316 lines
39 KiB
XML
316 lines
39 KiB
XML
<document ID-DOI="10.11646/zootaxa.5264.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836674" ID-ZooBank="286F4C82-D3A7-455B-8174-354BFBF5D28F" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1681728586229" checkinUser="plazi" docAuthor="Shear, William A. & Marek, Paul E." docDate="2023" docId="CC6F87FEFFF260400AB8FF57FDA8B8D7" docLanguage="en" docName="zootaxa.5264.3.2.pdf" docOrigin="Zootaxa 5264 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miniaria ramifera Shear & Marek 2023, n. sp." docType="treatment" docVersion="2" lastPageNumber="328" masterDocId="3056FF86FFF160450A2FFFC0FF9CB922" masterDocTitle="The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)" masterLastPageNumber="340" masterPageNumber="323" pageNumber="326" updateTime="1681729003861" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The millipede family Striariidae Bollman, 1893. VIII. Three new genera and four new species of minute millipedes from Oregon and Washington, USA (Diplopoda, Chordeumatida, Striarioidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shear, William A.</mods:namePart>
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<mods:affiliation>Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943 USA, current address: 1950 Price Drive, Farmville VA 23901 USA.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:affiliation>Department of Entomology, Virginia Tech, Blacksburg VA 24061 USA.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5264</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>323</mods:start>
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<mods:end>340</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.3.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5264.3.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7836674</mods:identifier>
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<mods:identifier type="ZooBank">286F4C82-D3A7-455B-8174-354BFBF5D28F</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836678" ID-Zenodo-Dep="7836678" LSID="urn:lsid:plazi:treatment:CC6F87FEFFF260400AB8FF57FDA8B8D7" httpUri="http://treatment.plazi.org/id/CC6F87FEFFF260400AB8FF57FDA8B8D7" lastPageId="5" lastPageNumber="328" pageId="3" pageNumber="326">
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<subSubSection box="[151,657,151,177]" pageId="3" pageNumber="326" type="nomenclature">
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<paragraph blockId="3.[151,657,151,214]" box="[151,657,151,177]" pageId="3" pageNumber="326">
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<heading bold="true" box="[151,657,151,177]" fontSize="11" level="1" pageId="3" pageNumber="326" reason="1">
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<emphasis bold="true" box="[151,657,151,177]" pageId="3" pageNumber="326">
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<taxonomicName authority="Shear & Marek" authorityName="Shear & Marek" authorityYear="2023" box="[151,577,151,177]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="326" phylum="Arthropoda" rank="species" species="ramifera" status="n. sp.">
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<emphasis bold="true" box="[151,372,151,177]" italics="true" pageId="3" pageNumber="326">Miniaria ramifera</emphasis>
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Shear & Marek
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</taxonomicName>
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,
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<taxonomicNameLabel box="[590,657,151,177]" pageId="3" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="326" type="description">
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<paragraph blockId="3.[151,657,151,214]" box="[151,346,187,214]" pageId="3" pageNumber="326">
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<figureCitation box="[151,263,187,213]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–8" captionStart-2="FIGURES 9–14" captionStartId-0="3.[151,264,1908,1932]" captionStartId-1="4.[151,264,1492,1516]" captionStartId-2="6.[151,264,1848,1872]" captionTargetBox-0="[188,1400,525,1884]" captionTargetBox-1="[155,1434,181,1467]" captionTargetBox-2="[174,1414,181,1823]" captionTargetId-0="figure-137@3.[188,1400,525,1884]" captionTargetId-1="figure-20@4.[153,1434,181,1468]" captionTargetId-2="figure-20@6.[174,1414,181,1823]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURES 1–4. Miniaria ramifera n. gen., n. sp., male holotype. 1. Head and legpairs 1 and 2, anterioventral view. 2. Left leg 1, posterior view. 3. Coxae of legpair 2, ventral view. 4. Legpairs 3 and 4, ventral view. Abbreviations: cf, coxal flask of coxa 3; cx3, body of coxa 3; lab, labrum; L (numeral), numbered legpair; m, mandibular stipes; om, ommatidium; pf (numeral), prefemur of numbered legpair; vd, opening of vas deferens." captionText-1="FIGURES 5–8. Miniaria ramifera n. gen., n. sp., male holotype. 5. Legpairs 2 and 3, ventral view. 6. Tarsus of left leg 3, lateral view. 7. Comb seta of tarsus 3. 8. Legpairs 4–6, ventral view. Abbreviations: cf, coxal flask of leg 3; L (numeral), numbered legpair;pf(numeral),prefemurofnumberedleg;w,accessory clawoftarsus3,x,combsetae;y,recessedporeplate;z,sensory seta." captionText-2="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi-0="http://doi.org/10.5281/zenodo.7836676" figureDoi-1="http://doi.org/10.5281/zenodo.7836680" figureDoi-2="http://doi.org/10.5281/zenodo.7836682" httpUri-0="https://zenodo.org/record/7836676/files/figure.png" httpUri-1="https://zenodo.org/record/7836680/files/figure.png" httpUri-2="https://zenodo.org/record/7836682/files/figure.png" pageId="3" pageNumber="326">Figs 1–11</figureCitation>
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,
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<figureCitation box="[275,346,187,214]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="3" pageNumber="326">39–45</figureCitation>
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</paragraph>
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<paragraph blockId="3.[151,1436,259,502]" pageId="3" pageNumber="326">
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<emphasis box="[151,223,260,285]" italics="true" pageId="3" pageNumber="326">Types:</emphasis>
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<materialsCitation collectingDate="2004-03-19" collectionCode="CAS" collectorName="W. Leonard & C. Richart. Parts" county="Pierce Co." latitude="46.86695" location="Mashel River" longLatPrecision="1" longitude="-122.33645" municipality="Male" pageId="3" pageNumber="326" specimenCount="14" specimenCount-female="6" specimenCount-male="6" stateProvince="Washington" typeStatus="holotype">
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All specimens deposited in
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<collectionCode box="[553,611,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="3" pageNumber="326" type="Museum">CAS</collectionCode>
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.
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<collectingMunicipality box="[624,682,259,285]" pageId="3" pageNumber="326">Male</collectingMunicipality>
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<typeStatus box="[692,793,259,285]" pageId="3" pageNumber="326">holotype</typeStatus>
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,
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<specimenCount box="[807,885,259,285]" pageId="3" pageNumber="326" type="male">6 male</specimenCount>
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<typeStatus box="[895,1002,260,286]" pageId="3" pageNumber="326">paratypes</typeStatus>
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and
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<specimenCount box="[1062,1162,259,285]" pageId="3" pageNumber="326" type="female">6 female</specimenCount>
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<typeStatus box="[1172,1279,260,286]" pageId="3" pageNumber="326">paratypes</typeStatus>
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from
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<location LSID="urn:lsid:plazi:treatment:CC6F87FEFFF260400AB8FF57FDA8B8D7:41196033FFF260460F65FEC3FF49B863" county="Pierce Co." latitude="46.86695" longLatPrecision="1" longitude="-122.33645" municipality="Male" name="Mashel River" pageId="3" pageNumber="326" stateProvince="Washington">Mashel River</location>
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at
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<location LSID="urn:lsid:plazi:treatment:CC6F87FEFFF260400AB8FF57FDA8B8D7:41196033FFF260460AD6FEE7FEE0B860" box="[249,380,295,322]" county="Pierce Co." latitude="46.86695" longLatPrecision="1" longitude="-122.33645" municipality="Male" name="State Route" pageId="3" pageNumber="326" stateProvince="Washington">State Route</location>
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7,
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<geoCoordinate box="[416,556,295,322]" degrees="46.86695" direction="north" orientation="latitude" pageId="3" pageNumber="326" precision="1" value="46.86695">46.86695°N</geoCoordinate>
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, -
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<geoCoordinate box="[575,734,295,322]" degrees="122.33645" direction="west" orientation="longitude" pageId="3" pageNumber="326" precision="1" value="-122.33645">122.33645°W</geoCoordinate>
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, 600′asl,
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<collectingCounty box="[840,959,295,321]" pageId="3" pageNumber="326">Pierce Co.</collectingCounty>
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,
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<collectingRegion box="[971,1107,295,321]" country="United States of America" name="Washington" pageId="3" pageNumber="326">Washington</collectingRegion>
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, collected
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<date box="[1228,1400,295,321]" pageId="3" pageNumber="326" value="2004-03-19">
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<collectingDate box="[1228,1400,295,321]" pageId="3" pageNumber="326" value="2004-03-19">19 March 2004</collectingDate>
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</date>
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by
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<collectorName box="[151,282,331,358]" pageId="3" pageNumber="326">W. Leonard</collectorName>
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and
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<collectorName box="[337,522,331,358]" pageId="3" pageNumber="326">C. Richart. Parts</collectorName>
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of the
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<typeStatus box="[600,698,331,357]" pageId="3" pageNumber="326">holotype</typeStatus>
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are on SEM stub WS36-8
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="326" type="diagnosis">
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<paragraph blockId="3.[151,1436,259,502]" pageId="3" pageNumber="326">
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<emphasis bold="true" box="[199,314,367,393]" pageId="3" pageNumber="326">Diagnosis</emphasis>
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. Distinct from
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<taxonomicName authority="Shear & Marek, 2023" authorityName="Shear & Marek" authorityYear="2023" box="[499,627,367,393]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="326" phylum="Arthropoda" rank="species" species="richarti" status="n. sp.">
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<emphasis box="[499,627,367,393]" italics="true" pageId="3" pageNumber="326">M. richarti</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[639,706,367,393]" pageId="3" pageNumber="326">
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<taxonomicNameLabel box="[639,706,367,393]" pageId="3" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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in having the gonopod anterior angiocoxite with a prominent subterminal posterior branch (
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<emphasis bold="true" box="[492,540,403,429]" pageId="3" pageNumber="326">aac,</emphasis>
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<figureCitation box="[547,661,403,429]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="3" pageNumber="326">Figs 9, 11</figureCitation>
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,
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<figureCitation box="[674,749,403,429]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="3" pageNumber="326">39–44</figureCitation>
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). The gonopods are widely separated mesally, while those of
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<taxonomicName authority="Shear & Marek, 2023" authorityName="Shear & Marek" authorityYear="2023" box="[151,273,439,465]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="326" phylum="Arthropoda" rank="species" species="richarti" status="n. sp.">
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<emphasis box="[151,273,439,465]" italics="true" pageId="3" pageNumber="326">M. richarti</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[278,340,439,465]" pageId="3" pageNumber="326">
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<taxonomicNameLabel box="[278,340,439,465]" pageId="3" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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are appressed to each other in the midline. The anterior angiocoxites are evenly curved, not sharply elbowed as in
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<taxonomicName authority="Shear & Marek, 2023" authorityName="Shear & Marek" authorityYear="2023" box="[312,435,475,501]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="326" phylum="Arthropoda" rank="species" species="richarti" status="n. sp.">
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<emphasis box="[312,435,475,501]" italics="true" pageId="3" pageNumber="326">M. richarti</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[442,505,475,501]" pageId="3" pageNumber="326">
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<taxonomicNameLabel box="[442,505,475,501]" pageId="3" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7836676" ID-Zenodo-Dep="7836676" httpUri="https://zenodo.org/record/7836676/files/figure.png" pageId="3" pageNumber="326" startId="3.[151,264,1908,1932]" targetBox="[188,1400,525,1884]" targetPageId="3">
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<paragraph blockId="3.[151,1436,1908,2040]" pageId="3" pageNumber="326">
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<emphasis bold="true" box="[151,314,1908,1932]" pageId="3" pageNumber="326">FIGURES 1–4.</emphasis>
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<taxonomicName authority="Shear & Marek, 2023" authorityName="Shear & Marek" authorityYear="2023" box="[321,506,1908,1932]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="326" phylum="Arthropoda" rank="species" species="ramifera" status="n. gen., n. sp.">
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<emphasis box="[321,506,1908,1932]" italics="true" pageId="3" pageNumber="326">Miniaria ramifera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[512,653,1908,1932]" pageId="3" pageNumber="326" rank="species">
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<emphasis bold="true" box="[512,584,1908,1932]" pageId="3" pageNumber="326">n. gen.</emphasis>
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,
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<emphasis bold="true" box="[596,653,1908,1932]" pageId="3" pageNumber="326">n. sp.</emphasis>
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</taxonomicNameLabel>
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, male holotype.
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<emphasis bold="true" box="[822,842,1908,1932]" pageId="3" pageNumber="326">1.</emphasis>
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Head and legpairs 1 and 2, anterioventral view.
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<emphasis bold="true" box="[1331,1350,1908,1932]" pageId="3" pageNumber="326">2.</emphasis>
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Left leg 1, posterior view.
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<emphasis bold="true" box="[334,353,1944,1968]" pageId="3" pageNumber="326">3.</emphasis>
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Coxae of legpair 2, ventral view.
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<emphasis bold="true" box="[698,716,1944,1968]" pageId="3" pageNumber="326">4.</emphasis>
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Legpairs 3 and 4, ventral view.
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<emphasis bold="true" box="[1043,1231,1944,1968]" pageId="3" pageNumber="326">Abbreviations: cf</emphasis>
|
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, coxal flask of coxa 3;
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<emphasis bold="true" box="[178,215,1980,2004]" pageId="3" pageNumber="326">cx3</emphasis>
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, body of coxa 3;
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<emphasis bold="true" box="[398,432,1980,2004]" pageId="3" pageNumber="326">lab</emphasis>
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, labrum;
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<emphasis bold="true" box="[532,663,1980,2004]" pageId="3" pageNumber="326">L (numeral)</emphasis>
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, numbered legpair;
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<emphasis bold="true" box="[870,891,1980,2004]" pageId="3" pageNumber="326">m</emphasis>
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, mandibular stipes;
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<emphasis bold="true" box="[1100,1134,1980,2004]" pageId="3" pageNumber="326">om</emphasis>
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, ommatidium;
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<emphasis bold="true" box="[1292,1430,1980,2004]" pageId="3" pageNumber="326">pf (numeral)</emphasis>
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, prefemur of numbered legpair;
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<emphasis bold="true" box="[469,496,2016,2040]" pageId="3" pageNumber="326">vd</emphasis>
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, opening of
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<emphasis box="[624,749,2016,2040]" italics="true" pageId="3" pageNumber="326">vas deferens</emphasis>
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.
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</paragraph>
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</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836680" ID-Zenodo-Dep="7836680" httpUri="https://zenodo.org/record/7836680/files/figure.png" pageId="4" pageNumber="327" startId="4.[151,264,1492,1516]" targetBox="[155,1434,181,1467]" targetPageId="4">
|
||
<paragraph blockId="4.[151,1437,1492,1589]" pageId="4" pageNumber="327">
|
||
<emphasis bold="true" box="[151,314,1492,1516]" pageId="4" pageNumber="327">FIGURES 5–8.</emphasis>
|
||
<taxonomicName authority="Shear & Marek, 2023" authorityName="Shear & Marek" authorityYear="2023" box="[319,503,1492,1516]" class="Diplopoda" family="Striariidae" genus="Miniaria" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="327" phylum="Arthropoda" rank="species" species="ramifera" status="n. gen., n. sp.">
|
||
<emphasis box="[319,503,1492,1516]" italics="true" pageId="4" pageNumber="327">Miniaria ramifera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[509,647,1492,1516]" pageId="4" pageNumber="327" rank="species">
|
||
<emphasis bold="true" box="[509,579,1492,1516]" pageId="4" pageNumber="327">n. gen.</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[590,647,1492,1516]" pageId="4" pageNumber="327">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype.
|
||
<emphasis bold="true" box="[813,826,1492,1516]" pageId="4" pageNumber="327">5</emphasis>
|
||
. Legpairs 2 and 3, ventral view.
|
||
<emphasis bold="true" box="[1151,1164,1492,1516]" pageId="4" pageNumber="327">6</emphasis>
|
||
. Tarsus of left leg 3, lateral view.
|
||
<emphasis bold="true" box="[212,225,1528,1552]" pageId="4" pageNumber="327">7</emphasis>
|
||
. Comb seta of tarsus 3.
|
||
<emphasis bold="true" box="[472,485,1528,1552]" pageId="4" pageNumber="327">8</emphasis>
|
||
. Legpairs 4–6, ventral view.
|
||
<emphasis bold="true" box="[781,969,1528,1552]" pageId="4" pageNumber="327">Abbreviations: cf</emphasis>
|
||
, coxal flask of leg 3;
|
||
<emphasis bold="true" box="[1191,1322,1528,1552]" pageId="4" pageNumber="327">L (numeral)</emphasis>
|
||
, numbered legpair;
|
||
<emphasis bold="true" box="[232,366,1564,1588]" pageId="4" pageNumber="327">pf(numeral)</emphasis>
|
||
, prefemur of numbered leg;
|
||
<emphasis bold="true" box="[640,658,1564,1588]" pageId="4" pageNumber="327">w</emphasis>
|
||
, accessory claw of tarsus 3,
|
||
<emphasis bold="true" box="[929,942,1564,1588]" pageId="4" pageNumber="327">x</emphasis>
|
||
, comb setae;
|
||
<emphasis bold="true" box="[1071,1084,1564,1588]" pageId="4" pageNumber="327">y</emphasis>
|
||
, recessed pore plate;
|
||
<emphasis bold="true" box="[1289,1300,1564,1588]" pageId="4" pageNumber="327">z</emphasis>
|
||
, sensory seta.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="4" pageNumber="327" type="etymology">
|
||
<paragraph blockId="4.[151,1437,1634,2021]" pageId="4" pageNumber="327">
|
||
<emphasis bold="true" box="[199,328,1634,1660]" pageId="4" pageNumber="327">Etymology</emphasis>
|
||
. The species name is a Latin adjective, meaning “branch-bearing,” and refers to the subterminal posterior branch of the gonopod anterior angiocoxite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="327" type="description">
|
||
<paragraph blockId="4.[151,1437,1634,2021]" pageId="4" pageNumber="327">
|
||
<emphasis bold="true" box="[199,337,1706,1732]" pageId="4" pageNumber="327">Description</emphasis>
|
||
. Male
|
||
<typeStatus box="[414,510,1707,1733]" pageId="4" pageNumber="327">paratype</typeStatus>
|
||
from Mashel River. Length
|
||
<quantity box="[825,912,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="327" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, greatest width
|
||
<quantity box="[1086,1185,1706,1733]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="327" unit="mm" value="0.31">0.31 mm</quantity>
|
||
. Nonsexual characters and secondary sexual modifications as described for genus.Gonopods (
|
||
<figureCitation box="[929,1037,1742,1768]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="4" pageNumber="327">Figs 9, 11</figureCitation>
|
||
,
|
||
<figureCitation box="[1047,1118,1742,1768]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="4" pageNumber="327">39–44</figureCitation>
|
||
) small; anterior angiocoxites (
|
||
<emphasis bold="true" box="[161,201,1778,1804]" pageId="4" pageNumber="327">aac</emphasis>
|
||
,
|
||
<figureCitation box="[218,288,1778,1804]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="4" pageNumber="327">Fig. 9</figureCitation>
|
||
) curved posteriad, slightly knobbed at tips, distally with prominent fringed margin, long, flattened subterminal process. Posterior angiocoxites (
|
||
<emphasis bold="true" box="[655,697,1814,1840]" pageId="4" pageNumber="327">pac</emphasis>
|
||
,
|
||
<figureCitation box="[711,777,1814,1840]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="4" pageNumber="327">Fig. 9</figureCitation>
|
||
) less than half length of anterior ones, fimbriate especially along posterior margin, sheathing single curved flagellocoxite or flagellum. Colpocoxite reduced to single branch with membranous base. (
|
||
<emphasis bold="true" box="[433,465,1886,1912]" pageId="4" pageNumber="327">cc,</emphasis>
|
||
<figureCitation box="[472,601,1886,1912]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="4" pageNumber="327">Figs 39–44</figureCitation>
|
||
). Ninth legs (
|
||
<figureCitation box="[751,833,1886,1912]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1848,1872]" captionTargetBox="[174,1414,181,1823]" captionTargetId="figure-20@6.[174,1414,181,1823]" captionTargetPageId="6" captionText="FIGURES 9–14. Miniaria n. gen. species. 9–11. Miniaria ramifera n. gen., n. sp., male holotype. 9. Gonopods, posterior view. 10. Legpair 9 and coxae of legpair 10, anterior view. 11. Tip of gonopod anterior angiocoxite, ventral view. 12–14. Miniaria richarti n. gen., n. sp., male from Germany Creek. 12. Head and legpair 1, ventrolateral view. 13. Pleurotergites 11 and 12, lateral view. 14. Pleurotergites 12 and 13, dorsal view. Abbreviations: aac, anterior angiocoxite; cx, gonopod coxa; cx (numeral), coxa of numbered legpair; lab, labrum; L1, legpair 1; m, mandibular stipes; numerals, ring numbers; om, ommatidium." figureDoi="http://doi.org/10.5281/zenodo.7836682" httpUri="https://zenodo.org/record/7836682/files/figure.png" pageId="4" pageNumber="327">Fig. 10</figureCitation>
|
||
) as described for genus.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1634,2021]" box="[199,768,1922,1949]" pageId="4" pageNumber="327">Females similar to males in all nonsexual respects.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="328" pageId="4" pageNumber="327" type="distribution">
|
||
<paragraph blockId="4.[151,1437,1634,2021]" lastBlockId="5.[151,1437,150,501]" lastPageId="5" lastPageNumber="328" pageId="4" pageNumber="327">
|
||
<emphasis bold="true" box="[199,345,1958,1984]" pageId="4" pageNumber="327">Distribution</emphasis>
|
||
.
|
||
<collectingRegion box="[369,490,1958,1984]" country="United States of America" name="Oregon" pageId="4" pageNumber="327">OREGON</collectingRegion>
|
||
: Multnomah Co.: Ainsworth State Park,
|
||
<geoCoordinate box="[999,1111,1958,1985]" degrees="45.596" direction="north" orientation="latitude" pageId="4" pageNumber="327" precision="55" value="45.596">45.596°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate box="[1139,1270,1958,1985]" degrees="122.051" direction="west" orientation="longitude" pageId="4" pageNumber="327" precision="55" value="-122.051">122.051°W</geoCoordinate>
|
||
, 210′ asl,
|
||
<date pageId="4" pageNumber="327" value="1967-11-15">15 November 1967</date>
|
||
, E. M. Benedict,
|
||
<quantity box="[536,605,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="327" unit="mm" value="2.0">2 mm</quantity>
|
||
. Gobbert Butte, Gresham,
|
||
<geoCoordinate box="[909,1034,1994,2021]" degrees="45.4731" direction="north" orientation="latitude" pageId="4" pageNumber="327" precision="5" value="45.4731">45.4731°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate box="[1054,1199,1994,2020]" degrees="122.4378" direction="west" orientation="longitude" pageId="4" pageNumber="327" precision="5" value="-122.4378">122.4378°W</geoCoordinate>
|
||
,
|
||
<date box="[1211,1404,1994,2020]" pageId="4" pageNumber="327" value="2016-10-30">30 October 2016</date>
|
||
, P. Nosler, m.
|
||
<collectingRegion box="[276,409,151,177]" country="United States of America" name="Washington" pageId="5" pageNumber="328">Washington</collectingRegion>
|
||
Co.:
|
||
<quantity box="[473,546,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="5" pageNumber="328" unit="mi" value="1.7">1.7 mi</quantity>
|
||
W of Timber, T3N, R5W, sec.?, 300′ asl,
|
||
<date box="[1024,1244,151,177]" pageId="5" pageNumber="328" value="1971-11-27">27 November 1971</date>
|
||
, E. M. Benedict, m.
|
||
<collectingRegion box="[190,381,187,213]" country="United States of America" name="Washington" pageId="5" pageNumber="328">WASHINGTON</collectingRegion>
|
||
: Pierce Co.: Ohop Valley Road at State Route 7, 600′ asl,
|
||
<geoCoordinate box="[1073,1212,186,213]" degrees="46.86695" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="1" value="46.86695">46.86695°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate box="[1235,1393,186,213]" degrees="122.33645" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="1" value="-122.33645">122.33645°W</geoCoordinate>
|
||
,
|
||
<date pageId="5" pageNumber="328" value="2004-01-12">12 January 2004</date>
|
||
, W. Leonard, m. Skamania Co.: Cook-Underwood Road,
|
||
<quantity box="[966,1040,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.2530816" pageId="5" pageNumber="328" unit="mi" value="1.4">1.4 mi</quantity>
|
||
NE of Cook, 450′ asl,
|
||
<geoCoordinate box="[1307,1432,222,249]" degrees="45.7239" direction="north" orientation="latitude" pageId="5" pageNumber="328" precision="5" value="45.7239">45.7239°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate box="[159,318,258,285]" degrees="121.64765" direction="west" orientation="longitude" pageId="5" pageNumber="328" precision="1" value="-121.64765">121.64765°W</geoCoordinate>
|
||
,
|
||
<date box="[329,548,259,285]" pageId="5" pageNumber="328" value="2003-11-30">30 November 2003</date>
|
||
, W. Leonard,
|
||
<quantity box="[703,771,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="328" unit="mm" value="4.0">4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,150,501]" pageId="5" pageNumber="328">
|
||
<emphasis bold="true" box="[199,266,295,321]" pageId="5" pageNumber="328">Notes</emphasis>
|
||
. Gonopod variation in this species focuses on the degree to which the anterior angiocoxite is terminally knobbed and the extensiveness of the fimbriated posterior distal margin. In the Ainsworth State Park male (
|
||
<figureCitation box="[1340,1423,331,357]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="5" pageNumber="328">Fig. 43</figureCitation>
|
||
), the fimbriate part appears as a coalesced process. In the Timber (
|
||
<figureCitation box="[883,964,367,393]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="5" pageNumber="328">Fig. 44</figureCitation>
|
||
) and Mashel River (
|
||
<figureCitation box="[1196,1326,367,393]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="5" pageNumber="328">Figs 39, 40</figureCitation>
|
||
) males, it is more extensive and does not protrude, and while in the male from Cook-Underwood Road (
|
||
<figureCitation box="[1195,1322,403,429]" captionStart="FIGURES 39–46" captionStartId="13.[151,262,1817,1841]" captionTargetBox="[216,1378,182,1788]" captionTargetId="figure-17@13.[206,1382,181,1792]" captionTargetPageId="13" captionText="FIGURES 39–46. Gonopods and ninth legs of male striariid millipedes. Figs. 39–45. Miniaria ramifera n. gen., n. sp. 39. Right gonopod of specimen from Mashel River, mesal view. 40. Same, lateral view. 41. Right gonopod of specimen from Cook- Underwood Road, mesal view. 42. Same, lateral view. 43, Right gonopod of specimen fromAinsworth State Park, mesal view. 44. Right gonopod of specimen from Timber, mesal view. 45. Ninth legpair of specimen from Ainsworth State Park, posterior view. 46. Miniaria richarti n. gen., n. sp., right gonopod of specimen from Hospital Creek, mesal view. Abbreviations: aac, anterior angiocoxite; cc, colpocoxite; cx9, ninth leg coxa; cxp, coxal process; pac, posterior angiocoxite; t9, telopodite of ninth leg." figureDoi="http://doi.org/10.5281/zenodo.7836702" httpUri="https://zenodo.org/record/7836702/files/figure.png" pageId="5" pageNumber="328">Figs 41, 42</figureCitation>
|
||
) there is a broad fimbriate region. We do not have enough localities to determine if this is geographic variation or simply due to positioning on a microscope slide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |