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<document id="1DE9BFD8F9EFE83595D3AD59EF3B9B6B" ID-CLB-Dataset="45456" ID-DOI="10.11646/zootaxa.3815.4.4" ID-GBIF-Dataset="b2e39573-c2df-4ad5-9dec-93eb67d58e6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="228698" ID-ZooBank="172C594B-7321-4F1C-B8CC-158A195A7D73" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438354307" checkinUser="plazi" docAuthor="Väisänen, Rauno" docDate="2014" docId="03998794FFFFF706FF4067CDFAD7FE6C" docLanguage="en" docName="zt03815p540.pdf" docOrigin="Zootaxa 3815 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mycomya shimai Väisänen, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="537" masterDocId="FFA0FFECFFF5F70DFFD76339FFB3FF99" masterDocTitle="New Mycomya species from South-East Asia (Diptera, Mycetophilidae)" masterLastPageNumber="540" masterPageNumber="526" pageNumber="536" updateTime="1698332543223" updateUser="ExternalLinkService">
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<mods:title id="806DFABA57A24671325B45686EBEFD0B">New Mycomya species from South-East Asia (Diptera, Mycetophilidae)</mods:title>
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<mods:namePart id="4579D09035329E5D0B88B66F799A1311">Väisänen, Rauno</mods:namePart>
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<mods:date id="ACECDB141323C2F0C86222F027F0F873">2014</mods:date>
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<mods:identifier id="AD2B8167208F58B9C47DDEC03A044194" type="DOI">10.11646/zootaxa.3815.4.4</mods:identifier>
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<treatment id="03998794FFFFF706FF4067CDFAD7FE6C" ID-DOI="http://doi.org/10.5281/zenodo.6134646" ID-GBIF-Taxon="119507175" ID-Zenodo-Dep="6134646" LSID="urn:lsid:plazi:treatment:03998794FFFFF706FF4067CDFAD7FE6C" httpUri="http://treatment.plazi.org/id/03998794FFFFF706FF4067CDFAD7FE6C" lastPageId="11" lastPageNumber="537" pageId="10" pageNumber="536">
<subSubSection id="C32A6509FFFFF707FF4067CDFEAAFAA9" pageId="10" pageNumber="536" type="nomenclature">
<paragraph id="8B8F3682FFFFF707FF4067CDFE11FA94" blockId="10.[151,418,1267,1328]" box="[151,418,1267,1293]" pageId="10" pageNumber="536">
<heading id="D0C781EEFFFFF707FF4067CDFE11FA94" bold="true" box="[151,418,1267,1293]" fontSize="10" level="2" pageId="10" pageNumber="536" reason="2">
<emphasis id="B944EA90FFFFF707FF4067CDFE11FA94" bold="true" box="[151,418,1267,1293]" pageId="10" pageNumber="536">
<taxonomicName id="4C304D01FFFFF707FF4067CDFEEFFA95" box="[151,348,1267,1292]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="536" phylum="Arthropoda" rank="species" species="shimai" status="sp. nov.">
<emphasis id="B944EA90FFFFF707FF4067CDFEEFFA95" bold="true" box="[151,348,1267,1292]" italics="true" pageId="10" pageNumber="536">Mycomya shimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757EBFFFFF707FEB467CDFE11FA94" box="[355,418,1268,1293]" pageId="10" pageNumber="536" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F3682FFFFF707FF40662EFEAAFAA9" blockId="10.[151,418,1267,1328]" box="[151,281,1303,1328]" pageId="10" pageNumber="536">
<figureCitation id="130B2A07FFFFF707FF40662EFF51FAA9" box="[151,226,1303,1328]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Figs. 7</figureCitation>
A–H
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFFFF707FF406666FDB8FA05" pageId="10" pageNumber="536" type="materials_examined">
<paragraph id="8B8F3682FFFFF707FF406666FDB8FA05" blockId="10.[151,1437,1375,2024]" pageId="10" pageNumber="536">
<emphasis id="B944EA90FFFFF707FF406666FEDAFAE1" bold="true" box="[151,361,1375,1400]" pageId="10" pageNumber="536">Material studied.</emphasis>
<emphasis id="B944EA90FFFFF707FEA36659FE69FAE1" box="[372,474,1376,1400]" italics="true" pageId="10" pageNumber="536">
<typeStatus id="548B8820FFFFF707FEA36659FE69FAE1" box="[372,474,1376,1400]" pageId="10" pageNumber="536" type="holotype">Holotype</typeStatus>
</emphasis>
. ♂.
<collectingCountry id="F3277612FFFFF707FDC46659FD1DFAE1" box="[531,686,1376,1400]" name="Indonesia" pageId="10" pageNumber="536">INDONESIA</collectingCountry>
, Java, Puncak
<quantity id="4CC89B67FFFFF707FC886659FC08FAE1" box="[863,955,1376,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="536" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FF8E1042FFFFF707FC1C6659FBDEFAE1" box="[971,1133,1376,1400]" pageId="10" pageNumber="536" value="1973-11-14">14 Nov. 1973</date>
. M. Shima (KUC).
<emphasis id="B944EA90FFFFF707FA8C6659FF4EFA05" italics="true" pageId="10" pageNumber="536">Other material.</emphasis>
Same data, 1 ♀ (KUC).
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFFFF706FF10669EFD84FF61" lastPageId="11" lastPageNumber="537" pageId="10" pageNumber="536" type="description">
<paragraph id="8B8F3682FFFFF707FF10669EFA2FF87E" blockId="10.[151,1437,1375,2024]" pageId="10" pageNumber="536">
<emphasis id="B944EA90FFFFF707FF10669EFEE2FA59" bold="true" box="[199,337,1447,1472]" pageId="10" pageNumber="536">Description</emphasis>
. ♂.
<emphasis id="B944EA90FFFFF707FE56669EFE7BFA59" bold="true" box="[385,456,1447,1472]" pageId="10" pageNumber="536">Head.</emphasis>
Palp, other mouthparts and face yellow, posterior parts of head light brownish. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. 1st flagellomere about
<date id="FF8E1042FFFFF707FB9B66F5FBDBFA7D" box="[1100,1128,1484,1508]" pageId="10" pageNumber="536">3x</date>
its width, 2nd flagellomere
<date id="FF8E1042FFFFF707FF4066C9FF00F991" box="[151,179,1520,1544]" pageId="10" pageNumber="536">2x</date>
its width.
<emphasis id="B944EA90FFFFF707FEF566D6FE31F991" bold="true" box="[290,386,1519,1544]" pageId="10" pageNumber="536">Thorax.</emphasis>
Pronotum yellow, with 4 long setae. Scutum yellow with 3 brownish longitudinal stripes. Anepisternum and preepisternum yellow. Scutellum yellowish, with 4 long setae. Laterotergite yellowish. Mediotergite yellowish, bare.
<emphasis id="B944EA90FFFFF707FE3D650EFD82F9C9" bold="true" box="[490,561,1591,1616]" pageId="10" pageNumber="536">Wing.</emphasis>
Length 4.0 mm. Wing hyaline. Sc ending in C near middle of small cell, Sc2 ending in R1 distad of middle of small cell. Apical part of Sc bearing 10 small setae. Small cell about 1.5x as long as wide. Cu fork below M fork. M ratios: 0.48, 0.60. Cu ratios: 0.87, 1.33. Small setae: M petiole: 0; M1: +; M2: +; Cu petiole: +; Cu1: +; Cu2: +. Halter pale yellowish, apex brownish.
<emphasis id="B944EA90FFFFF707FC796595FC5EF95C" bold="true" box="[942,1005,1708,1733]" pageId="10" pageNumber="536">Legs.</emphasis>
Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 with relatively short, shorter than half of coxa 2, curved spur. Spur with 1 small apical tooth. Leg ratios: bt1:t1 =? (structure missing), bt2:t2 = 0.70, bt3:t3 = 0.42.
<emphasis id="B944EA90FFFFF707FBE065CEFB01F889" bold="true" box="[1079,1202,1783,1808]" pageId="10" pageNumber="536">Abdomen.</emphasis>
Tergites 1–5 yellow, with brown dorsal line and posterior margins, 6–7 brownish to brown. Sternites 1–5 yellow, 6–7 brownish.
<emphasis id="B944EA90FFFFF707FF406406FE8AF8C1" bold="true" box="[151,313,1855,1880]" pageId="10" pageNumber="536">Hypopygium.</emphasis>
<figureCitation id="130B2A07FFFFF707FE966406FE3EF8C1" box="[321,397,1855,1880]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Figs. 7</figureCitation>
A–H, yellow. Tergal fork almost H-shaped, its apex about 1.4x as wide as its base, branches almost parallel, very thin, about
<date id="FF8E1042FFFFF707FDC9645AFDFBF8E2" box="[542,584,1891,1915]" pageId="10" pageNumber="536" value="2010-10">10x</date>
as long as wide, with short apical setae, and with lateral lobes curved backwards (
<figureCitation id="130B2A07FFFFF707FEC864BEFEDEF806" box="[287,365,1927,1952]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Figs. 7</figureCitation>
C–D). Sternal lateral appendages with some flattened setae (
<figureCitation id="130B2A07FFFFF707FBCC64BEFBECF806" box="[1051,1119,1927,1952]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Fig. 7</figureCitation>
F). Gonostylus bilobed with two long, very slender branches, shorter branch straight, about
<date id="FF8E1042FFFFF707FCBF6495FC21F85D" box="[872,914,1964,1988]" pageId="10" pageNumber="536" value="2010-10">10x</date>
as long as its width and about ½ of length of longer, curved branch, which is about
<date id="FF8E1042FFFFF707FD9364F6FDDDF871" box="[580,622,1999,2024]" pageId="10" pageNumber="536" value="1915-10">15x</date>
as long as its width (
<figureCitation id="130B2A07FFFFF707FC8A64F6FC13F87E" box="[861,928,1999,2024]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Fig. 7</figureCitation>
G). Aedeagus with two short spurs (
<figureCitation id="130B2A07FFFFF707FAE264F6FACBF87E" box="[1333,1400,1999,2024]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="10" pageNumber="536">Fig. 7</figureCitation>
H).
</paragraph>
<paragraph id="8B8F3682FFFEF706FF4063AEFD84FF61" blockId="11.[151,1436,151,501]" pageId="11" pageNumber="537">
<emphasis id="B944EA90FFFEF706FF4063AEFF46FF29" bold="true" box="[151,245,151,176]" pageId="11" pageNumber="537">Female.</emphasis>
Wing length
<quantity id="4CC89B67FFFEF706FE4663A1FE5FFF36" box="[401,492,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="537" unit="mm" value="4.2">4.2 mm</quantity>
. Thorax yellow to yellowish. Colouration of abdomen as in ♂. Leg ratios: bt1:t1 = 1.09, bt2:t2 = 0.73, bt3:t3 = 0.62. Terminalia yellow. Hypogynal valve triangular, apically tapered, with several setae. Apical segment of cercus oval.
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFFEF706FF10623DFC67FE49" pageId="11" pageNumber="537" type="discussion">
<paragraph id="8B8F3682FFFEF706FF10623DFC67FE49" blockId="11.[151,1436,151,501]" pageId="11" pageNumber="537">
<emphasis id="B944EA90FFFEF706FF10623DFEFFFE84" bold="true" box="[199,332,260,285]" pageId="11" pageNumber="537">Discussion.</emphasis>
<taxonomicName id="4C304D01FFFEF706FE85623CFDA2FE85" box="[338,529,260,284]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="537" phylum="Arthropoda" rank="species" species="shimai">
<emphasis id="B944EA90FFFEF706FE85623CFDA2FE85" box="[338,529,260,284]" italics="true" pageId="11" pageNumber="537">Mycomya shimai</emphasis>
</taxonomicName>
belongs to the subgenus
<taxonomicName id="4C304D01FFFEF706FCFF623DFC7BFE85" box="[808,968,260,284]" class="Insecta" family="Mycetophilidae" genus="Calomycomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA90FFFEF706FCFF623DFC7BFE85" box="[808,968,260,284]" italics="true" pageId="11" pageNumber="537">Calomycomya</emphasis>
</taxonomicName>
(based on the presence of the curved spur of the coxa 2 and the tergal fork of the hypopygium) and is the only known representative of the subgenus from the South East Asian islands.
<taxonomicName id="4C304D01FFFEF706FE0D6275FDC9FEFD" box="[474,634,332,356]" class="Insecta" family="Mycetophilidae" genus="Calomycomya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA90FFFEF706FE0D6275FDC9FEFD" box="[474,634,332,356]" italics="true" pageId="11" pageNumber="537">Calomycomya</emphasis>
</taxonomicName>
is a widely distributed subgenus in the Palaearctic, Nearctic and Neotropical regions, with three species known from the Himalayas (
<bibRefCitation id="EFA14B73FFFEF706FC656256FBC3FE11" author="Vaisanen" box="[946,1136,367,392]" pageId="11" pageNumber="537" refString="Vaisanen, R. (2013 a) New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 2. Subgenera Calomycomya, Cymomya, Neomycomya and Pavomya subg. n. Zootaxa, 3666 (3), 301 - 318. http: // dx. doi. org / 10.11646 / zootaxa. 3666.3.1" type="journal article" year="2013" yearSuffix="a">Väisänen 2013a</bibRefCitation>
). It differs from the other species of the subgenus in the almost H-shaped tergal fork with lateral lobes curved inwards (
<figureCitation id="130B2A07FFFEF706FB6662ADFB4DFE35" box="[1201,1278,404,429]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="11" pageNumber="537">Figs. 7</figureCitation>
A, 7C–D) and in its characteristic gonostylus, with two long, slender branches (
<figureCitation id="130B2A07FFFEF706FCBA628EFC03FE49" box="[877,944,439,464]" captionStart="FIGURE 7" captionStartId="11.[151,250,1940,1962]" captionTargetBox="[198,1390,555,1909]" captionTargetId="figure@11.[186,1401,547,1918]" captionTargetPageId="11" captionText="FIGURE 7. Mycomya shimai sp. n., male hypopygium. A – B tergal and sternal views, C tergal fork and associated structures, lateral view, D tergal fork, tergal view, twice enlarged, E detail of tergite 9, twice enlarged, F gonostylus, twice enlarged, G apex of sternal lateral appendage, twice enlarged, H apex of aedeagus, twice enlarged. Scale 0.2 mm." httpUri="https://zenodo.org/record/228705/files/figure.png" pageId="11" pageNumber="537">Fig. 7</figureCitation>
G).
</paragraph>
</subSubSection>
<subSubSection id="C32A6509FFFEF706FF1062E5FAD7FE6C" box="[199,1380,476,501]" pageId="11" pageNumber="537" type="etymology">
<paragraph id="8B8F3682FFFEF706FF1062E5FAD7FE6C" blockId="11.[151,1436,151,501]" box="[199,1380,476,501]" pageId="11" pageNumber="537">
<emphasis id="B944EA90FFFEF706FF1062E5FEFDFE6C" bold="true" box="[199,334,476,501]" pageId="11" pageNumber="537">Etymology.</emphasis>
The species is named after the Japanese entomologist M. Shima, who collected the
<typeStatus id="548B8820FFFEF706FB2C62E5FAD2FE6C" box="[1275,1377,476,501]" pageId="11" pageNumber="537" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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