377 lines
48 KiB
XML
377 lines
48 KiB
XML
<document id="CFE9EEA13A5A7B3845F192770B045778" ID-DOI="10.5252/g2012n2a6" ID-ISSN="1638-9395" ID-Zenodo-Dep="5378837" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630286459456" checkinUser="marcus" docAuthor="Petit, Gilles & Charbonnier, Sylvain" docDate="2012" docId="03C2C703FFBFD500FF34F6513379DD7A" docLanguage="en" docName="Geodiversitas.2012.34.2.359-372.pdf" docOrigin="Geodiversitas 34 (2)" docSource="http://dx.doi.org/10.5252/g2012n2a6" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Carpopenaeus garassinoi Petit & Charbonnier 2012, n. sp." docType="treatment" docVersion="3" lastPageNumber="364" masterDocId="FFFBBF7BFFBCD505FFB9F23F371CDE77" masterDocTitle="Fossil sponge gemmules, epibionts of Carpopenaeus garassinoi n. sp. (Crustacea, Decapoda) from the Sahel Alma Lagerstätte (Late Cretaceous, Lebanon)" masterLastPageNumber="372" masterPageNumber="359" pageNumber="362" updateTime="1699214653279" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="B993F4D1888ED3E17779ABE41625EF66">Fossil sponge gemmules, epibionts of Carpopenaeus garassinoi n. sp. (Crustacea, Decapoda) from the Sahel Alma Lagerstätte (Late Cretaceous, Lebanon)</mods:title>
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<mods:namePart id="F43701BA819FDCE680CD754A100D0DD5">Petit, Gilles</mods:namePart>
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<mods:namePart id="2F3970EE7D3014E4C8B7C3C739C2D284">Charbonnier, Sylvain</mods:namePart>
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<mods:title id="F7ECD479CD85B7748B82F59ABC2C085A">Geodiversitas</mods:title>
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<mods:part id="A890754011BCB58CAFFB2EB441D83B32">
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<mods:date id="1BB5AADF69280EEF4B964FB29D4D705F">2012</mods:date>
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<mods:number id="1730E9520D7D84975FC44BBB32933249">2012-06-30</mods:number>
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<mods:number id="4AEF95959A95001E739443ABC8D23568">34</mods:number>
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<mods:url id="BCFB25441D6377A45BBC5CEC233B153F">http://dx.doi.org/10.5252/g2012n2a6</mods:url>
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<mods:identifier id="B5694F03457DCBF532460DD858EA9E4E" type="DOI">10.5252/g2012n2a6</mods:identifier>
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<mods:identifier id="D8E0623412907CCAE27816B732B7268F" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03C2C703FFBFD500FF34F6513379DD7A" LSID="urn:lsid:plazi:treatment:03C2C703FFBFD500FF34F6513379DD7A" httpUri="http://treatment.plazi.org/id/03C2C703FFBFD500FF34F6513379DD7A" lastPageId="5" lastPageNumber="364" pageId="3" pageNumber="362">
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<subSubSection id="C371259EFFBFD506FF34F65136B8DADF" pageId="3" pageNumber="362" type="nomenclature">
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<paragraph id="8BD47615FFBFD506FF34F65136B8DADF" blockId="3.[141,619,1134,1192]" pageId="3" pageNumber="362">
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<heading id="D09CC179FFBFD506FF34F65136B8DADF" centered="true" fontSize="11" level="2" pageId="3" pageNumber="362" reason="2">
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<taxonomicName id="4C6B0D96FFBFD506FF34F6513535DAFF" authority="Charbonnier" authorityName="Petit & Charbonnier" authorityYear="2012" box="[141,553,1134,1160]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="362" phylum="Arthropoda" rank="species" species="garassinoi" status="sp. nov.">
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<emphasis id="B91FAA07FFBFD506FF34F651368BDAFF" bold="true" box="[141,407,1134,1160]" italics="true" pageId="3" pageNumber="362">Carpopenaeus garassinoi</emphasis>
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Charbonnier
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</taxonomicName>
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<taxonomicNameLabel id="A22C177CFFBFD506FD89F6503577DAFE" box="[560,619,1135,1161]" pageId="3" pageNumber="362" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="13506A90FFBFD506FEE2F6B13685DADF" box="[347,409,1166,1192]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1</figureCitation>
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)
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C371259EFFBFD506FFC8F6EE36B8DBB2" pageId="3" pageNumber="362" type="materials_examined">
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||
<paragraph id="8BD47615FFBFD506FFC8F6EE3626DB74" blockId="3.[113,646,1233,1283]" pageId="3" pageNumber="362">
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<materialsCitation id="3B037C48FFBFD506FFC8F6EE3626DB74" accessionNumber="A33532" collectionCode="MNHN" pageId="3" pageNumber="362" specimenCount="1" typeStatus="holotype">
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TYPE MATERIAL. —
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<typeStatus id="54D0C8B7FFBFD506FE88F6EE3692DA9E" box="[305,398,1233,1257]" pageId="3" pageNumber="362" type="holotype">Holotype</typeStatus>
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||
(
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||
<collectionCode id="ED7AEED0FFBFD506FE25F6EE36F1DA9E" box="[412,493,1233,1257]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="362">MNHN</collectionCode>
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.F.
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<accessionNumber id="9438EBF6FFBFD506FDB8F6EE3549DA9E" box="[513,597,1233,1257]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A33532" pageId="3" pageNumber="362" type="EnaNcbi">A33532</accessionNumber>
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, collection Arambourg).
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BD47615FFBFD506FFC8F71A3682DB20" blockId="3.[113,646,1316,1367]" pageId="3" pageNumber="362">
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<typeStatus id="54D0C8B7FFBFD506FFC8F71A37BFDB4D" box="[113,163,1317,1340]" pageId="3" pageNumber="362">TYPE</typeStatus>
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LOCALITY. — Sahel Alma Lagerstätte (
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<emphasis id="B91FAA07FFBFD506FD94F71A352ADB4B" box="[557,566,1317,1340]" italics="true" pageId="3" pageNumber="362">c</emphasis>
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||
.
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<quantity id="4C93DBF0FFBFD506FDFDF71A359ADB4B" box="[580,646,1316,1341]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="362" unit="km" value="20.0">20 km</quantity>
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northeast of
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<collectingRegion id="49AFB8F7FFBFD506FF56F700362CDB20" box="[239,304,1343,1367]" country="Lebanon" name="Beyrouth" pageId="3" pageNumber="362">Beirut</collectingRegion>
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),
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<collectingCountry id="F37C3685FFBFD506FEF8F7003686DB20" box="[321,410,1343,1367]" name="Lebanon" pageId="3" pageNumber="362">Lebanon</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BD47615FFBFD506FFC8F74636B8DBB2" blockId="3.[113,648,1400,1477]" pageId="3" pageNumber="362">
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<typeStatus id="54D0C8B7FFBFD506FFC8F74637B8DBF9" box="[113,164,1401,1424]" pageId="3" pageNumber="362">TYPE</typeStatus>
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HORIZON. — Late Cretaceous sublithographic limestones from the Sahel Alma quarry (Late Santonian after
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<bibRefCitation id="EFFA0BE4FFBFD506FF1DF792368BDBB2" author="EJEL F. & DUBERTRET L." box="[164,407,1453,1477]" pageId="3" pageNumber="362" pagination="353 - 354" refId="ref6552" refString="EJEL F. & DUBERTRET L. 1966. - Sur l'age precis du gisement de poissons et de crustaces cretaces de Sahel Alma (Liban). Comptes rendus sommaires des Seances de la Societe geologique de France 9: 353 - 354." type="journal article" year="1966">Ejel & Dubertret [1966]</bibRefCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C371259EFFBFD506FD1CF2E735F3DF19" pageId="3" pageNumber="362" type="etymology">
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<paragraph id="8BD47615FFBFD506FD1CF2E733A4DF54" blockId="3.[677,1209,215,292]" pageId="3" pageNumber="362">
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ETYMOLOGY. — In honor of Dr. Alessandro Garassino (Museo Civico di Storia Naturale di Milano,
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<collectingCountry id="F37C3685FFBFD506FBD4F2CE3381DF7E" box="[1133,1181,241,265]" name="Italy" pageId="3" pageNumber="362">Italy</collectingCountry>
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), a specialist in the palaeontology of decapod crustaceans.
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||
</paragraph>
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<paragraph id="8BD47615FFBFD506FD1CF30235F3DF19" blockId="3.[677,1209,316,366]" pageId="3" pageNumber="362">
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STUDIED MATERIAL. — Only the
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||
<typeStatus id="54D0C8B7FFBFD506FC57F3033358DF23" box="[1006,1092,316,340]" pageId="3" pageNumber="362" type="holotype">holotype</typeStatus>
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||
is currently known.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C371259EFFBFD506FD1CF3B833B1DFE9" box="[677,1197,390,415]" pageId="3" pageNumber="362" type="description">
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||
<paragraph id="8BD47615FFBFD506FD1CF3B833B1DFE9" blockId="3.[677,1197,390,415]" box="[677,1197,390,415]" pageId="3" pageNumber="362">
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||
OCCURENCES. — Known only from the
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||
<typeStatus id="54D0C8B7FFBFD506FB88F3B93347DFE9" box="[1073,1115,390,414]" pageId="3" pageNumber="362">type</typeStatus>
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||
locality.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C371259EFFBFD506FD1CF38934AEDCD6" pageId="3" pageNumber="362" type="diagnosis">
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||
<paragraph id="8BD47615FFBFD506FD1CF38934AEDCD6" blockId="3.[675,1212,437,673]" pageId="3" pageNumber="362">
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DIAGNOSIS. — Large species of
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<taxonomicName id="4C6B0D96FFBFD506FC64F3893343DFBA" authorityName="Glaessner" authorityYear="1945" box="[989,1119,438,461]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="362" phylum="Arthropoda" rank="genus">
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<emphasis id="B91FAA07FFBFD506FC64F3893343DFBA" box="[989,1119,438,461]" italics="true" pageId="3" pageNumber="362">Carpopenaeus</emphasis>
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</taxonomicName>
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; rostrum very elongated, curved upward distally, with four basal suprarostral spines, one median suprarostral spine, one median subrostral spine and three distal spines; third maxilliped well-developed, with serrated ventral margin; pereiopods 2 and 3 very elongated, with multiarticulate carpus; pereiopods 4 and 5 thin and achelate; basipodite well-developed; exopod with rounded diaresis and distal part bearing very small pits.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C371259EFFBFD500FD1CF0EC3531DD1B" lastPageId="5" lastPageNumber="364" pageId="3" pageNumber="362" type="description">
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||
<paragraph id="8BD47615FFBFD506FD1CF0EC3429DC9C" blockId="3.[674,1212,723,1478]" box="[677,821,723,749]" pageId="3" pageNumber="362">DESCRIPTION</paragraph>
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<paragraph id="8BD47615FFBFD506FD1CF0CD3309DA5E" blockId="3.[674,1212,723,1478]" pageId="3" pageNumber="362">
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<materialsCitation id="3B037C48FFBFD506FD1CF0CD346BDD1B" collectionCode="CL" country="Switzerland" location="CH" pageId="3" pageNumber="362" specimenCount="1" typeStatus="holotype">
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Subrectangular cephalothorax, height nearly equal with length (
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<typeStatus id="54D0C8B7FFBFD506FC80F12D34BEDD5B" box="[825,930,786,812]" pageId="3" pageNumber="362" type="holotype">holotype</typeStatus>
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:
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<collectionCode id="ED7AEED0FFBFD506FC16F12D34CFDD5B" box="[943,979,786,812]" pageId="3" pageNumber="362">CL</collectionCode>
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=
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<emphasis id="B91FAA07FFBFD506FC4CF12C331ADD5B" box="[1013,1030,787,812]" italics="true" pageId="3" pageNumber="362">c.</emphasis>
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<quantity id="4C93DBF0FFBFD506FBA9F12D337ADD5A" box="[1040,1126,786,813]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="362" unit="mm" value="22.0">22 mm</quantity>
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||
,
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||
<collectingCountry id="F37C3685FFBFD506FBCCF12D33BCDD5B" box="[1141,1184,786,812]" name="Switzerland" pageId="3" pageNumber="362">CH</collectingCountry>
|
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=
|
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<emphasis id="B91FAA07FFBFD506FD1EF10C35A4DD3B" box="[679,696,819,844]" italics="true" pageId="3" pageNumber="362">c.</emphasis>
|
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<quantity id="4C93DBF0FFBFD506FD07F10D340DDD3B" box="[702,785,818,844]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="362" unit="mm" value="21.0">21 mm</quantity>
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||
), with short longitudinal carina at midheight (
|
||
<figureCitation id="13506A90FFBFD506FD40F16E3473DD1B" box="[761,879,849,876]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1A, B</figureCitation>
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||
)
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||
</materialsCitation>
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||
; anterior margin fragmented; ventral margin not clearly visible; posterior margin slightly sinuous with a thin ridge; very elongated rostrum curved upward distally, with four basal suprarostral spines, one median suprarostral spine, one median subrostral spine and three distal spines (
|
||
<figureCitation id="13506A90FFBFD506FD13F63035E1DA5D" box="[682,765,1039,1066]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1C</figureCitation>
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||
); ornamentation smooth.
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||
</paragraph>
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||
<paragraph id="8BD47615FFBFD506FD02F610345CDBB1" blockId="3.[674,1212,723,1478]" pageId="3" pageNumber="362">
|
||
Cephalic appendages well-preserved (
|
||
<figureCitation id="13506A90FFBFD506FBE7F61035A6DA1E" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1A, B</figureCitation>
|
||
): antennulae (a1) composed of long antennular peduncle with three segments diminishing in length distally and two long and thin multi-articulated flagellae (preserved length:
|
||
<emphasis id="B91FAA07FFBFD506FC70F69034CEDABF" box="[969,978,1199,1224]" italics="true" pageId="3" pageNumber="362">c</emphasis>
|
||
.
|
||
<quantity id="4C93DBF0FFBFD506FC59F691332EDABF" box="[992,1074,1198,1225]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="362" unit="mm" value="25.0">25 mm</quantity>
|
||
,
|
||
<figureCitation id="13506A90FFBFD506FB85F69133B1DABF" box="[1084,1197,1198,1224]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1B, C</figureCitation>
|
||
); antennae (a2) composed of antennal peduncle with a very long multi-articulated flagellum (preserved length:
|
||
<emphasis id="B91FAA07FFBFD506FD4AF731341FDB50" box="[755,771,1294,1319]" italics="true" pageId="3" pageNumber="362">c.</emphasis>
|
||
<quantity id="4C93DBF0FFBFD506FCB1F732344BDB50" box="[776,855,1293,1319]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="362" unit="mm" value="90.0">90 mm</quantity>
|
||
); scaphocerite well-developed, oblong and pointed with a longitudinal lateral carina. – Third maxilliped relatively long ending in dactylus (
|
||
<figureCitation id="13506A90FFBFD506FD13F753343CDBF1" box="[682,800,1388,1414]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="3" pageNumber="362">Fig. 1B, C</figureCitation>
|
||
), ventral margin of carpus, propodus and dactylus bearing one row of small teeth (serrated pattern);
|
||
</paragraph>
|
||
<caption id="DF14269DFFBFD506FFC8F43733A7D8D6" pageId="3" pageNumber="362" startId="3.[113,124,1544,1561]">
|
||
<paragraph id="8BD47615FFBFD506FFC8F43733A7D8D6" blockId="3.[113,1211,1544,1697]" pageId="3" pageNumber="362">
|
||
FIG. 1. —
|
||
<taxonomicName id="4C6B0D96FFBFD506FF7DF4373518D86E" authority="Charbonnier" authorityName="Petit & Charbonnier" authorityYear="2012" box="[196,516,1544,1561]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="362" phylum="Arthropoda" rank="species" species="garassinoi" status="sp. nov.">
|
||
<emphasis id="B91FAA07FFBFD506FF7DF4373684D86E" box="[196,408,1544,1561]" italics="true" pageId="3" pageNumber="362">Carpopenaeus garassinoi</emphasis>
|
||
Charbonnier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C177CFFBFD506FDB2F4373527D86E" box="[523,571,1544,1561]" pageId="3" pageNumber="362" rank="species">n. sp.</taxonomicNameLabel>
|
||
(holotype, MNHN.F.
|
||
<accessionNumber id="9438EBF6FFBFD506FD59F437343FD86E" box="[736,803,1544,1561]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A33532" pageId="3" pageNumber="362" type="EnaNcbi">A33532</accessionNumber>
|
||
, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon):
|
||
<emphasis id="B91FAA07FFBFD506FE11F42036A9D847" bold="true" box="[424,437,1567,1584]" pageId="3" pageNumber="362">A</emphasis>
|
||
, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;
|
||
<emphasis id="B91FAA07FFBFD506FE77F40A36C7D831" bold="true" box="[462,475,1589,1606]" pageId="3" pageNumber="362">B</emphasis>
|
||
, line drawing of the holotype;
|
||
<emphasis id="B91FAA07FFBFD506FD6BF40A35FCD831" bold="true" box="[722,736,1589,1606]" pageId="3" pageNumber="362">C</emphasis>
|
||
, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus;
|
||
<emphasis id="B91FAA07FFBFD506FEECF45C367FD803" bold="true" box="[341,355,1635,1652]" pageId="3" pageNumber="362">D</emphasis>
|
||
, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations:
|
||
<emphasis id="B91FAA07FFBFD506FC6BF44634FBD8FD" bold="true" box="[978,999,1657,1674]" pageId="3" pageNumber="362">a1</emphasis>
|
||
, antennula;
|
||
<emphasis id="B91FAA07FFBFD506FBE9F4463379D8FD" bold="true" box="[1104,1125,1657,1674]" pageId="3" pageNumber="362">a2</emphasis>
|
||
, antenna;
|
||
<emphasis id="B91FAA07FFBFD506FFCAF4AF3789D8D6" bold="true" box="[115,149,1680,1697]" pageId="3" pageNumber="362">bap</emphasis>
|
||
, basipodite;
|
||
<emphasis id="B91FAA07FFBFD506FF44F4AF360ED8D6" bold="true" box="[253,274,1680,1697]" pageId="3" pageNumber="362">ca</emphasis>
|
||
, carpus;
|
||
<emphasis id="B91FAA07FFBFD506FEE7F4AF3673D8D6" bold="true" box="[350,367,1680,1697]" pageId="3" pageNumber="362">ct</emphasis>
|
||
, cephalothorax;
|
||
<emphasis id="B91FAA07FFBFD506FE41F4AF3512D8D6" bold="true" box="[504,526,1680,1697]" pageId="3" pageNumber="362">da</emphasis>
|
||
, dactylus;
|
||
<emphasis id="B91FAA07FFBFD506FDDEF4AF356BD8D6" bold="true" box="[615,631,1680,1697]" pageId="3" pageNumber="362">di</emphasis>
|
||
, diaresis;
|
||
<emphasis id="B91FAA07FFBFD506FD73F4AF35FCD8D6" bold="true" box="[714,736,1680,1697]" pageId="3" pageNumber="362">en</emphasis>
|
||
, endopod;
|
||
<emphasis id="B91FAA07FFBFD506FC84F4AF344ED8D6" bold="true" box="[829,850,1680,1697]" pageId="3" pageNumber="362">ex</emphasis>
|
||
, exopod;
|
||
<emphasis id="B91FAA07FFBFD506FC1AF4AF34AED8D6" bold="true" box="[931,946,1680,1697]" pageId="3" pageNumber="362">is</emphasis>
|
||
, ischium;
|
||
<emphasis id="B91FAA07FFBFD506FBBCF4AF3309D8D6" bold="true" box="[1029,1045,1680,1697]" pageId="3" pageNumber="362">lc</emphasis>
|
||
, longitudinal carina;
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD47615FFB8D501FFC8F44635EBD8D6" blockId="4.[113,1211,1657,1697]" pageId="4" pageNumber="363">
|
||
<emphasis id="B91FAA07FFB8D501FFC8F4463791D8FD" bold="true" box="[113,141,1657,1674]" pageId="4" pageNumber="363">me</emphasis>
|
||
, merus;
|
||
<emphasis id="B91FAA07FFB8D501FF63F44637E3D8FD" bold="true" box="[218,255,1657,1674]" pageId="4" pageNumber="363">mx3</emphasis>
|
||
, maxilliped 3;
|
||
<emphasis id="B91FAA07FFB8D501FEC7F44636AED8FD" bold="true" box="[382,434,1657,1674]" pageId="4" pageNumber="363">p2-p5</emphasis>
|
||
, pereiopods 2 to 5;
|
||
<emphasis id="B91FAA07FFB8D501FDDAF446356AD8FD" bold="true" box="[611,630,1657,1674]" pageId="4" pageNumber="363">pr</emphasis>
|
||
, propodus;
|
||
<emphasis id="B91FAA07FFB8D501FD66F44635E3D8FD" bold="true" box="[735,767,1657,1674]" pageId="4" pageNumber="363">pa1</emphasis>
|
||
, antennular peduncle;
|
||
<emphasis id="B91FAA07FFB8D501FC7FF44634D1D8FD" bold="true" box="[966,973,1657,1674]" pageId="4" pageNumber="363">r</emphasis>
|
||
, rostrum;
|
||
<emphasis id="B91FAA07FFB8D501FB9FF4463327D8FD" bold="true" box="[1062,1083,1657,1674]" pageId="4" pageNumber="363">sc</emphasis>
|
||
, scaphocerite;
|
||
<emphasis id="B91FAA07FFB8D501FFCAF4AF37B8D8D6" bold="true" box="[115,164,1680,1697]" pageId="4" pageNumber="363">s1-s6</emphasis>
|
||
, somites 1 to 6;
|
||
<emphasis id="B91FAA07FFB8D501FE97F4AF365FD8D6" bold="true" box="[302,323,1680,1697]" pageId="4" pageNumber="363">sp</emphasis>
|
||
, spine;
|
||
<emphasis id="B91FAA07FFB8D501FE3CF4AF3690D8D6" bold="true" box="[389,396,1680,1697]" pageId="4" pageNumber="363">t</emphasis>
|
||
, telson. Scale bars: A, B,
|
||
<quantity id="4C93DBF0FFB8D501FDDAF4AF358CD8D6" box="[611,656,1680,1697]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="363" unit="cm" value="2.0">2 cm</quantity>
|
||
; C, D,
|
||
<quantity id="4C93DBF0FFB8D501FD7EF4AF35EFD8D6" box="[711,755,1680,1697]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="363" unit="cm" value="1.0">1 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD47615FFB9D500FFC8F2E736C8DF07" blockId="5.[111,648,216,876]" pageId="5" pageNumber="364">Pereiopod 1 not preserved; pereiopods 2 and 3 very elongated, both displaying antennae-like multiarticulate carpus, ending in minute chela; pereiopods 4 and 5 almost equal in size, thin and smooth, achelate, ending in small dactyli.</paragraph>
|
||
<paragraph id="8BD47615FFB9D500FFC8F3493531DD1B" blockId="5.[111,648,216,876]" pageId="5" pageNumber="364">
|
||
Abdomen convergent distally with smooth ornamentation; somites 1 and 6 very reduced, somite 2 badly preserved, shorter than somite3, somite 3 large and well-developed, somites 4 and 5 short; pleurae of somites ventrally smooth and rounded (
|
||
<figureCitation id="13506A90FFB9D500FD93F3CA3564DC78" box="[554,632,501,527]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="5" pageNumber="364">Fig. 1B</figureCitation>
|
||
); pleopods not preserved; telson short (length:
|
||
<emphasis id="B91FAA07FFB9D500FDEEF02A3574DC59" box="[599,616,533,558]" italics="true" pageId="5" pageNumber="364">c.</emphasis>
|
||
⅔ length of uropods), pointed, only known in section; basipodite of the uropods well-developed, subcircular (
|
||
<figureCitation id="13506A90FFB9D500FFCEF04B37DBDCF9" box="[119,199,628,654]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="5" pageNumber="364">Fig.1D</figureCitation>
|
||
); uropodal exopod and endopod elongated, almost equal in length with lateral margins smooth and rounded inferior margins; exopod with median longitudinal carina and a well-developed diaresis with crescent-shaped suture delimiting a distal part uniformly covered with very small pits; endopod slender, with median longitudinal carina and distal part also bearing very small pits (
|
||
<figureCitation id="13506A90FFB9D500FE73F16E3503DD1B" box="[458,543,849,876]" captionStart="FIG" captionStartId="3.[113,124,1544,1561]" captionText="FIG. 1. — Carpopenaeus garassinoi Charbonnier n. sp. (holotype, MNHN.F.A33532, collection Arambourg) from the Sahel Alma Lagerstätte (Late Santonian, Lebanon): A, specimen flattened in right lateral view, note the exquisite preservation with cephalothorax, abdomen and appendages in connection;B, line drawing of the holotype;C, detail of the cephalic region showing the rostrum with its spines (black arrows), the antennulae, the base of one antenna,the 3rd maxilliped with serrated ventral margin and the 2nd pereiopod with multiarticulate carpus; D, detail of the tail fan, note the well-developed basipodite, the rounded uropods with diaresis on exopod and the distal part of each uropod uniformly covered by very small pits (black arrows). Abbreviations: a1, antennula; a2, antenna; bap, basipodite; ca, carpus; ct, cephalothorax;da, dactylus;di, diaresis; en, endopod; ex, exopod;is, ischium; lc, longitudinal carina;" pageId="5" pageNumber="364">Fig. 1D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C371259EFFB9D500FFC8F1AE3379DD7A" pageId="5" pageNumber="364" type="discussion">
|
||
<paragraph id="8BD47615FFB9D500FFC8F1AE37EFDDDE" blockId="5.[111,649,913,1700]" box="[113,243,913,939]" pageId="5" pageNumber="364">DISCUSSION</paragraph>
|
||
<paragraph id="8BD47615FFB9D500FFC8F18E34A2DC78" blockId="5.[111,649,913,1700]" lastBlockId="5.[674,1212,216,781]" pageId="5" pageNumber="364">
|
||
We agree with the diagnosis of
|
||
<taxonomicName id="4C6B0D96FFB9D500FE7DF18E354DDDBC" authorityName="Glaessner" authorityYear="1945" box="[452,593,945,971]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FAA07FFB9D500FE7DF18E354DDDBC" box="[452,593,945,971]" italics="true" pageId="5" pageNumber="364">Carpopenaeus</emphasis>
|
||
</taxonomicName>
|
||
proposed by
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FF6BF1EF369FDD9C" author="GLAESSNER M. - F." box="[210,387,976,1003]" pageId="5" pageNumber="364" pagination="694 - 707" refId="ref7202" refString="GLAESSNER M. - F. 1945. - Cretaceous Crustacea from Mount Lebanon, Syria. Annals and Magazine of Natural History 11 (12): 694 - 707." type="journal article" year="1945">Glaessner (1945)</bibRefCitation>
|
||
and revised by
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FDA6F1EF37A7DA7D" author="GARASSINO A." pageId="5" pageNumber="364" pagination="3 - 27" refId="ref6978" refString="GARASSINO A. 1994. - The macruran decapod crustaceans of the Upper Cretaceous of Lebanon. Paleontologia Lombarda (nuova serie) 3: 3 - 27." type="journal article" year="1994">Garassino (1994)</bibRefCitation>
|
||
. The newly described form here is clearly a representative of this genus based upon the following main features:very long rostrum with one subrostral tooth; longitudinal carina on the carapace,pereiopods 2 and 3with multiarticulated carpus, and exopod with a diaresis.
|
||
<taxonomicName id="4C6B0D96FFB9D500FF6AF6B03556DADF" authority="Charbonnier" authorityName="Petit & Charbonnier" authorityYear="2012" box="[211,586,1166,1193]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="garassinoi" status="sp. nov.">
|
||
<emphasis id="B91FAA07FFB9D500FF6AF6B036DCDADF" box="[211,448,1167,1193]" italics="true" pageId="5" pageNumber="364">Carpopenaeus garassinoi</emphasis>
|
||
Charbonnier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C177CFFB9D500FDF7F6B03599DADE" box="[590,645,1167,1193]" pageId="5" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from theTithonian species
|
||
<taxonomicName id="4C6B0D96FFB9D500FE78F69136E2DA9F" authority="Schweigert & Garassino, 2005" authorityName="Schweigert & Garassino" authorityYear="2005" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="peterbuergeri">
|
||
<emphasis id="B91FAA07FFB9D500FE78F69137DDDA90" italics="true" pageId="5" pageNumber="364">Carpopenaeus peterbuergeri</emphasis>
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FF7FF6F236E2DA9F" author="SCHWEIGERT G. & GARASSINO A." box="[198,510,1229,1256]" pageId="5" pageNumber="364" pagination="490 - 502" refId="ref8027" refString="SCHWEIGERT G. & GARASSINO A. 2005. - First record of the shrimp genus Carpopenaeus Glaessner, 1945 (Crustacea: Decapoda: Carpopenaeidae) from the Upper Jurassic. Neues Jahrbuch fur Geologie und Paleontologie Monatshefte 2005 (8): 490 - 502." type="journal article" year="2005">
|
||
Schweigert & Garassino,
|
||
<quantity id="4C93DBF0FFB9D500FE71F6F236E2DA9F" box="[456,510,1229,1256]" metricMagnitude="1" metricUnit="m" metricValue="5.0927" pageId="5" pageNumber="364" unit="in" value="2005.0">2005</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in that it has: 1) much thinner pereiopods 4 and 5; 2) pereiopods 2 and 3 of equal length whereas in
|
||
<taxonomicName id="4C6B0D96FFB9D500FE5FF732359ADB50" authorityName="Schweigert & Garassino" authorityYear="2005" box="[486,646,1293,1319]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="peterbuergeri">
|
||
<emphasis id="B91FAA07FFB9D500FE5FF732359ADB50" box="[486,646,1293,1319]" italics="true" pageId="5" pageNumber="364">C. peterbuergeri</emphasis>
|
||
</taxonomicName>
|
||
pereiopods 2 longer than pereiopods 3; and 3) the absence of a branchiocardiac carina well-developed in
|
||
<taxonomicName id="4C6B0D96FFB9D500FFD6F753361BDBF1" authorityName="Schweigert & Garassino" authorityYear="2005" box="[111,263,1388,1414]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="peterbuergeri">
|
||
<emphasis id="B91FAA07FFB9D500FFD6F753361BDBF1" box="[111,263,1388,1414]" italics="true" pageId="5" pageNumber="364">C.peterbuergeri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6B0D96FFB9D500FEACF753359BDBF1" authority="Charbonnier" authorityName="Petit & Charbonnier" authorityYear="2012" box="[277,647,1388,1414]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="garassinoi" status="sp. nov.">
|
||
<emphasis id="B91FAA07FFB9D500FEACF75336E3DBF1" box="[277,511,1388,1414]" italics="true" pageId="5" pageNumber="364">Carpopenaeus garassinoi</emphasis>
|
||
Charbonnier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C177CFFB9D500FFD6F7B337BADBD1" box="[111,166,1420,1446]" pageId="5" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from both Cenomanian species
|
||
<taxonomicName id="4C6B0D96FFB9D500FD8DF7B3364CDBB1" authority="Glaessner, 1945" authorityName="Glaessner" authorityYear="1945" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="callirostris">
|
||
<emphasis id="B91FAA07FFB9D500FD8DF7B3355ADBD1" box="[564,582,1420,1446]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FDEDF7B337B6DBB2" italics="true" pageId="5" pageNumber="364">callirostris</emphasis>
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FF16F794364CDBB1" author="GLAESSNER M. - F." box="[175,336,1451,1478]" pageId="5" pageNumber="364" pagination="694 - 707" refId="ref7202" refString="GLAESSNER M. - F. 1945. - Cretaceous Crustacea from Mount Lebanon, Syria. Annals and Magazine of Natural History 11 (12): 694 - 707." type="journal article" year="1945">Glaessner, 1945</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B0D96FFB9D500FEC6F79337ACDB91" authority="(Dames, 1886)" baseAuthorityName="Dames" baseAuthorityYear="1886" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FEC6F793368CDBB1" box="[383,400,1452,1478]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FE25F7933537DBB2" box="[412,555,1452,1477]" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FD8FF79437B4DB91" author="DAMES W." pageId="5" pageNumber="364" pagination="551 - 576" refId="ref6365" refString="DAMES W. 1886. - Ueber einige Crustaceen aus den Kreideablagerungen des Libanon. Zeitschrift der Deutschen Geologischen Gesellschaft 38 (3): 551 - 576." type="journal article" year="1886">Dames, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in that it has: 1) a rostrum with four basal suprarostral spines and only one median suprarostral spine whereas the rostrum of
|
||
<taxonomicName id="4C6B0D96FFB9D500FE21F434353FD853" authorityName="Glaessner" authorityYear="1945" box="[408,547,1546,1573]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="callirostris">
|
||
<emphasis id="B91FAA07FFB9D500FE21F43436B6D852" box="[408,426,1547,1573]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FE0EF435353FD853" box="[439,547,1546,1572]" italics="true" pageId="5" pageNumber="364">Callirostris</emphasis>
|
||
</taxonomicName>
|
||
shows six suprarostal spines uniformly distributed and that of
|
||
<taxonomicName id="4C6B0D96FFB9D500FFC8F475363BD813" baseAuthorityName="Dames" baseAuthorityYear="1886" box="[113,295,1610,1636]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FFC8F475379FD813" box="[113,131,1610,1636]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FF2AF474363BD813" box="[147,295,1611,1636]" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
</taxonomicName>
|
||
is shorter with seven suprarostal spines and one distal subrostral spine; 2) the short longitudinal lateral carina on the carapace that is more elongated in
|
||
<taxonomicName id="4C6B0D96FFB9D500FC91F2E734A9DE85" authorityName="Glaessner" authorityYear="1945" box="[808,949,216,242]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="callirostris">
|
||
<emphasis id="B91FAA07FFB9D500FC91F2E73426DE85" box="[808,826,216,242]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FCF0F2E734A9DE85" box="[841,949,216,242]" italics="true" pageId="5" pageNumber="364">Callirostris</emphasis>
|
||
</taxonomicName>
|
||
and duplicated in
|
||
<taxonomicName id="4C6B0D96FFB9D500FBCFF2E73404DF66" baseAuthorityName="Dames" baseAuthorityYear="1886" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FBCFF2E73394DE85" box="[1142,1160,216,242]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FB2DF2E63404DF66" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
</taxonomicName>
|
||
; 3) a third maxilliped with ventral margin serrated and composed of small rounded teeth whereas those of
|
||
<taxonomicName id="4C6B0D96FFB9D500FD41F3083460DF26" authorityName="Glaessner" authorityYear="1945" box="[760,892,311,337]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="callirostris">
|
||
<emphasis id="B91FAA07FFB9D500FD41F3083416DF26" box="[760,778,311,337]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FCA1F3083460DF26" box="[792,892,311,337]" italics="true" pageId="5" pageNumber="364">callirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B0D96FFB9D500FC14F3083347DF26" baseAuthorityName="Dames" baseAuthorityYear="1886" box="[941,1115,311,337]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FC14F30834A3DF26" box="[941,959,311,337]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FC73F3073347DF26" box="[970,1115,312,337]" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
</taxonomicName>
|
||
display a row of small spines; 4) the well-developed basipodite; and 5) the distal part of each uropod is covered by small pits whereas the ornamentation is smooth on the uropods of
|
||
<taxonomicName id="4C6B0D96FFB9D500FCFEF38934D7DFB8" authorityName="Glaessner" authorityYear="1945" box="[839,971,437,464]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="callirostris">
|
||
<emphasis id="B91FAA07FFB9D500FCFEF3893445DFA7" box="[839,857,438,464]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FCDEF38A34D7DFB8" box="[871,971,437,463]" italics="true" pageId="5" pageNumber="364">callirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B0D96FFB9D500FBB8F38933AEDFB8" baseAuthorityName="Dames" baseAuthorityYear="1886" box="[1025,1202,438,464]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FBB8F389330FDFA7" box="[1025,1043,438,464]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FB98F38933AEDFB8" box="[1057,1202,438,463]" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the uropodal exopod of
|
||
<taxonomicName id="4C6B0D96FFB9D500FBB3F3E933A5DF98" baseAuthorityName="Dames" baseAuthorityYear="1886" box="[1034,1209,470,496]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="septemspinatus">
|
||
<emphasis id="B91FAA07FFB9D500FBB3F3E93300DF87" box="[1034,1052,470,496]" italics="true" pageId="5" pageNumber="364">C</emphasis>
|
||
.
|
||
<emphasis id="B91FAA07FFB9D500FB91F3E933A5DF98" box="[1064,1209,470,495]" italics="true" pageId="5" pageNumber="364">septemspinatus</emphasis>
|
||
</taxonomicName>
|
||
does not show any diaresis.
|
||
</paragraph>
|
||
<paragraph id="8BD47615FFB9D500FD02F02A3379DD7A" blockId="5.[674,1212,216,781]" pageId="5" pageNumber="364">
|
||
In conclusion,
|
||
<taxonomicName id="4C6B0D96FFB9D500FCEEF02A35D3DC39" authority="Charbonnier" authorityName="Petit & Charbonnier" authorityYear="2012" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="species" species="garassinoi" status="sp. nov.">
|
||
<emphasis id="B91FAA07FFB9D500FCEEF02A3350DC59" box="[855,1100,533,559]" italics="true" pageId="5" pageNumber="364">Carpopenaeus garassinoi</emphasis>
|
||
Charbonnier
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C177CFFB9D500FD6FF00A340EDC38" box="[726,786,565,591]" pageId="5" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
|
||
from the Santonian Sahel Alma Lagerstätte is the youngest representative of the genus
|
||
<taxonomicName id="4C6B0D96FFB9D500FD1FF04B34C0DCF9" authority="Glaessner, 1945" authorityName="Glaessner" authorityYear="1945" box="[678,988,627,654]" class="Malacostraca" family="Carpopenaeidae" genus="Carpopenaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FAA07FFB9D500FD1FF04B342EDCF9" box="[678,818,628,654]" italics="true" pageId="5" pageNumber="364">Carpopenaeus</emphasis>
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FC8EF04B34C0DCF9" author="GLAESSNER M. - F." box="[823,988,627,654]" pageId="5" pageNumber="364" pagination="694 - 707" refId="ref7202" refString="GLAESSNER M. - F. 1945. - Cretaceous Crustacea from Mount Lebanon, Syria. Annals and Magazine of Natural History 11 (12): 694 - 707." type="journal article" year="1945">Glaessner, 1945</bibRefCitation>
|
||
</taxonomicName>
|
||
which was previously known from the Tithonian of the Lithographic Limestones of Zandt,
|
||
<collectingCountry id="F37C3685FFB9D500FC1CF08C3312DCBA" box="[933,1038,691,717]" name="Germany" pageId="5" pageNumber="364">Germany</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFFA0BE4FFB9D500FBA7F08C3445DC9A" author="SCHWEIGERT G. & GARASSINO A." pageId="5" pageNumber="364" pagination="490 - 502" refId="ref8027" refString="SCHWEIGERT G. & GARASSINO A. 2005. - First record of the shrimp genus Carpopenaeus Glaessner, 1945 (Crustacea: Decapoda: Carpopenaeidae) from the Upper Jurassic. Neues Jahrbuch fur Geologie und Paleontologie Monatshefte 2005 (8): 490 - 502." type="journal article" year="2005">Schweigert & Garassino 2005</bibRefCitation>
|
||
) and from the Cenomanian of
|
||
<collectingCountry id="F37C3685FFB9D500FD1EF0CD341ADD7B" box="[679,774,754,780]" name="Lebanon" pageId="5" pageNumber="364">Lebanon</collectingCountry>
|
||
(Hakel, Hadjoula Lagerstätten).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |