treatments-xml/data/03/8D/87/038D87F8FFBEFFE37AB1BB44FBA26CAB.xml

<document id="0B2118529A2DFB082CFC40BF85332B44" ID-CLB-Dataset="304005" ID-DOI="10.11646/phytotaxa.446.2.7" ID-GBIF-Dataset="8a3c3ece-f415-4e9a-a2a9-f4f0a77a89ed" ID-ISSN="1179-3163" ID-Zenodo-Dep="13877969" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727829247194" checkinUser="felipe" docAuthor="Tandang, Danilo N., Bustamante, Rene Alfredanton, Ferreras, Ulysses, Hadsall, Annalee S., Pym-Lyon, Stephanie &amp; Robinson, Alastair S." docDate="2020" docId="038D87F8FFBEFFE37AB1BB44FBA26CAB" docLanguage="en" docName="phytotaxa.446.2.7.pdf" docOrigin="Phytotaxa 446 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.446.2.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Corybas circinatus Tandang &amp; R. Bustam. 2020, spec. nov." docType="treatment" docVersion="2" lastPageNumber="139" masterDocId="FFB4FF80FFBFFFE77A39B929FF916B08" masterDocTitle="Corybas circinatus (Orchidaceae), a new species from Palawan, the Philippines" masterLastPageNumber="140" masterPageNumber="135" pageNumber="136" updateTime="1727835546902" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8A753FD4789324DA33B1174F6F03B668">Corybas circinatus (Orchidaceae), a new species from Palawan, the Philippines</mods:title>
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<mods:namePart id="F606FAEBAF5B6B5DED58EB98493EFC90">Tandang, Danilo N.</mods:namePart>
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<mods:affiliation id="0C6BF76E2482882D6BFEC38B1281B960">Institute of Biological Sciences, College of Arts and Sciences, University of the Philippines, Los Banos, College, Laguna 4031, Philippines.</mods:affiliation>
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<mods:affiliation id="D7C5D47F755781B55EFBEF24201BFAEA">Department of Botany, University of Wisconsin-Madison, Madison, WI 53706, USA.</mods:affiliation>
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<mods:namePart id="633DE6C7D894ECE4AF49D27673C5F44A">Robinson, Alastair S.</mods:namePart>
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<mods:affiliation id="8AE5D41D7FBE3B3AF092DE19E595889D">National Herbarium of Victoria, Royal Botanic Gardens, Melbourne, VIC 3004, Australia.</mods:affiliation>
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<mods:date id="246ABD40421BB0DFCF5C193220FDDAD2">2020</mods:date>
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<taxonomicName id="4C244D6DFFBEFFE67AB1BB44FDF36980" ID-CoL="9V7M8" authority="Tandang &amp; R.Bustam." authorityName="Tandang &amp; R. Bustam." authorityYear="2020" box="[136,610,621,648]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="136" phylum="Tracheophyta" rank="species" species="circinatus" status="sp. nov.">
<emphasis id="B950EAFCFFBEFFE67AB1BB44FEF0698F" bold="true" box="[136,353,621,648]" italics="true" pageId="1" pageNumber="136">Corybas circinatus</emphasis>
Tandang &amp; R.Bustam.
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(
<figureCitation id="131F2A6BFFBEFFE678DCBB44FCDF6980" box="[741,846,621,648]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[136,229,1336,1358]" captionStartId-1="3.[136,229,1493,1515]" captionStartId-2="4.[136,229,1105,1127]" captionTargetBox-0="[233,1354,187,1307]" captionTargetBox-1="[305,1267,242,1417]" captionTargetBox-2="[327,1261,209,1080]" captionTargetId-0="figure-26@2.[233,1354,187,1308]" captionTargetId-1="figure-18@3.[286,1302,221,1445]" captionTargetId-2="figure-26@4.[327,1261,209,1080]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Corybas circinatus, Tandang &amp; R.Bustam. spec. nov. A. Sterile plant in situ; B. Habit in situ; C–D. Lateral and front view at anthesis; E. Developing bloom; F. Tuber; Photos:A by Y. P.Ang; B.–F. by D. Tandang" captionText-1="FIGURE 2. Corybas circinatus, Tandang &amp; R.Bustam. spec. nov. A. Tuber; B. C. D. Habit, various views; E. Leaf and flower; F. Floral bracts; G. Petals and lateral sepals, flattened; H. I. Petals and lateral sepals, various shapes; J. Labellum, longitudinal section; K. Labellum, Flattened; L. Dorsal sepal." captionText-2="FIGURE 3. Distribution of Corybas circinatus on the island of Palawan, Philippines. Map generated using DIVA-GI." figureDoi-0="http://doi.org/10.5281/zenodo.13877971" figureDoi-1="http://doi.org/10.5281/zenodo.13877975" figureDoi-2="http://doi.org/10.5281/zenodo.13877977" httpUri-0="https://zenodo.org/record/13877971/files/figure.png" httpUri-1="https://zenodo.org/record/13877975/files/figure.png" httpUri-2="https://zenodo.org/record/13877977/files/figure.png" pageId="1" pageNumber="136">Figs. 1–3</figureCitation>
)
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</paragraph>
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<emphasis id="B950EAFCFFBEFFE67AB1BB90FF4B69C6" bold="true" box="[136,218,697,718]" pageId="1" pageNumber="136">Type:—</emphasis>
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, 
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, Mt. Bloomfield, 
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elevation, along trail, 
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, 
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, 
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, D. 
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. Tandang 2060, 
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255637 (
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<emphasis id="B950EAFCFFBEFFE67AB1BA28FE8E681F" bold="true" box="[136,287,769,791]" pageId="1" pageNumber="136">Description:—</emphasis>
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<emphasis id="B950EAFCFFBEFFE67B26BA28FE45681F" box="[287,468,769,791]" italics="true" pageId="1" pageNumber="136">Corybas circinatus</emphasis>
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is distinct from all known members of this genus in having a four-lobed labellum with two longer, upper lobes incurved over the two shorter lobes, a strongly cucullate dorsal sepal with a retuse apex, and a small, glandular protuberance at the front of the pedicel.
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Diminutive terrestrial herb, 
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tall, growing solitarily or in clonal colonies. Raphides present in almost all plant parts. Tuber ellipsoid, 3.0–6.0 mm long × 2.0–3.5 wide, densely villous. Below-ground axes terete, 
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long, densely muricate, narrow near tuber, 
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thick, becoming broader near the root collar, 
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thick. Stem solitary, erect, terete, 
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long × 
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thick, glabrous, green with purple streak along the edges, enclosed at base by a glabrous tubular sheath (cataphyll) 
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long with a short triangular blade. 
<emphasis id="B950EAFCFFBEFFE67F43BD34FA3D6F3F" box="[1402,1452,1053,1079]" italics="true" pageId="1" pageNumber="136">Leaf</emphasis>
solitary, borne horizontally above the ground, slightly fleshy, glabrous on both sides, broadly cordate, 
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from point of attachment to apex, up to 
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long including basal lobes, 
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at widest point, basal lobes sparsely overlapping, margins entire, often undulate, apex mucronate; adaxial surface minutely puncticulate, dark green with light pink to dark metallic magenta veins, sunken above and raised below, venation brochidodromous, midrib extending all the way to the apex and 4–5 lateral veins on either side of midrib, abaxial surface dark magenta. 
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solitary, nearly sessile, to 9.0– 
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high. 
<emphasis id="B950EAFCFFBEFFE6781CBDDCFD3E6E07" box="[549,687,1269,1295]" italics="true" pageId="1" pageNumber="136">Floral bract</emphasis>
erect at base, incurved or spreading at apex, linear-lanceolate, 
<quantity id="4CDC9B0BFFBEFFE67F4ABDDCFF256E3C" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="1" pageNumber="136" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
long × 1.0– 
<quantity id="4CDC9B0BFFBEFFE67B04BC30FE006E3C" box="[317,401,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="136" unit="mm" value="1.5">1.5 mm</quantity>
wide at base, translucent, magenta, keeled, margin crenate, apex acuminate. 
<emphasis id="B950EAFCFFBEFFE67ED7BC30FAD36E3B" box="[1262,1346,1305,1331]" italics="true" pageId="1" pageNumber="136">Pedicel</emphasis>
enclosed by floral bract, very short, 0.4–0.6 long × 
<quantity id="4CDC9B0BFFBEFFE67863BC14FD706E50" box="[602,737,1341,1368]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="136" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.6–0.7 mm</quantity>
wide at top, with a small glandular protuberance, 
<quantity id="4CDC9B0BFFBEFFE67F36BC17FA046E50" box="[1295,1429,1342,1368]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="1" pageNumber="136" unit="mm" value="0.25" valueMax="0.4" valueMin="0.1">0.1–0.4 mm</quantity>
× 
<quantity id="4CDC9B0BFFBEFFE67AB1BC4BFE806E74" box="[136,273,1378,1404]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="136" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
, arising from the ventral base. 
<emphasis id="B950EAFCFFBEFFE67857BC4BFD256E74" box="[622,692,1378,1404]" italics="true" pageId="1" pageNumber="136">Ovary</emphasis>
erect, 3.0– 
<quantity id="4CDC9B0BFFBEFFE67908BC48FC166E74" box="[817,903,1377,1404]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="136" unit="mm" value="4.5">4.5 mm</quantity>
long × 
<quantity id="4CDC9B0BFFBEFFE679E6BC48FBF76E74" box="[991,1126,1377,1404]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="1" pageNumber="136" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
wide, magenta, with 
<emphasis id="B950EAFCFFBEFFE67F60BC4BFAEB6E73" box="[1369,1402,1378,1403]" italics="true" pageId="1" pageNumber="136">ca.</emphasis>
5–6 longitudinal, slightly carinate ridges. 
<emphasis id="B950EAFCFFBEFFE67815BCACFD2D6E97" box="[556,700,1413,1439]" italics="true" pageId="1" pageNumber="136">Dorsal sepal</emphasis>
strongly cucullate, concave, incurved through almost 350 degrees, nearly enclosing the entire labellum, oblong when flattened, inconspicuously 3–5 at base, becoming 9 parallel-veined at tip, 
<quantity id="4CDC9B0BFFBEFFE67AECBCE7FEC06EE0" box="[213,337,1486,1512]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.9" metricValueMin="1.4" pageId="1" pageNumber="136" unit="mm" value="16.5" valueMax="19.0" valueMin="14.0">14–19 mm</quantity>
long, 
<quantity id="4CDC9B0BFFBEFFE67BA4BCE7FD836EE0" box="[413,530,1486,1512]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="1" pageNumber="136" unit="mm" value="1.7" valueMax="2.0" valueMin="1.4">1.4–2 mm</quantity>
wide at base, gradually widening to 
<quantity id="4CDC9B0BFFBEFFE679FABCE7FBB26EE0" box="[963,1059,1486,1512]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="136" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">
<emphasis id="B950EAFCFFBEFFE679FABCE7FC406EE0" box="[963,977,1486,1512]" italics="true" pageId="1" pageNumber="136">7</emphasis>
–8 mm
</quantity>
above the middle then narrowing gradually towards the blunt, downward-pointing apex, to 
<quantity id="4CDC9B0BFFBEFFE6792DBCDBFC166D04" box="[788,903,1522,1548]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.8" metricValueMin="2.0" pageId="1" pageNumber="136" unit="mm" value="2.4" valueMax="2.8" valueMin="2.0">2–2.8 mm</quantity>
wide, retuse; margin otherwise entire, incurved; abaxial (outer) surface densely papillate, basal half white with occasional dark purplish ruby to carmine red streaks, apical half longitudinally banded with dark, parallel veins of a similar hue on a richly coloured purplish ruby to carmine red background. 
<emphasis id="B950EAFCFFBEFFE67B98BF74FDAE6D7F" box="[417,575,1629,1655]" italics="true" pageId="1" pageNumber="136">Lateral sepals</emphasis>
and 
<emphasis id="B950EAFCFFBEFFE67848BF74FD256D7F" box="[625,692,1629,1655]" italics="true" pageId="1" pageNumber="136">petals</emphasis>
free, short, deep carmine, often hidden behind labellum spurs; sepals narrowly to broadly falcate, 1.0– 
<quantity id="4CDC9B0BFFBEFFE67BC1BFABFDDE6D94" box="[504,591,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="136" unit="mm" value="2.2">2.2 mm</quantity>
long × 
<quantity id="4CDC9B0BFFBEFFE6789FBFABFCBC6D94" box="[678,813,1666,1692]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="1" pageNumber="136" unit="mm" value="0.5" valueMax="0.8" valueMin="0.2">0.2–0.8 mm</quantity>
wide; petals similar to lateral sepals, linear-lanceolate to slightly falcate, 
<quantity id="4CDC9B0BFFBEFFE67B04BF8CFE556DC8" box="[317,452,1701,1728]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="3.5" metricValueMin="1.2" pageId="1" pageNumber="136" unit="mm" value="2.35" valueMax="3.5" valueMin="1.2">1.2–3.5 mm</quantity>
× 
<quantity id="4CDC9B0BFFBEFFE67BDBBF8FFDFB6DC8" box="[482,618,1702,1728]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="1" pageNumber="136" unit="mm" value="0.44999999999999996" valueMax="0.7" valueMin="0.2">0.2–0.7 mm</quantity>
wide. 
<emphasis id="B950EAFCFFBEFFE6788FBF8CFCB16DB7" box="[694,800,1701,1727]" italics="true" pageId="1" pageNumber="136">Labellum</emphasis>
from an erect, tubular basal part 3.6–4.3 x 4.0– 
<quantity id="4CDC9B0BFFBEFFE67F00BF8FFA1E6DC8" box="[1337,1423,1702,1728]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="136" unit="mm" value="4.8">4.8 mm</quantity>
in diameter with a 
<quantity id="4CDC9B0BFFBEFFE67B07BFE3FE506DEC" box="[318,449,1738,1764]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="1" pageNumber="136" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8-3.2 mm</quantity>
notch, becoming reflexed towards distal half, 
<quantity id="4CDC9B0BFFBEFFE679F2BFE3FB9D6DEC" box="[971,1036,1738,1764]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="136" unit="mm" value="8.0">8 mm</quantity>
long × 
<quantity id="4CDC9B0BFFBEFFE67E5ABFE0FB2B6DEC" box="[1123,1210,1737,1764]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="136" unit="mm" value="6.8">6.8 mm</quantity>
wide when flattened, 8-veined at base becoming 14-veined towards apex, anterior margin bilabiate with two longer, upper, lateral lobes, 
<quantity id="4CDC9B0BFFBEFFE67FA4BFC4FF256C24" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="136" unit="mm" value="5.0">5 mm</quantity>
× 
<quantity id="4CDC9B0BFFBEFFE67AEABE3BFE876C24" box="[211,278,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="136" unit="mm" value="2.0">2 mm</quantity>
, incurved over a shorter, lower, central, 1.5 × 
<quantity id="4CDC9B0BFFBEFFE67927BE3BFCF16C24" box="[798,864,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="136" unit="mm" value="4.0">4 mm</quantity>
lobe, the latter 3- veined, reflexed, broadly bilobed; abaxial (outer) surface glabrous and glossy, adaxial (inner) surface densely short hairy, whitish, longitudinally banded with 8 parallel dark purplish red stripes; at base with two distinct, horn-like spurs angled downwards, whitish streaked with fine purplish red lines, 
<quantity id="4CDC9B0BFFBEFFE67B87BE54FDD56C90" box="[446,580,1917,1944]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.6" metricValueMin="1.8" pageId="1" pageNumber="136" unit="mm" value="2.2" valueMax="2.6" valueMin="1.8">1.8–2.6 mm</quantity>
long from ventral base of flower to tip, 
<quantity id="4CDC9B0BFFBEFFE679C2BE54FB106C90" box="[1019,1153,1917,1944]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="1" pageNumber="136" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
wide at base. 
<emphasis id="B950EAFCFFBEFFE67F22BE54FAE56C9F" box="[1307,1396,1917,1943]" italics="true" pageId="1" pageNumber="136">Column</emphasis>
very short, 
<quantity id="4CDC9B0BFFBEFFE67AF4BE8BFE9D6CB4" box="[205,268,1954,1980]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="136" unit="mm" value="2.0">2 mm</quantity>
high × 
<quantity id="4CDC9B0BFFBEFFE67B66BE88FE226CB4" box="[351,435,1953,1980]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="136" unit="mm" value="1.5">1.5 mm</quantity>
wide, slightly curved; anther cap 
<quantity id="4CDC9B0BFFBEFFE6791EBE8BFCED6CB4" box="[807,892,1954,1980]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="136" unit="mm" value="0.9">0.9 mm</quantity>
long × 
<quantity id="4CDC9B0BFFBEFFE679F7BE8BFBB26CB4" box="[974,1059,1954,1980]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="136" unit="mm" value="1.1">1.1 mm</quantity>
wide, heart-shaped when flattened, keeled, with a short mucronate apiculum produced between the lobes; stigma orbicular with swollen apical margin, 
<quantity id="4CDC9B0BFFBEFFE67FB1BEEFFF25630C" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="136" unit="mm" value="1.2">1.2 mm</quantity>
wide; pollinia 4, in 2 pairs, 
<quantity id="4CDC9B0BFFBEFFE67BC9BEC0FDE6630C" box="[496,631,2025,2052]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="136" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
× 
<quantity id="4CDC9B0BFFBEFFE678ACBEC3FC8F630C" box="[661,798,2026,2052]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="136" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
, the two outer pollinia raised, the two inner pollinia lower and reduced, apparently adnate to anther cap.
</paragraph>
<caption id="DF5B6666FFBDFFE57AB1BC11FC5A6E7A" ID-DOI="http://doi.org/10.5281/zenodo.13877971" ID-Zenodo-Dep="13877971" httpUri="https://zenodo.org/record/13877971/files/figure.png" pageId="2" pageNumber="137" startId="2.[136,229,1336,1358]" targetBox="[233,1354,187,1307]" targetPageId="2" targetType="figure">
<paragraph id="8B9B36EEFFBDFFE57AB1BC11FC5A6E7A" blockId="2.[136,1451,1336,1394]" pageId="2" pageNumber="137">
<emphasis id="B950EAFCFFBDFFE57AB1BC11FF6F6E45" bold="true" box="[136,254,1336,1358]" pageId="2" pageNumber="137">FIGURE 1.</emphasis>
<taxonomicName id="4C244D6DFFBDFFE57B3ABC11FD0D6E46" authority="Tandang &amp; R.Bustam." authorityName="Tandang &amp; R. Bustam." authorityYear="2020" box="[259,668,1336,1358]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="species" species="circinatus" status="sp. nov.">
<emphasis id="B950EAFCFFBDFFE57B3ABC11FE286E46" box="[259,441,1336,1358]" italics="true" pageId="2" pageNumber="137">Corybas circinatus</emphasis>
, Tandang &amp; R.Bustam.
</taxonomicName>
<emphasis id="B950EAFCFFBDFFE57898BC10FD6E6E46" box="[673,767,1337,1358]" italics="true" pageId="2" pageNumber="137">
<taxonomicNameLabel id="A2635787FFBDFFE57898BC10FD6E6E46" box="[673,767,1337,1358]" pageId="2" pageNumber="137" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
A. Sterile plant 
<emphasis id="B950EAFCFFBDFFE57998BC10FC4D6E46" box="[929,988,1337,1358]" italics="true" pageId="2" pageNumber="137">in situ</emphasis>
; B. Habit 
<emphasis id="B950EAFCFFBDFFE57E06BC10FBEB6E46" box="[1087,1146,1337,1358]" italics="true" pageId="2" pageNumber="137">in situ</emphasis>
; C–D. Lateral and front view at anthesis; E. Developing bloom; F. Tuber; Photos: A by Y. P. Ang; B.–F. by D. Tandang
</paragraph>
</caption>
<paragraph id="8B9B36EEFFBDFFE57A84BC89FB476ED7" blockId="2.[136,1452,1440,2079]" pageId="2" pageNumber="137">
<emphasis id="B950EAFCFFBDFFE57A84BC89FD3A6EB2" bold="true" box="[189,683,1440,1466]" pageId="2" pageNumber="137">Additional Representative Specimens:—</emphasis>
<collectingCountry id="F333767EFFBDFFE57892BC88FCBD6EB3" box="[683,812,1441,1467]" name="Philippines" pageId="2" pageNumber="137">Philippines</collectingCountry>
, Island of 
<collectingRegion id="49E0F80CFFBDFFE57988BC88FB826EB3" box="[945,1043,1441,1467]" country="Philippines" name="Palawan" pageId="2" pageNumber="137">Palawan</collectingRegion>
, Mt. Bloomfield, 
<quantity id="4CDC9B0BFFBDFFE57ED2BC88FAA36EB3" box="[1259,1330,1441,1467]" metricMagnitude="2" metricUnit="m" metricValue="4.89" pageId="2" pageNumber="137" unit="m" value="489.0">489 m</quantity>
elevation, along trail, 
<geoCoordinate id="EE105029FFBDFFE57B31BCEFFE386ED7" box="[264,425,1477,1505]" degrees="10" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="137" precision="1" seconds="45.3" value="10.195917">10º11′45.3″ N</geoCoordinate>
, 
<geoCoordinate id="EE105029FFBDFFE57B8DBCEFFDCF6ED7" box="[436,606,1477,1505]" degrees="118" direction="east" minutes="52" orientation="longitude" pageId="2" pageNumber="137" precision="1" seconds="36.5" value="118.87681">118º52′36.5″ E</geoCoordinate>
, flower buds, 
<date id="FF9A102EFFBDFFE578C7BCECFC076ED7" box="[766,918,1477,1503]" pageId="2" pageNumber="137" value="2014-06-17">17 June 2014</date>
, D.N. Tandang 2061 (PNH).
</paragraph>
</subSubSection>
<subSubSection id="C33E6565FFBDFFE57A84BCC0FB1D6D9B" pageId="2" pageNumber="137" type="etymology">
<paragraph id="8B9B36EEFFBDFFE57A84BCC0FB366D2F" blockId="2.[136,1452,1440,2079]" pageId="2" pageNumber="137">
<emphasis id="B950EAFCFFBDFFE57A84BCC0FEF56D0B" bold="true" box="[189,356,1513,1539]" pageId="2" pageNumber="137">Etymology:—</emphasis>
The specific epithet 
<taxonomicName id="4C244D6DFFBDFFE5787EBCC0FD276D0A" box="[583,694,1513,1538]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBDFFE5787EBCC0FD276D0A" box="[583,694,1513,1538]" italics="true" pageId="2" pageNumber="137">circinatus</emphasis>
</taxonomicName>
is the perfect passive participle of the Latin verb 
<emphasis id="B950EAFCFFBDFFE57EE7BCC0FAD26D0A" box="[1246,1347,1513,1538]" italics="true" pageId="2" pageNumber="137">circinare</emphasis>
(to make round), meaning 
<emphasis id="B950EAFCFFBDFFE57B71BF24FE356D2F" box="[328,420,1549,1575]" italics="true" pageId="2" pageNumber="137">rounded</emphasis>
, in reference to the highly distinctive strongly incurved dorsal sepal.
</paragraph>
<paragraph id="8B9B36EEFFBDFFE57A84BF18FB1D6D9B" blockId="2.[136,1452,1440,2079]" pageId="2" pageNumber="137">
<emphasis id="B950EAFCFFBDFFE57A84BF18FECE6D43" bold="true" box="[189,351,1585,1611]" pageId="2" pageNumber="137">Phenology:—</emphasis>
Anthesis in 
<taxonomicName id="4C244D6DFFBDFFE57BDFBF18FD2A6D42" box="[486,699,1585,1611]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBDFFE57BDFBF18FD2A6D42" box="[486,699,1585,1611]" italics="true" pageId="2" pageNumber="137">Corybas circinatus</emphasis>
</taxonomicName>
has been observed on Mt. Bloomfield in 
<date id="FF9A102EFFBDFFE57EB6BF18FA906D43" box="[1167,1281,1585,1611]" pageId="2" pageNumber="137" value="2012-05">May 2012</date>
and 
<date id="FF9A102EFFBDFFE57F00BF18FA3A6D43" box="[1337,1451,1585,1611]" pageId="2" pageNumber="137" value="2014-06">June 2014</date>
(D.T. pers. observ.). It was also photographed in flower on Mt. 
<collectingRegion id="49E0F80CFFBDFFE57961BF7CFC236D67" box="[856,946,1621,1647]" country="Australia" name="Victoria" pageId="2" pageNumber="137">Victoria</collectingRegion>
in 
<date id="FF9A102EFFBDFFE579E1BF7CFBDF6D67" box="[984,1102,1621,1647]" pageId="2" pageNumber="137" value="2007-06">June 2007</date>
, on Mt. Mantalingahan in 
<date id="FF9A102EFFBDFFE57F44BF7CFF526D9B" pageId="2" pageNumber="137" value="2007-07">July 2007</date>
, on Sultan Peak in 
<date id="FF9A102EFFBDFFE57BA3BF50FD9E6D9B" box="[410,527,1657,1683]" pageId="2" pageNumber="137" value="2013-06">June 2013</date>
, and on Mt. Bahile in 
<date id="FF9A102EFFBDFFE57931BF50FCEB6D9B" box="[776,890,1657,1683]" pageId="2" pageNumber="137" value="2018-06">June 2018</date>
(all A.R., pers. observ.).
</paragraph>
</subSubSection>
<subSubSection id="C33E6565FFBDFFE37A84BFB5FBA26CAB" lastPageId="4" lastPageNumber="139" pageId="2" pageNumber="137" type="distribution">
<paragraph id="8B9B36EEFFBDFFE57A84BFB5FC146317" blockId="2.[136,1452,1440,2079]" pageId="2" pageNumber="137">
<emphasis id="B950EAFCFFBDFFE57A84BFB5FD846DBF" bold="true" box="[189,533,1692,1719]" pageId="2" pageNumber="137">Distribution and Ecology:—</emphasis>
<taxonomicName id="4C244D6DFFBDFFE5782CBFB4FD7F6DBE" box="[533,750,1693,1719]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBDFFE5782CBFB4FD7F6DBE" box="[533,750,1693,1719]" italics="true" pageId="2" pageNumber="137">Corybas circinatus</emphasis>
</taxonomicName>
is presently known exclusively from five ultramafic sites in 
<collectingRegion id="49E0F80CFFBDFFE57AB1BFE8FF7B6DD3" box="[136,234,1729,1755]" country="Philippines" name="Palawan" pageId="2" pageNumber="137">Palawan</collectingRegion>
, including: Mt. Bloomfield, 
<collectingRegion id="49E0F80CFFBDFFE5781EBFE8FD486DD3" box="[551,729,1729,1755]" country="Philippines" name="Puerto Princesa" pageId="2" pageNumber="137">Puerto Princesa</collectingRegion>
, at 
<quantity id="4CDC9B0BFFBDFFE578C4BFE8FCED6DD3" box="[765,892,1729,1755]" metricMagnitude="2" metricUnit="m" metricValue="4.1" metricValueMax="4.9" metricValueMin="3.3" pageId="2" pageNumber="137" unit="m" value="410.0" valueMax="490.0" valueMin="330.0">330–490 m</quantity>
elevation in shaded forest habitats in soils derived from peridotites such as harzburgite and dunite; Mt. Bahile, 
<collectingRegion id="49E0F80CFFBDFFE57900BFCCFC7C6DF7" box="[825,1005,1765,1791]" country="Philippines" name="Puerto Princesa" pageId="2" pageNumber="137">Puerto Princesa</collectingRegion>
, at 
<quantity id="4CDC9B0BFFBDFFE57E2FBFCDFB066DF7" box="[1046,1175,1764,1791]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="2" pageNumber="137" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300–450 m</quantity>
elevation in moderately to strongly shaded sites amongst stunted trees and scrub, particularly on moss covered clay embankments derived from degraded ultramafic rock; Mt. Mantalingahan, 
<collectingRegion id="49E0F80CFFBDFFE578EEBE04FC116C4F" box="[727,896,1837,1863]" country="Philippines" name="Rizal" pageId="2" pageNumber="137">Rizal Province</collectingRegion>
, at 
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elevation on sheltered banks close to a watercourse near Kawayan Camp; on Mt. 
<collectingRegion id="49E0F80CFFBDFFE57858BE78FD2F6C63" box="[609,702,1873,1899]" country="Australia" name="Victoria" pageId="2" pageNumber="137">Victoria</collectingRegion>
, Municipality of Narra, at 
<quantity id="4CDC9B0BFFBDFFE579D0BE78FBA16C63" box="[1001,1072,1873,1899]" metricMagnitude="2" metricUnit="m" metricValue="2.01" pageId="2" pageNumber="137" unit="m" value="201.0">201 m</quantity>
elevation on skeletal, humic soils overlying mafic rocks bordering 
<taxonomicName id="4C244D6DFFBDFFE57BCEBE5CFD246C87" authority="Camp" authorityName="Camp" box="[503,693,1909,1935]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="genus">
<emphasis id="B950EAFCFFBDFFE57BCEBE5CFDFD6C87" box="[503,620,1909,1935]" italics="true" pageId="2" pageNumber="137">Nepenthes</emphasis>
Camp
</taxonomicName>
on the Buhawi River; and on Sultan Peak, Municipality of Narra, at 
<quantity id="4CDC9B0BFFBDFFE57AB1BEB0FF5E6CBB" box="[136,207,1945,1971]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="2" pageNumber="137" unit="m" value="874.0">874 m</quantity>
elevation in leaf litter amongst stunted, mid-montane trees close to the waterfall camp. As with most species of 
<taxonomicName id="4C244D6DFFBDFFE57AB1BE94FF766CDF" authorityName="Salisbury" authorityYear="1807" box="[136,231,1981,2007]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="137" phylum="Tracheophyta" rank="genus">
<emphasis id="B950EAFCFFBDFFE57AB1BE94FF766CDF" box="[136,231,1981,2007]" italics="true" pageId="2" pageNumber="137">Corybas</emphasis>
</taxonomicName>
, this taxon appears to have rather specific habitat requirements and narrow geographic ranges that are difficult to predict even when modelled for the presence of suitable habitat. The exact suite of environmental parameters required for their survival and recruitment are still largely unknown.
</paragraph>
<caption id="DF5B6666FFBCFFE47AB1BCFCFE166D3B" ID-DOI="http://doi.org/10.5281/zenodo.13877975" ID-Zenodo-Dep="13877975" httpUri="https://zenodo.org/record/13877975/files/figure.png" pageId="3" pageNumber="138" startId="3.[136,229,1493,1515]" targetBox="[305,1267,242,1417]" targetPageId="3" targetType="figure">
<paragraph id="8B9B36EEFFBCFFE47AB1BCFCFE166D3B" blockId="3.[136,1452,1493,1587]" pageId="3" pageNumber="138">
<emphasis id="B950EAFCFFBCFFE47AB1BCFCFF6F6EE2" bold="true" box="[136,254,1493,1515]" pageId="3" pageNumber="138">FIGURE 2.</emphasis>
<taxonomicName id="4C244D6DFFBCFFE47B3DBCFCFD0F6EE3" authority="Tandang &amp; R.Bustam." authorityName="Tandang &amp; R. Bustam." authorityYear="2020" box="[260,670,1493,1515]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="138" phylum="Tracheophyta" rank="species" species="circinatus" status="sp. nov.">
<emphasis id="B950EAFCFFBCFFE47B3DBCFCFE2B6EE3" box="[260,442,1493,1515]" italics="true" pageId="3" pageNumber="138">Corybas circinatus</emphasis>
, Tandang &amp; R.Bustam.
</taxonomicName>
<emphasis id="B950EAFCFFBCFFE4789ABCFFFC906EE3" box="[675,769,1494,1515]" italics="true" pageId="3" pageNumber="138">
<taxonomicNameLabel id="A2635787FFBCFFE4789ABCFFFC906EE3" box="[675,769,1494,1515]" pageId="3" pageNumber="138" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
A. Tuber; B. C. D. Habit, various views; E. Leaf and flower; F. Floral bracts; G. Petals and lateral sepals, flattened; H. I. Petals and lateral sepals, various shapes; J. Labellum, longitudinal section; K. Labellum, Flattened; L. Dorsal sepal.
</paragraph>
</caption>
<paragraph id="8B9B36EEFFBCFFE47A84BF4BFCE96300" blockId="3.[136,1452,1634,2056]" pageId="3" pageNumber="138">
<emphasis id="B950EAFCFFBCFFE47A84BF4BFE476D74" bold="true" box="[189,470,1634,1660]" pageId="3" pageNumber="138">Conservation Status:—</emphasis>
The species has been documented in small numbers at five localities scattered across a 
<quantity id="4CDC9B0BFFBCFFE47AB1BFAFFF4C6DA8" box="[136,221,1670,1696]" metricMagnitude="5" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="138" unit="km" value="180.0">180 km</quantity>
range, the smallest population consisting of three plants in active growth, and the largest consisting of several colonies of 5–20 plants in the surveyed area. All observations were made on the flanks of peaks derived from the Mount Beaufort Ultramafics geological terrane, a series of outcrops of Eocene origin (
<bibRefCitation id="EFB54B1FFFBCFFE47E05BFE7FB586DE0" author="Okubo, Y." box="[1084,1225,1742,1768]" pageId="3" pageNumber="138" refId="ref3611" refString="Okubo, Y. (1989) The mineral exploration - mineral deposits and tectonics of two contrasting geologic environments in the Republic of the Philippines - consolidated report on Palawan area. Japan International Cooperation Agency, Metal Mining Agency of Japan and Department of Environment and Natural Resources (Philippines) Joint Committee, Philippines, 182 pp." type="book" year="1989">Okubo 1989</bibRefCitation>
) that account for the majority of the mountains of 
<collectingRegion id="49E0F80CFFBCFFE47BEABFDBFDA46C04" box="[467,565,1778,1804]" country="Philippines" name="Palawan" pageId="3" pageNumber="138">Palawan</collectingRegion>
, including the peaks of the Mt. 
<collectingRegion id="49E0F80CFFBCFFE479A3BFDBFC656C04" box="[922,1012,1778,1804]" country="Australia" name="Victoria" pageId="3" pageNumber="138">Victoria</collectingRegion>
Massif (Brow Shoulder, Mt. Shumkat, Sultan Peak, Sagpaw and the eponymous Mt. 
<collectingRegion id="49E0F80CFFBCFFE478B7BE3FFD7C6C38" box="[654,749,1814,1840]" country="Australia" name="Victoria" pageId="3" pageNumber="138">Victoria</collectingRegion>
), which represents the largest contiguous exposed land area of this terrane (
<bibRefCitation id="EFB54B1FFFBCFFE47B2FBE13FD996C5C" author="Robinson, A. S. &amp; Gironella, E. P. &amp; Cervancia, J. M." box="[278,520,1850,1876]" pageId="3" pageNumber="138" pagination="99 - 113" refId="ref3803" refString="Robinson, A. S., Gironella, E. P. &amp; Cervancia, J. M. (2016) New orchid species of Stigmatodactylus (Orchidoideae; Diurideae) and a new record of Cryptostylis carinata from central Palawan, Philippines. Phytotaxa 252 (2): 99 - 113. https: // doi. org / 10.11646 / phytotaxa. 252.2.2" type="journal article" year="2016">
Robinson 
<emphasis id="B950EAFCFFBCFFE47BB2BE12FE2C6C5C" box="[395,445,1850,1876]" italics="true" pageId="3" pageNumber="138">et al</emphasis>
. 2016
</bibRefCitation>
), as well as Mt. Mantalingahan, Escapardo Peak, Mt. Beaufort and Mt. Bahile. As such, the possibility remains for additional sites of this species being located, with the caveat that the lower reaches of this range are also currently the areas most threatened on 
<collectingRegion id="49E0F80CFFBCFFE47975BEABFC3A6C94" box="[844,939,1922,1948]" country="Philippines" name="Palawan" pageId="3" pageNumber="138">Palawan</collectingRegion>
by encroaching agricultural and subsistence practices. Such practices include the harvesting of timber for construction and for fuel, as well as the clearing of land for crop production, all of which lead to ecological shifts incompatible with the long term persistence of moisture and shade loving members of rainforest understorey herb communities.
</paragraph>
<caption id="DF5B6666FFBBFFE37AB1BD78FB646F6E" ID-DOI="http://doi.org/10.5281/zenodo.13877977" ID-Zenodo-Dep="13877977" box="[136,1269,1105,1127]" httpUri="https://zenodo.org/record/13877977/files/figure.png" pageId="4" pageNumber="139" startId="4.[136,229,1105,1127]" targetBox="[327,1261,209,1080]" targetPageId="4" targetType="figure">
<paragraph id="8B9B36EEFFBBFFE37AB1BD78FB646F6E" blockId="4.[136,1269,1105,1127]" box="[136,1269,1105,1127]" pageId="4" pageNumber="139">
<emphasis id="B950EAFCFFBBFFE37AB1BD78FF6F6F6E" bold="true" box="[136,254,1105,1127]" pageId="4" pageNumber="139">FIGURE 3.</emphasis>
Distribution of 
<taxonomicName id="4C244D6DFFBBFFE37BAEBD78FDDF6F6F" box="[407,590,1105,1127]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBBFFE37BAEBD78FDDF6F6F" box="[407,590,1105,1127]" italics="true" pageId="4" pageNumber="139">Corybas circinatus</emphasis>
</taxonomicName>
on the island of Palawan, Philippines. Map generated using DIVA-GI.
</paragraph>
</caption>
<paragraph id="8B9B36EEFFBBFFE37A84BDBFFE686E37" blockId="4.[136,1452,1173,1956]" pageId="4" pageNumber="139">
Current observations of 
<taxonomicName id="4C244D6DFFBBFFE37BF5BDBCFD306FA7" box="[460,673,1173,1199]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBBFFE37BF5BDBCFD306FA7" box="[460,673,1173,1199]" italics="true" pageId="4" pageNumber="139">Corybas circinatus</emphasis>
</taxonomicName>
made 
<emphasis id="B950EAFCFFBBFFE378D2BDBFFCBE6FA7" box="[747,815,1174,1199]" italics="true" pageId="4" pageNumber="139">in situ</emphasis>
satisfy the IUCN 3.1 Red List EN (Endangered) Criteria B1ab(ii,iii,iv)+2ab(i,ii,iii) (
<bibRefCitation id="EFB54B1FFFBBFFE37B83BD93FDDF6FDC" author="IUCN" box="[442,590,1210,1236]" pageId="4" pageNumber="139" refId="ref3524" refString="IUCN (2012) IUCN Red List Categories and Criteria Version 3.1, 2 nd ed. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. Available from: http: // www. iucnredlist. org / static / categories _ criteria _ 3 _ 1 (accessed 4 February 2020)" type="url" year="2012">IUCN, 2012</bibRefCitation>
), i.e. the species has an EOO &lt;
<quantity id="4CDC9B0BFFBBFFE379F4BD90FBA26FDB" box="[973,1075,1209,1236]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="139" unit="km" value="5000.0">5000 km</quantity>
<superScript id="7C519BA6FFBBFFE37E0ABD90FBAA6FCF" attach="left" box="[1075,1083,1209,1223]" fontSize="6" pageId="4" pageNumber="139">2</superScript>
and is known from 5 locations with inferred declines in area of occupancy and quality of habitat as a result of human activities causing observed degradation to two of its localities, and an AOO &lt;
<quantity id="4CDC9B0BFFBBFFE37888BC28FC946E13" box="[689,773,1281,1308]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="139" unit="km" value="500.0">500 km</quantity>
<superScript id="7C519BA6FFBBFFE3793CBC28FC9C6E07" attach="left" box="[773,781,1281,1295]" fontSize="6" pageId="4" pageNumber="139">2</superScript>
with a projected reduction in area of occurrence, occupancy, and extent and quality of habitat.
</paragraph>
<paragraph id="8B9B36EEFFBBFFE37A84BC63FBA26CAB" blockId="4.[136,1452,1173,1956]" pageId="4" pageNumber="139">
<emphasis id="B950EAFCFFBBFFE37A84BC63FEB76E6C" bold="true" box="[189,294,1354,1380]" pageId="4" pageNumber="139">Notes:—</emphasis>
<taxonomicName id="4C244D6DFFBBFFE37B1FBC60FE696E6B" box="[294,504,1353,1379]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBBFFE37B1FBC60FE696E6B" box="[294,504,1353,1379]" italics="true" pageId="4" pageNumber="139">Corybas circinatus</emphasis>
</taxonomicName>
resembles several other 
<taxonomicName id="4C244D6DFFBBFFE37938BC60FC246E6B" box="[769,949,1353,1379]" pageId="4" pageNumber="139">
<emphasis id="B950EAFCFFBBFFE37938BC60FCF16E6B" box="[769,864,1353,1379]" italics="true" pageId="4" pageNumber="139">Corybas</emphasis>
species
</taxonomicName>
in having a strongly curved, markedly hooded dorsal sepal that encloses the labellum. Only two species with this morphology have been previously described from regions west of Wallace’s line: 
<taxonomicName id="4C244D6DFFBBFFE37BDDBCB8FD4A6EA3" authorityName="Schlechter" authorityYear="1923" baseAuthorityName="J. J. Sm." box="[484,731,1425,1451]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="imperatorius">
<emphasis id="B950EAFCFFBBFFE37BDDBCB8FD4A6EA3" box="[484,731,1425,1451]" italics="true" pageId="4" pageNumber="139">Corybas imperatorius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C244D6DFFBBFFE37928BCB8FB666EA3" authority="J. Dransf. &amp; J. B. Comber" authorityName="J. Dransf. &amp; J. B. Comber" box="[785,1271,1425,1452]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="umbrosus">
<emphasis id="B950EAFCFFBBFFE37928BCB8FC736EA3" box="[785,994,1425,1451]" italics="true" pageId="4" pageNumber="139">Corybas umbrosus</emphasis>
J.Dransf. &amp; J.B.Comber
</taxonomicName>
in 
<bibRefCitation id="EFB54B1FFFBBFFE37F23BCB8FEA06EC7" author="Dransfield, J. &amp; Comber, J. B. &amp; Smith, G." pageId="4" pageNumber="139" pagination="575 - 613" refId="ref3471" refString="Dransfield, J., Comber, J. B. &amp; Smith, G. (1986) A Synopsis of Corybas (Orchidaceae) in West Malesia and Asia. Kew Bulletin 41 (3): 575 - 613. https: // doi. org / 10.2307 / 4103118" type="journal article" year="1986">
Dransfield 
<emphasis id="B950EAFCFFBBFFE37FAEBCBBFF0F6EC7" italics="true" pageId="4" pageNumber="139">et al</emphasis>
. (1986: 611)
</bibRefCitation>
, both endemic to the island of Java. While 
<taxonomicName id="4C244D6DFFBBFFE37920BC9FFC386EC7" box="[793,937,1462,1488]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBBFFE37920BC9FFC386EC7" box="[793,937,1462,1488]" italics="true" pageId="4" pageNumber="139">C. circinatus</emphasis>
</taxonomicName>
shares with these species dark green, broadly cordate leaves with contrasting veins, it differs in having a strongly 4-lobed labellum, a more curved dorsal sepal with a broad, retuse, incurved apex (vs. narrow, apiculate, recurved apex), and different colouration (dorsal sepal and labellum predominantly white with purplish carmine veins at the base vs. solidly purple at base). From 
<taxonomicName id="4C244D6DFFBBFFE37E90BF08FAEA6D33" box="[1193,1403,1569,1595]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="umbrosus">
<emphasis id="B950EAFCFFBBFFE37E90BF08FAEA6D33" box="[1193,1403,1569,1595]" italics="true" pageId="4" pageNumber="139">Corybas umbrosus</emphasis>
</taxonomicName>
, the new species is further differentiated by its smaller size (
<quantity id="4CDC9B0BFFBBFFE37935BF6FFCCF6D68" box="[780,862,1606,1632]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="139" unit="mm" value="10.0">10 mm</quantity>
vs. 
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tall). The new species occurs in shaded forest on ultramafic substrates at low elevations whereas both species from Java are found on other substrates at higher elevations (
<quantity id="4CDC9B0BFFBBFFE37B34BFA7FE3E6DA0" box="[269,431,1678,1704]" metricMagnitude="3" metricUnit="m" metricValue="1.685" metricValueMax="2.3" metricValueMin="1.0699999999999998" pageId="4" pageNumber="139" unit="m" value="1685.0" valueMax="2300.0" valueMin="1070.0">1070–2300 m</quantity>
elevation), with 
<taxonomicName id="4C244D6DFFBBFFE37842BFA4FCE66DAF" authorityName="Schlechter" authorityYear="1923" baseAuthorityName="J. J. Sm." box="[635,887,1677,1703]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="imperatorius">
<emphasis id="B950EAFCFFBBFFE37842BFA4FCE66DAF" box="[635,887,1677,1703]" italics="true" pageId="4" pageNumber="139">Corybas imperatorius</emphasis>
</taxonomicName>
found specifically in open grasslands (
<bibRefCitation id="EFB54B1FFFBBFFE37F73BFA4FF556DC4" author="Comber, J. B." pageId="4" pageNumber="139" refId="ref3452" refString="Comber, J. B. (1990) Orchids of Java. Royal Botanic Gardens, Kew." type="book" year="1990">Comber, 1990</bibRefCitation>
). The new species also appears similar to 
<taxonomicName id="4C244D6DFFBBFFE3788DBF98FB0A6DC4" authority="Schlechter (1923: 22)" authorityName="Schlechter" authorityPageNumber="22" authorityYear="1923" box="[692,1179,1713,1740]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="smithianus">
<emphasis id="B950EAFCFFBBFFE3788DBF98FC066DC3" box="[692,919,1713,1739]" italics="true" pageId="4" pageNumber="139">Corybas smithianus</emphasis>
<bibRefCitation id="EFB54B1FFFBBFFE37998BF98FB0A6DC4" author="Schlechter, F. R. R." box="[929,1179,1713,1740]" pageId="4" pageNumber="139" pagination="18 - 24" refId="ref3903" refString="Schlechter, F. R. R. (1923) Corybas Salisb. oder Corysanthes R. Br.? Repertorium specierum novarum regni vegetabilis. Centralblatt fur Sammlung und Veroffentlichung von Einzeldiagnosen neuer Pflanzen Beihefte, 19: 18 - 24. https: // doi. org / 10.1002 / fedr. 19230190104" type="journal article" year="1923">Schlechter (1923: 22)</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="4C244D6DFFBBFFE37EF0BF98FEAE6DF8" authority="Smith (1910: 7)" authorityName="Smith" authorityPageNumber="7" authorityYear="1910" class="Liliopsida" family="Orchidaceae" genus="Corysanthes" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="triloba">
<emphasis id="B950EAFCFFBBFFE37EF0BF98FA3A6DC3" box="[1225,1451,1713,1739]" italics="true" pageId="4" pageNumber="139">Corysanthes triloba</emphasis>
<bibRefCitation id="EFB54B1FFFBBFFE37AB1BFFCFEAE6DF8" author="Smith, J. J." box="[136,319,1749,1776]" pageId="4" pageNumber="139" pagination="1 - 22" refId="ref3959" refString="Smith, J. J. (1910) Vorlaufige Beschreibungen neuer papuanischer Orchideen. II. Bulletin du Departement de l'Agriculture aux Indes Neerlandaises 19: 1 - 22." type="journal article" year="1910">Smith (1910: 7)</bibRefCitation>
</taxonomicName>
(non Hook.f.)), a poorly known species described from western New 
<collectingCountry id="F333767EFFBBFFE37E55BFFCFB506DE7" box="[1132,1217,1749,1775]" name="Guinea-Bissau" pageId="4" pageNumber="139">Guinea</collectingCountry>
. With that species it shares both a strongly hooded dorsal sepal and a multi-lobed labellum with two lateral lobes incurved over a recurved central lobe. However, the shape of the dorsal sepal is different— 
<taxonomicName id="4C244D6DFFBBFFE37958BE37FC6B6C3F" authorityName="Schlechter" authorityYear="1923" box="[865,1018,1821,1848]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="smithianus">
<emphasis id="B950EAFCFFBBFFE37958BE37FC6B6C3F" box="[865,1018,1821,1848]" italics="true" pageId="4" pageNumber="139">C. smithianus</emphasis>
</taxonomicName>
, like the two species from Java, has an apiculate apex—and the middle lobe of the labellum is indistinctly bilobed with a crenulate-erose (vs. entire) margin. Furthermore, the protuberance/gland on the ventral pedicel of 
<taxonomicName id="4C244D6DFFBBFFE3797EBE4FFC496C77" box="[839,984,1894,1920]" class="Liliopsida" family="Orchidaceae" genus="Corybas" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="139" phylum="Tracheophyta" rank="species" species="circinatus">
<emphasis id="B950EAFCFFBBFFE3797EBE4FFC496C77" box="[839,984,1894,1920]" italics="true" pageId="4" pageNumber="139">C. circinatus</emphasis>
</taxonomicName>
appears to be a unique feature within the genus, though the significance and function of this structure are currently unknown.
</paragraph>
</subSubSection>
</treatment>
</document>