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<document id="C21135024291BB5FE88CD03BB0188622" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13845FF962B25FEE5FC1AFB0B" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium mindanao Darby 2020, sp. nov." docType="treatment" docUuid="05800208-5A5F-48D0-AC96-2616B8A34B4B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="89" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="88" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="5ACDBEF58327D763FEFF11FDF1FC8912">
<mods:title id="38B2474DD66E2CEE6F5462FAC0DC752C">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="5000DC74B8EC71D39685109DD59F7876" type="personal">
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<mods:roleTerm id="D9CD49DB05242D6A76937B7F5967C59C">Author</mods:roleTerm>
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<mods:namePart id="AA97300260C7CF6C63AFDE498AA787D0">Darby, Michael</mods:namePart>
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<mods:title id="D83E02FDD7E051D55CF704A61D5965C2">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BF7C7D7A73C6BCACD94C45B72D55033D">
<mods:date id="39C39131325AB3833103FCE3BC5892EE">2020</mods:date>
<mods:detail id="90A761C9CFB9BCDDF17F25DAF2897A12" type="pubDate">
<mods:number id="A3B4B78E858C07684976A3989AB52FAB">2020-04-02</mods:number>
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<mods:number id="B2630A2ABE77EF9B2BCAE223D9FAAA2C">622</mods:number>
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<mods:classification id="59BE7A18F223E1248355AF3605032E6C">journal article</mods:classification>
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<mods:identifier id="4A24A87A942022DDAEC780C420B5069A" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A13845FF962B25FEE5FC1AFB0B" ID-DOI="http://doi.org/10.5281/zenodo.3795586" ID-GBIF-Taxon="163877860" ID-Zenodo-Dep="3795586" LSID="urn:lsid:zoobank.org:act:05800208-5A5F-48D0-AC96-2616B8A34B4B" httpUri="http://treatment.plazi.org/id/039987A13845FF962B25FEE5FC1AFB0B" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
<subSubSection id="C32A653C3845FF992B25FEE5FB5EFED1" pageId="87" pageNumber="88" type="nomenclature">
<paragraph id="8B8F36B73845FF992B25FEE5FC54FEBE" blockId="87.[369,1218,267,365]" box="[620,968,267,294]" pageId="87" pageNumber="88">
<heading id="D0C781DB3845FF992B25FEE5FC54FEBE" box="[620,968,267,294]" centered="true" fontSize="11" level="2" pageId="87" pageNumber="88" reason="2">
<taxonomicName id="4C304D343845FF992B25FEE5FCFAFEBE" ID-CoL="8XXQV" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[620,870,267,294]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="mindanao" status="sp. nov.">
<emphasis id="B944EAA53845FF992B25FEE5FCFAFEBE" bold="true" box="[620,870,267,294]" italics="true" pageId="87" pageNumber="88">Cissidium mindanao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3845FF992A24FEE2FC54FEBE" box="[877,968,268,294]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B73845FF992838FEC0FB5EFED1" blockId="87.[369,1218,267,365]" box="[369,1218,302,329]" pageId="87" pageNumber="88">
<uri id="FFA13AB53845FF992838FEC0FB5EFED1" box="[369,1218,302,329]" pageId="87" pageNumber="88">
urn:lsid:zoobank.org:act:
<uuid id="FF960C623845FF992BD8FEC0FB5EFED1" box="[657,1218,302,329]" pageId="87" pageNumber="88">05800208-5A5F-48D0-AC96-2616B8A34B4B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3845FF992B9CFEBCFCC2FEF5" box="[725,862,338,365]" pageId="87" pageNumber="88" type="description">
<paragraph id="8B8F36B73845FF992B9CFEBCFCC2FEF5" blockId="87.[369,1218,267,365]" box="[725,862,338,365]" pageId="87" pageNumber="88">
<figureCitation id="130B2A323845FF992B9CFEBCFCA9FEF4" box="[725,821,338,364]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="87" pageNumber="88">Figs 6N</figureCitation>
,
<figureCitation id="130B2A323845FF992A08FEBCFCC2FEF5" box="[833,862,338,365]" captionStart="Fig" captionStartId="87.[189,232,1964,1990]" captionTargetBox="[193,1398,1011,1919]" captionTargetId="figure@87.[189,1398,1005,1919]" captionTargetPageId="87" captionText="Fig. 56. Cissidium mindanao sp. nov. A. Habitus. B. Pronotum, × 430. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660." figureDoi="http://doi.org/10.5281/zenodo.3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="87" pageNumber="88">56</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3845FF9929F4FE7DFE3AFE67" pageId="87" pageNumber="88" type="etymology">
<paragraph id="8B8F36B73845FF9929F4FE7DFED8FE35" blockId="87.[189,324,403,429]" box="[189,324,403,429]" pageId="87" pageNumber="88">
<heading id="D0C781DB3845FF9929F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
<emphasis id="B944EAA53845FF9929F4FE7DFED8FE35" bold="true" box="[189,324,403,429]" pageId="87" pageNumber="88">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73845FF9929F4FE2CFE3AFE67" blockId="87.[189,1399,450,511]" pageId="87" pageNumber="88">
Named after Mindanao, the second largest island in the
<collectingCountry id="F32776273845FF992A18FE2CFC49FE44" box="[849,981,450,476]" name="Philippines" pageId="87" pageNumber="88">Philippines</collectingCountry>
where the specimen was collected. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3845FF9929F4FDC8FE64FD2E" pageId="87" pageNumber="88" type="materials_examined">
<paragraph id="8B8F36B73845FF9929F4FDC8FE36FDD8" blockId="87.[189,426,550,576]" box="[189,426,550,576]" pageId="87" pageNumber="88">
<heading id="D0C781DB3845FF9929F4FDC8FE36FDD8" bold="true" box="[189,426,550,576]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
<emphasis id="B944EAA53845FF9929F4FDC8FE36FDD8" bold="true" box="[189,426,550,576]" pageId="87" pageNumber="88">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA3845FF9929A3FDBBFE64FD2E" ID-GBIF-Occurrence="2609493588" collectedFrom="flight dusk" collectingDate="1946-08-28" collectionCode="FMNH" collectorName="Werner" country="Philippines" elevation="1006" location="Mt McKinley" pageId="87" pageNumber="88" specimenCount="1" stateProvince="Davao Prov." typeStatus="holotype">
<paragraph id="8B8F36B73845FF9929A3FDBBFEC0FDF7" blockId="87.[189,1398,597,694]" box="[234,348,597,623]" pageId="87" pageNumber="88">
<emphasis id="B944EAA53845FF9929A3FDBBFEC0FDF7" bold="true" box="[234,348,597,623]" pageId="87" pageNumber="88">
<typeStatus id="548B88153845FF9929A3FDBBFEC0FDF7" box="[234,348,597,623]" pageId="87" pageNumber="88" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73845FF9929F4FD96FE64FD2E" blockId="87.[189,1398,597,694]" pageId="87" pageNumber="88">
<collectingCountry id="F32776273845FF9929F4FD96FEF6FD0A" box="[189,362,632,658]" name="Philippines" pageId="87" pageNumber="88">PHILIPPINES</collectingCountry>
•
<specimenCount id="9D36FD3E3845FF9928C8FD97FE05FD0B" box="[385,409,633,659]" pageId="87" pageNumber="88" type="male">♂</specimenCount>
; Mindanao, E slope of
<location id="8EEF606C3845FF992BECFD97FCDAFD0A" LSID="urn:lsid:plazi:treatment:039987A13845FF962B25FEE5FC1AFB0B:8EEF606C3845FF992BECFD97FCDAFD0A" box="[677,838,632,659]" country="Philippines" name="Mt McKinley" pageId="87" pageNumber="88" stateProvince="Davao Prov.">Mt McKinley</location>
,
<collectingRegion id="49F4F8553845FF992A19FD97FC78FD0B" box="[848,996,633,659]" country="Philippines" name="Davao" pageId="87" pageNumber="88">Davao Prov.</collectingRegion>
;
<elevation id="001DD1843845FF992AA6FD96FB1DFD0A" box="[1007,1153,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.0058399999999998" pageId="87" pageNumber="88" unit="ft" value="3300.0">
<quantity id="4CC89B523845FF992AA6FD96FBDEFD0A" box="[1007,1090,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.0058399999999998" pageId="87" pageNumber="88" unit="ft" value="3300.0">3300 ft</quantity>
a.s.l.
</elevation>
;
<date id="FF8E10773845FF992DC4FD96FAB1FD0A" box="[1165,1325,632,658]" pageId="87" pageNumber="88" value="1946-08-28">
<collectingDate id="EFCAE99F3845FF992DC4FD96FAB1FD0A" box="[1165,1325,632,658]" pageId="87" pageNumber="88" value="1946-08-28">28 Aug. 1946</collectingDate>
</date>
;
<collectedFrom id="382C754A3845FF992C71FD96FF64FD2D" pageId="87" pageNumber="88">flight dusk</collectedFrom>
;
<collectorName id="26C553613845FF99284AFD72FEC5FD2E" box="[259,345,668,694]" pageId="87" pageNumber="88">Werner</collectorName>
leg.;
<collectionCode id="ED21AE723845FF9928D2FD72FE68FD2E" box="[411,500,668,694]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="87" pageNumber="88" type="Museum">FMNH</collectionCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C32A653C3845FF9629F4FD33FCCFFC64" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88" type="description">
<paragraph id="8B8F36B73845FF9929F4FD33FED2FD6F" blockId="87.[189,334,733,759]" box="[189,334,733,759]" pageId="87" pageNumber="88">
<heading id="D0C781DB3845FF9929F4FD33FED2FD6F" bold="true" box="[189,334,733,759]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
<emphasis id="B944EAA53845FF9929F4FD33FED2FD6F" bold="true" box="[189,334,733,759]" pageId="87" pageNumber="88">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73845FF9929F4FCE5FD36FCBE" blockId="87.[189,682,779,806]" box="[189,682,779,806]" pageId="87" pageNumber="88">
SIZE. Habitus (
<figureCitation id="130B2A323845FF99282EFCE5FE48FCBE" box="[359,468,779,806]" captionStart="Fig" captionStartId="87.[189,232,1964,1990]" captionTargetBox="[193,1398,1011,1919]" captionTargetId="figure@87.[189,1398,1005,1919]" captionTargetPageId="87" captionText="Fig. 56. Cissidium mindanao sp. nov. A. Habitus. B. Pronotum, × 430. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660." figureDoi="http://doi.org/10.5281/zenodo.3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="87" pageNumber="88">Fig. 56A</figureCitation>
), length
<quantity id="4CC89B523845FF992B73FCE5FD39FCBE" box="[570,677,779,806]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="87" pageNumber="88" unit="mm" value="0.67">0.67 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73845FF9629F4FCA3FB18FED2" blockId="87.[189,978,845,871]" lastBlockId="88.[189,1398,267,330]" lastPageId="88" lastPageNumber="89" pageId="87" pageNumber="88">
COLOUR. Dark brown, pubescence, legs and antennae dusky yellow. HEAD. Densely setose; antennomeres III–XI length
<quantity id="4CC89B52384AFF962A7BFEE5FC3EFEBE" box="[818,930,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="88" pageNumber="89" unit="mm" value="0.37">0.37 mm</quantity>
, III–IX length
<quantity id="4CC89B52384AFF962D17FEE5FB53FEBE" box="[1118,1231,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="88" pageNumber="89" unit="mm" value="0.2">0.20 mm</quantity>
, X–XI length
<quantity id="4CC89B52384AFF9629F4FEDEFEB5FED2" box="[189,297,304,330]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="88" pageNumber="89" unit="mm" value="0.17">0.17 mm</quantity>
; width across eyes
<quantity id="4CC89B52384AFF962B42FEDEFDEBFED2" box="[523,631,304,330]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="88" pageNumber="89" unit="mm" value="0.22">0.22 mm</quantity>
; mentum parallel-sided, otherwise obscured.
</paragraph>
<caption id="DF4F663F3845FF9929F4F842FCF6F871" ID-DOI="http://doi.org/10.5281/zenodo.3777396" ID-Zenodo-Dep="3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="87" pageNumber="88" startId="87.[189,232,1964,1990]" targetBox="[193,1398,1011,1919]" targetPageId="87">
<paragraph id="8B8F36B73845FF9929F4F842FCF6F871" blockId="87.[189,1398,1964,2025]" pageId="87" pageNumber="88">
<emphasis id="B944EAA53845FF9929F4F842FE84F85E" bold="true" box="[189,280,1964,1990]" pageId="87" pageNumber="88">Fig. 56.</emphasis>
<taxonomicName id="4C304D343845FF992857F842FD93F85E" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[286,527,1964,1990]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="mindanao" status="sp. nov.">
<emphasis id="B944EAA53845FF992857F842FD93F85E" box="[286,527,1964,1990]" italics="true" pageId="87" pageNumber="88">Cissidium mindanao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3845FF992B5CF842FDF3F85E" box="[533,623,1964,1990]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B944EAA53845FF992B3CF842FD16F85E" bold="true" box="[629,650,1964,1990]" pageId="87" pageNumber="88">A</emphasis>
. Habitus.
<emphasis id="B944EAA53845FF992A49F842FC88F85E" bold="true" box="[768,788,1964,1990]" pageId="87" pageNumber="88">B</emphasis>
. Pronotum, × 430.
<emphasis id="B944EAA53845FF992ABAF842FB94F85E" bold="true" box="[1011,1032,1964,1990]" pageId="87" pageNumber="88">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660.
</paragraph>
</caption>
<paragraph id="8B8F36B7384AFF9629F4FE9DFC9FFE62" blockId="88.[189,1399,370,506]" pageId="88" pageNumber="89">
PRONOTUM. Length
<quantity id="4CC89B52384AFF9628EAFE9CFD8DFE15" box="[419,529,370,397]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="88" pageNumber="89" unit="mm" value="0.21">0.21 mm</quantity>
, width
<quantity id="4CC89B52384AFF962B22FE9CFD45FE15" box="[619,729,370,397]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="88" pageNumber="89" unit="mm" value="0.3">0.30 mm</quantity>
, densely pubescent and foveolate, without foveae along basal margin; sides sharply angled, concave to rectangular hind angles slightly concave to obtuse front angles, bordered, the border continuing along the entire posterior margin including the sinuous emargination in front of the scutellum (
<figureCitation id="130B2A32384AFF962BC3FE31FD6AFE62" box="[650,758,479,506]" captionStart="Fig" captionStartId="87.[189,232,1964,1990]" captionTargetBox="[193,1398,1011,1919]" captionTargetId="figure@87.[189,1398,1005,1919]" captionTargetPageId="87" captionText="Fig. 56. Cissidium mindanao sp. nov. A. Habitus. B. Pronotum, × 430. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660." figureDoi="http://doi.org/10.5281/zenodo.3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="88" pageNumber="89">Fig. 56B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FDCCFE02FDF9" blockId="88.[189,1398,546,609]" pageId="88" pageNumber="89">
ELYTRA. Length
<quantity id="4CC89B52384AFF9628C9FDCCFE73FDA4" box="[384,495,546,572]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="88" pageNumber="89" unit="mm" value="0.49">0.49 mm</quantity>
, width
<quantity id="4CC89B52384AFF962B03FDCCFD25FDA4" box="[586,697,546,572]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="88" pageNumber="89" unit="mm" value="0.4">0.40 mm</quantity>
, pubescent without distinct foveolae markedly contrasting with the pronotum.
</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FD67FE02FCAD" blockId="88.[189,1399,649,821]" pageId="88" pageNumber="89">
MESOVENTRITE. Median process of mesoventral collar narrow, parallel-sided; mid-keel anterior angles joining collar, posterior angles not reaching mesocoxal anterior borders, raised medially at junction with keel; keel with ± eight setae, parallel-sided, with rounded termination at the interruption of the mesocoxal borders; mesoventral lateral margins angled below humeri, serrate posteriorly; humeri toothed (
<figureCitation id="130B2A32384AFF96286CFCF4FE0DFCAD" box="[293,401,794,821]" captionStart="Fig" captionStartId="87.[189,232,1964,1990]" captionTargetBox="[193,1398,1011,1919]" captionTargetId="figure@87.[189,1398,1005,1919]" captionTargetPageId="87" captionText="Fig. 56. Cissidium mindanao sp. nov. A. Habitus. B. Pronotum, × 430. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 660." figureDoi="http://doi.org/10.5281/zenodo.3777396" httpUri="https://zenodo.org/record/3777396/files/figure.png" pageId="88" pageNumber="89">Fig. 56C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FCB3FB1FFCEF" blockId="88.[189,1155,861,887]" box="[189,1155,861,887]" pageId="88" pageNumber="89">
METAVENTRITE. Length
<quantity id="4CC89B52384AFF962881FCB3FDA8FCEF" box="[456,564,861,887]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="88" pageNumber="89" unit="mm" value="0.18">0.18 mm</quantity>
, setose, disc simple, width across spines
<quantity id="4CC89B52384AFF962D5DFCB3FBE3FCEF" box="[1044,1151,861,887]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="88" pageNumber="89" unit="mm" value="0.11">0.11 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FC4EFE5EFC22" blockId="88.[189,450,928,954]" box="[189,450,928,954]" pageId="88" pageNumber="89">WINGS. Macropterous.</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FC0CFCCFFC64" blockId="88.[189,851,994,1021]" box="[189,851,994,1021]" pageId="88" pageNumber="89">
GENITALIA. Male aedeagus (
<figureCitation id="130B2A32384AFF9628B7FC0CFDC1FC65" box="[510,605,994,1021]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="88" pageNumber="89">Fig. 6N</figureCitation>
). Female not known.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C384AFF9629F4FBCBFC1AFB0B" pageId="88" pageNumber="89" type="discussion">
<paragraph id="8B8F36B7384AFF9629F4FBCBFEB3FBA7" blockId="88.[189,303,1061,1087]" box="[189,303,1061,1087]" pageId="88" pageNumber="89">
<heading id="D0C781DB384AFF9629F4FBCBFEB3FBA7" bold="true" box="[189,303,1061,1087]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="B944EAA5384AFF9629F4FBCBFEB3FBA7" bold="true" box="[189,303,1061,1087]" pageId="88" pageNumber="89">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7384AFF9629F4FBBAFC1AFB0B" blockId="88.[189,1398,1108,1171]" pageId="88" pageNumber="89">
One of the only two species of
<taxonomicName id="4C304D34384AFF962B18FBBAFD76FBF6" authority="in" authorityName="In" box="[593,746,1108,1134]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA5384AFF962B18FBBAFD5AFBF6" box="[593,710,1108,1134]" italics="true" pageId="88" pageNumber="89">Cissidium</emphasis>
in
</taxonomicName>
this group from the
<collectingCountry id="F3277627384AFF962ABFFBBAFBE2FBF6" box="[1014,1150,1108,1134]" name="Philippines" pageId="88" pageNumber="89">Philippines</collectingCountry>
. Distinguished from
<taxonomicName id="4C304D34384AFF9629F4FB97FEA6FB0A" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[189,314,1145,1171]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="werneri" status="sp. nov.">
<emphasis id="B944EAA5384AFF9629F4FB97FEA6FB0A" box="[189,314,1145,1171]" italics="true" pageId="88" pageNumber="89">C. werneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE384AFF96280BFB97FE00FB0B" box="[322,412,1145,1171]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
by the more densely foveolate pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>
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