,399,426]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="75" pageNumber="76" unit="mm" value="0.19">0.19 mm</quantity>
, X–XI length
<quantity id="4CC89B523859FF852DFCFE61FAB8FE32" box="[1205,1316,399,426]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="75" pageNumber="76" unit="mm" value="0.19">0.19 mm</quantity>
; width across eyes
<quantity id="4CC89B523859FF852800FE5DFE29FE55" box="[329,437,435,461]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="75" pageNumber="76" unit="mm" value="0.21">0.21 mm</quantity>
; mentum and submentum (
<figureCitation id="130B2A323859FF852BBAFE5DFCCCFE55" box="[755,848,435,461]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="75" pageNumber="76">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FE1BFCF7FDCE" blockId="75.[189,1398,501,598]" pageId="75" pageNumber="76">
PRONOTUM. Length
<quantity id="4CC89B523859FF8528E2FE1BFD81FD97" box="[427,541,501,527]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="75" pageNumber="76" unit="mm" value="0.31">0.31 mm</quantity>
, width,
<quantity id="4CC89B523859FF852BCFFE1BFD64FD97" box="[646,760,501,527]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="75" pageNumber="76" unit="mm" value="0.2">0.20 mm</quantity>
, shining, pubescent; lateral margins slightly angled medially, concave to rectangular hind angles, bordered, the border continuing along the posterior margin, medial emargination before scutellum sinuous. (
<figureCitation id="130B2A323859FF852BBBFDD2FCC2FDCE" box="[754,862,572,598]" captionStart="Fig" captionStartId="74.[189,232,1964,1990]" captionTargetBox="[218,1368,1027,1927]" captionTargetId="figure@74.[210,1368,1017,1928]" captionTargetPageId="74" captionText="Fig. 47. Cissidium amboroense Darby, 2015. A. Habitus. B. Pronotum. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1220." figureDoi="http://doi.org/10.5281/zenodo.3777378" httpUri="https://zenodo.org/record/3777378/files/figure.png" pageId="75" pageNumber="76">Fig. 47B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FD90FD1FFD00" blockId="75.[189,643,638,664]" box="[189,643,638,664]" pageId="75" pageNumber="76">
ELYTRA.
<quantity id="4CC89B523859FF852869FD90FE15FD00" box="[288,393,638,664]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="75" pageNumber="76" unit="mm" value="0.47">0.47 mm</quantity>
long,
<quantity id="4CC89B523859FF85289BFD90FDA7FD00" box="[466,571,638,664]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="75" pageNumber="76" unit="mm" value="0.38">0.38 mm</quantity>
wide.
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FD2EFD4DFCDD" blockId="75.[189,1399,704,837]" pageId="75" pageNumber="76">
MESOVENTRITE. Medial extension of collar, broad, parallel-sided; mid-keel widest posteriorly, hind angles not reaching mesocoxal borders; keel long, with 4 setae, tapering narrowly to point at the base of the mesocoxae; mesoventral lateral margins evenly rounded without serrations; humeri toothed, posterior margin sloping strongly posteriad (
<figureCitation id="130B2A323859FF852B11FCC4FD58FCDD" box="[600,708,810,837]" captionStart="Fig" captionStartId="74.[189,232,1964,1990]" captionTargetBox="[218,1368,1027,1927]" captionTargetId="figure@74.[210,1368,1017,1928]" captionTargetPageId="74" captionText="Fig. 47. Cissidium amboroense Darby, 2015. A. Habitus. B. Pronotum. C. Mesoventrite showing median process of collar, mid-keel and keel, × 1220." figureDoi="http://doi.org/10.5281/zenodo.3777378" httpUri="https://zenodo.org/record/3777378/files/figure.png" pageId="75" pageNumber="76">Fig. 47C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FC83FB36FC1E" blockId="75.[189,1194,876,903]" box="[189,1194,876,903]" pageId="75" pageNumber="76">
METAVENTRITE. Pubescent, length
<quantity id="4CC89B523859FF852B0DFC83FD2CFC1F" box="[580,688,877,903]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="75" pageNumber="76" unit="mm" value="1.1">1.10 mm</quantity>
, width across spines
<quantity id="4CC89B523859FF852AEFFC82FB8EFC1F" box="[934,1042,876,903]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="75" pageNumber="76" unit="mm" value="0.85">0.85 mm</quantity>
, disc simple.
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FC40FC6AFC74" blockId="75.[189,1398,942,1005]" pageId="75" pageNumber="76">
GENITALIA. Female spermatheca globular (
<figureCitation id="130B2A323859FF852BECFC40FC9DFC51" box="[677,769,942,969]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="75" pageNumber="76">Fig. 6B</figureCitation>
). Male aedeagus pointed, not so beak-shaped as many species and with 15 short, thick, blunt setae on ventrite VIII (
<figureCitation id="130B2A323859FF852AC2FC3CFC75FC74" box="[907,1001,978,1004]" captionStart="Fig" captionStartId="8.[189,232,1861,1887]" captionTargetBox="[190,1393,524,1810]" captionTargetId="figure@8.[189,1398,523,1818]" captionTargetPageId="8" captionText="Fig. 7. Metaventral and male ventrite examples. A–B. Metaventrite posterior margin to show spines. A. Cissidium riparium sp. nov., × 610. B. Cissidium plaumanni sp. nov., × 970. C–D. Metaventral discs. C. Cissidium greensladei sp. nov., × 455. D. Cissidium insulare sp. nov., × 595. E. Cissidium obscenum sp. nov., × 840. F–G. Terminal male ventrites. F. Cissidium petri Darby, 2015, × 1580. G. Cissidium amboroense Darby, 2015, × 1380." figureDoi="http://doi.org/10.5281/zenodo.3777296" httpUri="https://zenodo.org/record/3777296/files/figure.png" pageId="75" pageNumber="76">Fig. 7G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3859FF8529F4FBFAFB6CFB18" pageId="75" pageNumber="76" type="discussion">
<paragraph id="8B8F36B73859FF8529F4FBFAFEB3FBB6" blockId="75.[189,303,1044,1070]" box="[189,303,1044,1070]" pageId="75" pageNumber="76">
<heading id="D0C781DB3859FF8529F4FBFAFEB3FBB6" bold="true" box="[189,303,1044,1070]" fontSize="11" level="3" pageId="75" pageNumber="76" reason="3">
<emphasis id="B944EAA53859FF8529F4FBFAFEB3FBB6" bold="true" box="[189,303,1044,1070]" pageId="75" pageNumber="76">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73859FF8529F4FBADFB6CFB18" blockId="75.[189,1399,1091,1153]" pageId="75" pageNumber="76">
The form of antennomere XI with a short median interruption and the sinuate posterior margin of the pronotum before the hind angles distinguish this species from the other Bolivian
<emphasis id="B944EAA53859FF852D3AFB88FB6CFB18" box="[1139,1264,1126,1152]" italics="true" pageId="75" pageNumber="76">
<taxonomicName id="4C304D343859FF852D3AFB88FB70FB18" box="[1139,1260,1126,1152]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">Cissidium</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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