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<document id="1988E5537095F5FA023CE66D084C2110" ID-DOI="10.11646/phytotaxa.292.1.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="13698007" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725569228876" checkinUser="felipe" docAuthor="Dittrich, Vinícius Antonio De Oliveira, Smith, Alan R. &amp; Gasper, André Luís De" docDate="2017" docId="039965528C25FFAB31F7FD048659F881" docLanguage="en" docName="phytotaxa.292.1.6.pdf" docOrigin="Phytotaxa 292 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.292.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Parablechnum paucipinna A. R. Sm. 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="66" masterDocId="FFA01D2A8C24FFAA317FFFF0833DFFEA" masterDocTitle="Parablechnum roraimense and P. paucipinna spp. nov. (Blechnaceae: Polypodiopsida), lectotypification of P. stuebelii, and citation corrections in the family" masterLastPageNumber="73" masterPageNumber="65" pageNumber="66" updateTime="1725574401031" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EA75655AA6CBBB0BD6DA1FEF3E16D06D">Parablechnum roraimense and P. paucipinna spp. nov. (Blechnaceae: Polypodiopsida), lectotypification of P. stuebelii, and citation corrections in the family</mods:title>
</mods:titleInfo>
<mods:name id="006B54B4081C5F7AB7E278110479BE74" type="personal">
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<mods:namePart id="6BB3571F40F726842BE2471C65AD848A">Dittrich, Vinícius Antonio De Oliveira</mods:namePart>
<mods:affiliation id="30D6CF6647F521D8F0F3D9C5B994C61C">Departamento de Botânica, Instituto de Ciências Biológicas, Universidade Federal de Juiz de Fora, Rua José Lourenço Kelmer, s. n.</mods:affiliation>
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<mods:namePart id="DA6020FCFC8BE61CE33D5688BF35ACBE">Smith, Alan R.</mods:namePart>
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<mods:role id="C80FDF8355C20F75F88D243638D18EFD">
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<mods:namePart id="4297A32904A942034A6BE3B2CCE1CD64">Gasper, André Luís De</mods:namePart>
<mods:affiliation id="2822804035919A5AE1F061B7DFC59F6F">Departamento de Botânica, Instituto de Ciências Biológicas, Universidade Federal de Juiz de Fora, Rua José Lourenço Kelmer, s. n. &amp; Departamento de Botânica, Instituto de Ciências Biológicas, Universidade Federal de Juiz de Fora, Rua José Lourenço Kelmer, s. n.</mods:affiliation>
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<mods:title id="AF5D80CC71BAAD08BB1A5E727B6D8645">Phytotaxa</mods:title>
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<mods:part id="AB67F2E2111A9C836B2047D928800B80">
<mods:date id="99FD1D00BEFA1A4A8AA72A96A6E013B8">2017</mods:date>
<mods:detail id="3D968C3C11C53C2D0ACEBBBD74370145" type="pubDate">
<mods:number id="10A6EB5BF7C6A7696C7756728DE675F6">2017-01-20</mods:number>
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<mods:identifier id="AF9266F46AD5ADFB6B83BCE0CC845EB6" type="DOI">10.11646/phytotaxa.292.1.6</mods:identifier>
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<treatment id="039965528C25FFAB31F7FD048659F881" ID-DOI="http://doi.org/10.5281/zenodo.13698037" ID-Zenodo-Dep="13698037" LSID="urn:lsid:plazi:treatment:039965528C25FFAB31F7FD048659F881" httpUri="http://treatment.plazi.org/id/039965528C25FFAB31F7FD048659F881" lastPageNumber="66" pageId="1" pageNumber="66">
<subSubSection id="C32A87CF8C25FFAB31F7FD0481DCFCE5" box="[136,737,756,783]" pageId="1" pageNumber="66" type="nomenclature">
<paragraph id="8B8FD4448C25FFAB31F7FD0481DCFCE5" blockId="1.[136,737,756,783]" box="[136,737,756,783]" pageId="1" pageNumber="66">
<heading id="D0C763288C25FFAB31F7FD0481DCFCE5" box="[136,737,756,783]" fontSize="11" level="2" pageId="1" pageNumber="66" reason="3">
<taxonomicName id="4C30AFC78C25FFAB31F7FD04811BFCE5" ID-CoL="4CMGQ" authority="A.R.Sm." authorityName="A. R. Sm." authorityYear="2017" box="[136,550,756,783]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="paucipinna" status="sp. nov.">
<emphasis id="B94408568C25FFAB31F7FD048280FCE4" bold="true" box="[136,445,756,783]" italics="true" pageId="1" pageNumber="66">Parablechnum paucipinna</emphasis>
A.R.Sm.
</taxonomicName>
,
<emphasis id="B94408568C25FFAB334DFD0681BAFCE5" box="[562,647,758,783]" italics="true" pageId="1" pageNumber="66">
<taxonomicNameLabel id="A277B52D8C25FFAB334DFD0681BAFCE5" box="[562,647,758,783]" pageId="1" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130BC8C18C25FFAB33EAFD0681EBFCE5" box="[661,726,758,783]" captionStart="FIGURE 1" captionStartId="2.[136,229,1955,1976]" captionTargetBox="[189,1398,190,1930]" captionTargetId="figure-15@2.[189,1399,190,1931]" captionTargetPageId="2" captionText="FIGURE 1. Photograph of the holotype of Parablechnum paucipinna (Y. Vivas, B. Holst, J. Tillet &amp; S. Fawcett 1553, UC)." figureDoi="http://doi.org/10.5281/zenodo.13698013" httpUri="https://zenodo.org/record/13698013/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A87CF8C25FFAB31F7FCB0800CF985" pageId="1" pageNumber="66" type="materials_examined">
<paragraph id="8B8FD4448C25FFAB31F7FCB0804DFC77" blockId="1.[136,1452,832,925]" pageId="1" pageNumber="66">
<emphasis id="B94408568C25FFAB31F7FCB083E7FCBF" bold="true" box="[136,218,832,853]" pageId="1" pageNumber="66">Type:—</emphasis>
<collectingCountry id="F32794D48C25FFAB31A5FCB08251FCBF" box="[218,364,832,853]" name="Venezuela" pageId="1" pageNumber="66">VENEZUELA</collectingCountry>
. Edo.
<collectingRegion id="49F41AA68C25FFAB30D4FCB082CAFCBF" box="[427,503,832,853]" country="Venezuela" name="Bolivar" pageId="1" pageNumber="66">Bolívar</collectingRegion>
: Sierra de Lema, sector La Escalera, al Este de la carretera, a unos
<quantity id="4CC879A18C25FFAB35EBFCB087EEFCBF" box="[1172,1235,832,853]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="66" unit="m" value="200.0">200 m</quantity>
del salto El Danto en dirección N-S,
<geoCoordinate id="EE04B2838C25FFAB3031FC9482FBFC93" box="[334,454,868,889]" degrees="05" direction="north" minutes="57" orientation="latitude" pageId="1" pageNumber="66" precision="15" seconds="54" value="5.9649997">05°5754”N</geoCoordinate>
,
<geoCoordinate id="EE04B2838C25FFAB30AFFC948176FC93" box="[464,587,868,889]" degrees="61" direction="west" minutes="23" orientation="longitude" pageId="1" pageNumber="66" precision="15" seconds="42" value="-61.395">61°2342”W</geoCoordinate>
, bosque de galería muy húmedo,
<quantity id="4CC879A18C25FFAB32F5FC94872DFC93" box="[906,1040,868,889]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="1" pageNumber="66" unit="m" value="1275.0" valueMax="1300.0" valueMin="1250.0">12501300 m</quantity>
,
<date id="FF8EF2848C25FFAB3566FC9487AFFC93" box="[1049,1170,868,889]" pageId="1" pageNumber="66" value="2006-04-23">23 Apr 2006</date>
,
<emphasis id="B94408568C25FFAB35E3FC948264FC77" italics="true" pageId="1" pageNumber="66">
Y. Vivas, B. Holst, J. Tillet &amp;
<collectionCode id="ED214C818C25FFAB31C2FC7883F1FC77" box="[189,204,904,925]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="66" type="Herbarium">S</collectionCode>
. Fawcett 1553
</emphasis>
(
<typeStatus id="548B6AE68C25FFAB301AFC788286FC77" box="[357,443,904,925]" pageId="1" pageNumber="66" type="holotype">holotype</typeStatus>
UC-2044673,
<typeStatus id="548B6AE68C25FFAB3335FC7881A4FC77" box="[586,665,904,925]" pageId="1" pageNumber="66" type="isotype">isotypes</typeStatus>
SEL, VEN, not seen).
</paragraph>
<paragraph id="8B8FD4448C25FFAB31F7FC3E800CF985" blockId="1.[136,1452,974,1899]" pageId="1" pageNumber="66">
<emphasis id="B94408568C25FFAB31F7FC3E83EDFC0D" box="[136,208,974,999]" italics="true" pageId="1" pageNumber="66">Plants</emphasis>
rupicolous;
<emphasis id="B94408568C25FFAB3024FC3E82FDFC0D" box="[347,448,974,999]" italics="true" pageId="1" pageNumber="66">rhizomes</emphasis>
compact, concealed by stipe bases and root mass, lacking stolons, caudex
<emphasis id="B94408568C25FFAB3580FC3E861CFC0D" box="[1279,1313,974,999]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
<quantity id="4CC879A18C25FFAB3458FC3E8663FC0D" box="[1319,1374,974,999]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="66" unit="cm" value="2.0">2 cm</quantity>
diam.; rhizome apex scales concolorous, tan to brownish, basifixed or nearly so, lanceolate to ovate-lanceolate with a caudate apex, 810 ×
<quantity id="4CC879A18C25FFAB305BFBE682A4FBC5" box="[292,409,1046,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="1" pageNumber="66" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">12.5 mm</quantity>
, margins entire or nearly so;
<emphasis id="B94408568C25FFAB339AFBE68016FBC5" box="[741,811,1046,1071]" italics="true" pageId="1" pageNumber="66">leaves</emphasis>
clustered, ca.
<quantity id="4CC879A18C25FFAB32B0FBE68728FBC5" box="[975,1045,1046,1071]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="66" unit="cm" value="35.0">35 cm</quantity>
long, the fertile and sterile strongly dimorphic, but similar in length;
<emphasis id="B94408568C25FFAB3082FBCA8100FBB9" box="[509,573,1082,1107]" italics="true" pageId="1" pageNumber="66">stipes</emphasis>
stramineous, or darkening with age, glabrous above the basal
<quantity id="4CC879A18C25FFAB3583FBCA866FFBB9" box="[1276,1362,1082,1107]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="66" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">23 cm</quantity>
, sulcate adaxially, with conspicuous transverse wrinkles throughout, the sterile to
<quantity id="4CC879A18C25FFAB32BAFBAE8736FB9D" box="[965,1035,1118,1143]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="66" unit="cm" value="20.0">20 cm</quantity>
×
<quantity id="4CC879A18C25FFAB3555FBAE87B7FB9D" box="[1066,1162,1118,1143]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="66" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, about the same length as the blades, the fertile to
<quantity id="4CC879A18C25FFAB30ECFB7282E6FB71" box="[403,475,1154,1179]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="66" unit="cm" value="25.0">25 cm</quantity>
, twice as long as blades, the basal scales to
<emphasis id="B94408568C25FFAB32C1FB7280E2FB71" box="[958,991,1154,1179]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
6 ×
<quantity id="4CC879A18C25FFAB356FFB72876FFB71" box="[1040,1106,1154,1179]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="66" unit="mm" value="2.0">2 mm</quantity>
, entire, lacking marginal setae, not or only weakly pectinate at bases, similar to those of rhizome apices but lighter tan;
<emphasis id="B94408568C25FFAB3523FB568798FB55" box="[1116,1189,1190,1215]" italics="true" pageId="1" pageNumber="66">blades</emphasis>
chartaceous, the sterile ca. 20 ×
<quantity id="4CC879A18C25FFAB3193FB3A8208FB09" box="[236,309,1226,1251]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="66" unit="cm" value="15.0">15 cm</quantity>
, deltate, lowest pinnae the longest or nearly so (lacking vestigial proximal pinnae), 1-pinnate, lateral pinnae 4 or 5 pairs, ascending ca. 45 degrees from rachis, to 9 ×
<quantity id="4CC879A18C25FFAB3216FB1E8084FAED" box="[873,953,1262,1287]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="66" unit="cm" value="1.6">1.6 cm</quantity>
, stalked
<quantity id="4CC879A18C25FFAB3563FB1E8741FAED" box="[1052,1148,1262,1287]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="1" pageNumber="66" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">15 mm</quantity>
, bases rounded to cuneate, each pinna with a low, blackish, tuberculiform aerophore at its base, pinna apices abruptly attenuate and minutely toothed at the tips, margins otherwise entire, not revolute; sterile blade apices conform or with a single basal lobe, ca. 8 ×
<quantity id="4CC879A18C25FFAB31C8FAAA8235FA99" box="[183,264,1370,1395]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="66" unit="cm" value="1.6">1.6 cm</quantity>
, fertile pinnae delicate,
<emphasis id="B94408568C25FFAB3365FAAA8101FA99" box="[538,572,1370,1395]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
4 pairs per frond, each
<emphasis id="B94408568C25FFAB3230FAAA804CFA99" box="[847,881,1370,1395]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
<quantity id="4CC879A18C25FFAB3206FAAA808FFA99" box="[889,946,1370,1395]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="66" unit="cm" value="5.0">5 cm</quantity>
×
<quantity id="4CC879A18C25FFAB32ACFAAA8724FA99" box="[979,1049,1370,1395]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="66" unit="mm" value="1.0">1 mm</quantity>
;
<emphasis id="B94408568C25FFAB3559FAAA87BEFA99" box="[1062,1155,1370,1395]" italics="true" pageId="1" pageNumber="66">rachises</emphasis>
stramineous or brownish, with transverse ridges like those of stipes, sulcate adaxially, bearing sparse tan scales
<emphasis id="B94408568C25FFAB352AFA8E874BFA7D" box="[1109,1142,1406,1431]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
0.53 ×
<quantity id="4CC879A18C25FFAB35A2FA8E8658FA7D" box="[1245,1381,1406,1431]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="1" pageNumber="66" unit="mm" value="0.25" valueMax="0.4" valueMin="0.1">0.10.4 mm</quantity>
along the costae abaxially, lacking hairs;
<emphasis id="B94408568C25FFAB336AFA528172FA51" box="[533,591,1442,1467]" italics="true" pageId="1" pageNumber="66">veins</emphasis>
simple or once-forked near their base, free, closely parallel (set
<emphasis id="B94408568C25FFAB3454FA528670FA51" box="[1323,1357,1442,1467]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
<quantity id="4CC879A18C25FFAB342AFA528691FA51" box="[1365,1452,1442,1467]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="66" unit="mm" value="0.5">0.5 mm</quantity>
apart midway between costae and pinna margin), without enlarged endings (hydathodes) adaxially, not significantly raised on either side, but darkened abaxially;
<emphasis id="B94408568C25FFAB33F8FA1A81F2F9E9" box="[647,719,1514,1539]" italics="true" pageId="1" pageNumber="66">costae</emphasis>
abaxially with rather sparse, filiform to lanceolate tan scales, the larger ones widened and pectinate at their bases, mostly
<quantity id="4CC879A18C25FFAB3278F9FE80B7F9CD" box="[775,906,1550,1575]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="66" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5-1.5 mm</quantity>
long, veins and laminar tissue lacking scales on both sides;
<emphasis id="B94408568C25FFAB307AF9C2820CF9A1" box="[261,305,1586,1611]" italics="true" pageId="1" pageNumber="66">sori</emphasis>
linear, continuous, parallel to the costae, bearing filiform scales along the costae abaxially; indusia very narrow, glabrous, entire, at least at maturity; spores not seen.
</paragraph>
</subSubSection>
<subSubSection id="C32A87CF8C25FFAB31C2F98A878BF95D" pageId="1" pageNumber="66" type="distribution">
<paragraph id="8B8FD4448C25FFAB31C2F98A878BF95D" blockId="1.[136,1452,974,1899]" pageId="1" pageNumber="66">
<emphasis id="B94408568C25FFAB31C2F98A8137F979" bold="true" box="[189,522,1658,1683]" pageId="1" pageNumber="66">Distribution and habitat:—</emphasis>
This species grows on rocks, with fronds clumped, in montane rain forests at ca.
<quantity id="4CC879A18C25FFAB31F7F96E83DDF95D" box="[136,224,1694,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="66" unit="m" value="1300.0">1300 m</quantity>
. The rhizomes are obscured by the compact, fine root mass. Known only from the
<typeStatus id="548B6AE68C25FFAB3500F96E878EF95D" box="[1151,1203,1694,1719]" pageId="1" pageNumber="66">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A87CF8C25FFAB31C2F9328659F881" pageId="1" pageNumber="66" type="etymology">
<paragraph id="8B8FD4448C25FFAB31C2F93280DAF931" blockId="1.[136,1452,974,1899]" box="[189,999,1730,1755]" pageId="1" pageNumber="66">
<emphasis id="B94408568C25FFAB31C2F9328259F931" bold="true" box="[189,356,1730,1755]" pageId="1" pageNumber="66">Etymology:—</emphasis>
Named for the few pinna pairs on relatively small fronds.
</paragraph>
<paragraph id="8B8FD4448C25FFAB31C2F9168659F881" blockId="1.[136,1452,974,1899]" pageId="1" pageNumber="66">
<emphasis id="B94408568C25FFAB31C2F916821BF915" bold="true" box="[189,294,1766,1791]" pageId="1" pageNumber="66">Notes:—</emphasis>
This may be the most delicate species yet described in
<taxonomicName id="4C30AFC78C25FFAB3207F9168720F915" box="[888,1053,1766,1791]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B94408568C25FFAB3207F9168720F915" box="[888,1053,1766,1791]" italics="true" pageId="1" pageNumber="66">Parablechnum</emphasis>
</taxonomicName>
—relatively thin-textured compared to other species in the genus, and virtually scaleless or with mainly tan, small, filiform, scales on costae and rachises. There are only few pinna pairs, up to
<emphasis id="B94408568C25FFAB3355F8DE8171F8AD" box="[554,588,1838,1863]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
5, in both sterile and fertile fronds; the fertile pinnae are exceedingly narrow, only about
<quantity id="4CC879A18C25FFAB3079F8A2827AF881" box="[262,327,1874,1899]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="66" unit="mm" value="1.0">1 mm</quantity>
wide, the narrowest for any known species of
<taxonomicName id="4C30AFC78C25FFAB322EF8A280CBF881" box="[849,1014,1874,1899]" class="Polypodiopsida" family="Blechnaceae" genus="Parablechnum" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B94408568C25FFAB322EF8A280CBF881" box="[849,1014,1874,1899]" italics="true" pageId="1" pageNumber="66">Parablechnum</emphasis>
</taxonomicName>
. Nearest affinities are uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>