633 lines
74 KiB
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633 lines
74 KiB
XML
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<mods:title id="22352C1644361A7C4C92669BD9C6680D">A revision of the Geothelphusa leIJicerIJix species group (Crustacea: Decapoda: Brachyura: Potamidae), with descriptions of three new species</mods:title>
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<mods:namePart id="E3D2E8730C418A59C659BE0054769D58">Naruse, Tohru</mods:namePart>
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<mods:namePart id="1E34FE4F85998455098A049AE91B4B34">Shokita, Shigemitsu</mods:namePart>
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<mods:namePart id="DA9FB6C45C8DD6E3BB540418717CF4CE">Ng, Peter K. L.</mods:namePart>
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<mods:title id="11F26D881BEBA88C39D0E950263DE49D">Journal of Natural History</mods:title>
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<mods:date id="E95A443EE951466A3101E9E881C8B5DE">2006</mods:date>
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<mods:number id="8DB25E37DE1D3BFE43A763A30798DF8E">2006-07-26</mods:number>
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<mods:number id="16153A8B933E251EBCBD1D622B17593F">13 - 14</mods:number>
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<subSubSection id="D6F99FC41D7FFFA822F8FF5171C02194" box="[418,909,137,161]" pageId="9" pageNumber="766" type="nomenclature">
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<paragraph id="9E5CCC4F1D7FFFA822F8FF5171C02194" blockId="9.[418,909,137,193]" box="[418,909,137,161]" pageId="9" pageNumber="766">
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<heading id="C5147B231D7FFFA822F8FF5171C02194" box="[418,909,137,161]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="766" reason="4">
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<emphasis id="AC97105D1D7FFFA822F8FF5170B62194" bold="true" box="[418,763,137,161]" italics="true" pageId="9" pageNumber="766">
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<taxonomicName id="59E3B7CC1D7FFFA822F8FF5170192194" ID-CoL="7NVV5" authorityName="Stimpson" authorityYear="1858" box="[418,596,137,161]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="766" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
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grandioIJata
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</emphasis>
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new species
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||
</heading>
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</paragraph>
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<subSubSection id="D6F99FC41D7FFFA8217CFF71713221C7" pageId="9" pageNumber="766" type="description">
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<paragraph id="9E5CCC4F1D7FFFA8217CFF7170B221F4" blockId="9.[418,909,137,193]" box="[550,767,169,193]" pageId="9" pageNumber="766">
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(
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<figureCitation id="06D8D0CA1D7FFFA82174FF7170D121F4" box="[558,668,169,193]" captionStart="Figure 4" captionStartId="10.[138,198,1513,1532]" captionTargetBox="[9,1186,133,1471]" captionTargetId="figure-1@10.[138,1187,133,1477]" captionTargetPageId="10" captionText="Figure 4. Geothelphusa grandioυata new species. (a) Dorsal view; (b) dorsal view of carapace; (c) ventral side of G1; (d) dorsal side of G1. (a) Paratype, ZRC 2006.0076, CL 44.8 mm; (b–d) holotype, ZLKU m. 1094, CL 40.5 mm. Scale bars: 10 mm (b); 5mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.4581770" httpUri="https://zenodo.org/record/4581770/files/figure.png" pageId="9" pageNumber="766">Figures 4</figureCitation>
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||
,
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||
<figureCitation id="06D8D0CA1D7FFFA821F1FF71708421F4" box="[683,713,169,193]" captionStart="Figure 5" captionStartId="11.[125,185,755,774]" captionTargetBox="[125,1172,133,719]" captionTargetId="figure-243@11.[125,1172,133,719]" captionTargetPageId="11" captionText="Figure 5. Fossils of Geothelphusa grandioυata new species and G. tenuimanus (Miyake and Minei, 1965). (a) Chela of G. grandioυata (RUMF, palm length 28.6+ mm, width 23.7+ mm); (b) chela of G. tenuimanus (RUMF-GF-11, palm length 18.6 mm, palm width 17.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.4581774" httpUri="https://zenodo.org/record/4581774/files/figure.png" pageId="9" pageNumber="766">5a</figureCitation>
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||
,
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||
<figureCitation id="06D8D0CA1D7FFFA8218DFF7170BB21F4" box="[727,758,169,193]" captionStart="Figure 8" captionStartId="19.[125,185,1284,1303]" captionTargetBox="[9,1174,135,1281]" captionTargetId="figure-87@19.[125,1174,133,1248]" captionTargetPageId="19" captionText="Figure 8. Frontal view of Geothelphusa leυicerυix (Rathbun, 1898) and G. grandioυata new species. (a) Geothelphusa leυicerυix, ZRC 2006.0074, male, CL 35.3 mm; (b) G. grandioυata, paratype, ZRC 2006.0076, male, CL 44.8 mm." figureDoi="http://doi.org/10.5281/zenodo.4581781" httpUri="https://zenodo.org/record/4581781/files/figure.png" pageId="9" pageNumber="766">8b</figureCitation>
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||
)
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||
</paragraph>
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||
<paragraph id="9E5CCC4F1D7FFFA82327FF03713221C7" blockId="9.[125,1173,218,530]" box="[125,895,218,242]" pageId="9" pageNumber="766">
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||
<taxonomicName id="59E3B7CC1D7FFFA82327FF0373C121C7" authority="" box="[125,396,218,242]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="766" phylum="Arthropoda" rank="subGenus" subGenus="Geothelphusa">
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||
<emphasis id="AC97105D1D7FFFA82327FF03729021C7" box="[125,221,219,242]" italics="true" pageId="9" pageNumber="766">Potamon</emphasis>
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||
(
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||
<emphasis id="AC97105D1D7FFFA823ABFF0273CF21C7" box="[241,386,218,242]" italics="true" pageId="9" pageNumber="766">Geothelphusa</emphasis>
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||
)
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||
</taxonomicName>
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||
<emphasis id="AC97105D1D7FFFA822CFFF0273B721C7" box="[405,506,218,242]" italics="true" pageId="9" pageNumber="766">leυicerυix</emphasis>
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||
:
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<bibRefCitation id="FA72B1BE1D7FFFA82156FF02716721C7" author="Miyake S & Minei H" box="[524,810,218,242]" pageId="9" pageNumber="766" pagination="377 - 382" refId="ref8786" refString="Miyake S, Minei H. 1965. A new fresh-water crab, Potamon (Geothelphusa) tenuimanus sp. nov. from Okinawajima, the Ryukyu Islands. Science Bulletin of the Faculty of Agriculture, Kyushu University 4: 377 - 382. Plate 21. (Jpn with Eng summary)." type="journal article" year="1965">Miyake and Minei 1965</bibRefCitation>
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||
, p 377.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="D6F99FC41D7FFFA82327FF2376102327" pageId="9" pageNumber="766" type="reference_group">
|
||
<paragraph id="9E5CCC4F1D7FFFA82327FF2376102327" blockId="9.[125,1173,218,530]" pageId="9" pageNumber="766">
|
||
<emphasis id="AC97105D1D7FFFA82327FF23733A2026" box="[125,375,251,275]" italics="true" pageId="9" pageNumber="766">
|
||
<taxonomicName id="59E3B7CC1D7FFFA82327FF2373412026" ID-CoL="7NVV5" authorityName="Stimpson" authorityYear="1858" box="[125,268,251,275]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="766" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
leυicerυix
|
||
</emphasis>
|
||
:
|
||
<bibRefCitation id="FA72B1BE1D7FFFA822DDFF23705C2026" author="Minei H" box="[391,529,251,275]" pageId="9" pageNumber="766" pagination="94 - 99" refId="ref8632" refString="Minei H. 1968. Fresh-water crabs of Japan. Nature Study 4: 94 - 99. (Jpn)." type="journal article" year="1968">Minei 1968</bibRefCitation>
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||
, p 99, Figure 11;
|
||
<bibRefCitation id="FA72B1BE1D7FFFA82180FF23712E2026" author="Minei H" box="[730,867,251,275]" pageId="9" pageNumber="766" pagination="203 - 226" refId="ref8654" refString="Minei H. 1973. Potamoid crabs of the Ryukyu Islands, with descriptions of five new species (Crustacea, Decapoda, Potamoidea). Journal of the Faculty of Agriculture, Kyushu University 17: 203 - 226." type="journal article" year="1973">Minei 1973</bibRefCitation>
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||
, p 211,
|
||
<figureCitation id="06D8D0CA1D7FFFA820E5FF2376642026" box="[959,1065,251,275]" captionStart="Figure 6" captionStartId="14.[138,198,1524,1543]" captionTargetBox="[9,1183,117,1467]" captionTargetId="figure-1@14.[138,1185,117,1477]" captionTargetPageId="14" captionText="Figure 6. Geothelphusa kumejima new species. (a) Dorsal view; (b) dorsal view of carpace; (c) ventral side of G1; (d) dorsal side of G1; (e) G2. (a) Paratype, ZRC 2006.0077, male, CL 35.0 mm; (b–e) holotype, RUMF-ZC-144, male, CL 39.5 mm. Scale bars: 10 mm (b); 5 mm (c–e)." figureDoi="http://doi.org/10.5281/zenodo.4581777" httpUri="https://zenodo.org/record/4581777/files/figure.png" pageId="9" pageNumber="766">Figures 6</figureCitation>
|
||
,
|
||
<figureCitation id="06D8D0CA1D7FFFA8276DFF23760A2026" box="[1079,1095,251,275]" captionStart="Figure 9" captionStartId="20.[138,198,1234,1253]" captionTargetBox="[138,1187,133,1198]" captionTargetId="figure-151@20.[138,1187,133,1198]" captionTargetPageId="20" captionText="Figure 9. Frontal view of Geothelphusa kumejima new species and Geothelphusa iheya new species. (a) Geothelphusa kumejima, paratype, ZRC 2006.0077, male, CL 35.0 mm; (b) G. iheya, paratype, ZRC 2006.0078, male, CL 28.7 mm." figureDoi="http://doi.org/10.5281/zenodo.4581783" httpUri="https://zenodo.org/record/4581783/files/figure.png" pageId="9" pageNumber="766">9</figureCitation>
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||
(C, D) (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8225FFEC373C82006" author="Sakai T" box="[261,389,283,307]" pageId="9" pageNumber="766" refId="ref9340" refString="Sakai T. 1976. Crabs of Japan and the adjacent seas. Tokyo: Kodansha. 773 p (Eng text), 461 p (Jpn text), 251 p (plate volume)." type="book" year="1976">Sakai 1976</bibRefCitation>
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||
, p 558 (English text), 346 (Japanese text) (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8209DFEC372982066" author="Gima H & Shokita S" pageId="9" pageNumber="766" pagination="9 - 15" refId="ref8401" refString="Gima H, Shokita S. 1980. Distribution of potamonid crabs in the Yona River of Okinawa Island, the Ryukyus. Biological Magazine, Okinawa 18: 9 - 15. (Jpn with Eng abstract)." type="journal article" year="1980">Gima and Shokita 1980</bibRefCitation>
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, p 9,
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<figureCitation id="06D8D0CA1D7FFFA82242FEE373CF2066" box="[280,386,315,339]" captionStart="Figure 7" captionStartId="17.[125,185,1485,1504]" captionTargetBox="[9,1174,135,1446]" captionTargetId="figure-0@17.[125,1174,133,1449]" captionTargetPageId="17" captionText="Figure 7. Geothelphusa iheya new species. (a) Dorsal view; (b) dorsal view of carpace; (c) ventral side of G1; (d) dorsal side of G1; (e) G2. (a) Paratype, ZRC 2006.0078, male, CL 28.7 mm; (b–e) holotype, RUMF-ZC-148, male, CL 32.6 mm. Scale bars: 10 mm (b); 1 mm (c–e)." figureDoi="http://doi.org/10.5281/zenodo.4581779" httpUri="https://zenodo.org/record/4581779/files/figure.png" pageId="9" pageNumber="766">Figures 7</figureCitation>
|
||
,
|
||
<figureCitation id="06D8D0CA1D7FFFA822C8FEE373EF2066" box="[402,418,315,339]" captionStart="Figure 8" captionStartId="19.[125,185,1284,1303]" captionTargetBox="[9,1174,135,1281]" captionTargetId="figure-87@19.[125,1174,133,1248]" captionTargetPageId="19" captionText="Figure 8. Frontal view of Geothelphusa leυicerυix (Rathbun, 1898) and G. grandioυata new species. (a) Geothelphusa leυicerυix, ZRC 2006.0074, male, CL 35.3 mm; (b) G. grandioυata, paratype, ZRC 2006.0076, male, CL 44.8 mm." figureDoi="http://doi.org/10.5281/zenodo.4581781" httpUri="https://zenodo.org/record/4581781/files/figure.png" pageId="9" pageNumber="766">8</figureCitation>
|
||
;
|
||
<bibRefCitation id="FA72B1BE1D7FFFA822E8FEE370732066" author="Minei H" box="[434,574,315,339]" pageId="9" pageNumber="766" pagination="79 - 92" refId="ref8695" refString="Minei H. 1981. Distribution and general habitat of the freshwater crabs of Japan. In: Yamaguchi T, editor. Ecological studies of coastal marine and freshwater crabs. Report for the Grant-in-Aid for Co-operative Research, 1978 - 1980. Tokyo: Ministry of Education. p 79 - 92. (Jpn)." type="book chapter" year="1981">Minei 1981</bibRefCitation>
|
||
, p 80, Tables 2, 3 Plate 3 (lower) (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8271AFEE372982046" author="Miyake S" pageId="9" pageNumber="766" refId="ref8754" refString="Miyake S. 1983. Japanese crustacean decapods and stomatopods in color. Volume 2, Brachyura (crabs), Osaka: Hoikusha. 277 p. (Jpn)." type="book" year="1983">Miyake 1983</bibRefCitation>
|
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, p 244, plate 51;
|
||
<bibRefCitation id="FA72B1BE1D7FFFA822C2FE83707F2046" author="Shokita S" box="[408,562,347,371]" pageId="9" pageNumber="766" pagination="3 - 13" refId="ref9449" refString="Shokita S. 1990. Inland-water macrofauna and their fluctuations in the northern part of Okinawa Island, the Ryukyus. Biological Magazine, Okinawa 27: 3 - 13. (Jpn with Eng abstract and captions)." type="journal article" year="1990">Shokita 1990</bibRefCitation>
|
||
, Table 2 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA82051FE8371972046" author="Anonymous" box="[779,986,347,371]" pageId="9" pageNumber="766" refId="ref8085" refString="Anonymous. 1991. Brackish and freshwater decapod crustaceans. In: Environment Agency, editor. Threatened wildlife in Japan: Red Data Book on invertebrates. Tokyo: Wildlife Research Center. 211 - 230. (Jpn)." type="book" year="1991">Anonymous 1991</bibRefCitation>
|
||
, p 219 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA823C2FEA373D520A6" author="Suzuki H & Sato M" box="[152,408,379,403]" pageId="9" pageNumber="766" refId="ref10004" refString="Suzuki H, Sato M. 1994. Kagoshima nature guide. Freshwater shrimps, crayfish, and crabs. Fukuoka: The Nishinippon Shimbun. 141 p. (Jpn)." type="book" year="1994">Suzuki and Sato 1994</bibRefCitation>
|
||
, p 114 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA82100FEA3714720A6" author="Shy J-Y & Ng PKL & Yu H-P" box="[602,778,379,403]" pageId="9" pageNumber="766" pagination="781 - 846" refId="ref9792" refString="Shy J-Y, Ng PKL, Yu H-P. 1994. Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology 42: 781 - 846." type="journal article" year="1994">Shy et al. 1994</bibRefCitation>
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||
, p 784;
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8203FFEA3761320A6" author="Ng PKL & Shokita S" box="[869,1118,379,403]" pageId="9" pageNumber="766" pagination="1 - 7" refId="ref9134" refString="Ng PKL, Shokita S. 1995. Ryukyum, a new genus of terrestrial crab from the Ryukyu Islands (Brachyura: Potamidae). Crustacean Research 24: 1 - 7." type="journal article" year="1995">Ng and Shokita 1995</bibRefCitation>
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||
, p 3;
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||
<bibRefCitation id="FA72B1BE1D7FFFA823C2FE4373242086" author="Anonymous" box="[152,361,411,435]" pageId="9" pageNumber="766" pagination="362 - 385" refId="ref8127" refString="Anonymous. 1996. Crustaceans. In: Nature Conservation Division (Department of Environment and Health, Okianawa Prefectural Government), editor: Red Data Okinawa. Naha: Nature Conservation Division (Department of Environment and Health, Okianawa Prefectural Government). p 362 - 385. (Jpn)." type="book chapter" year="1996">Anonymous 1996</bibRefCitation>
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, p 362 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8216BFE4370822086" author="Shokita S" box="[561,719,411,435]" pageId="9" pageNumber="766" pagination="343 - 353" refId="ref9489" refString="Shokita S. 1996. The origin of land-locked freshwater shrimps and potamoids from the Ryukyu Islands, Southern Japan. Journal of Geography 105: 343 - 353. (Jpn with Eng abstract and captions)." type="journal article" year="1996">Shokita 1996</bibRefCitation>
|
||
, p 348, Photograph 1;
|
||
<bibRefCitation id="FA72B1BE1D7FFFA82087FE4376DD2086" author="Yoshigou H" box="[989,1168,411,435]" pageId="9" pageNumber="766" pagination="17 - 26" refId="ref10127" refString="Yoshigou H. 1999. Potamidae (Crustacea: Decapoda: Brachyura) of Japan. Journal of the Hiba Society of Natural History 191: 17 - 26. Plates I, II. (Jpn)." type="journal article" year="1999">Yoshigou 1999</bibRefCitation>
|
||
, p 20, Plate 1(F) (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA822E0FE63701F20E6" author="Segawa R" box="[442,594,443,467]" pageId="9" pageNumber="766" pagination="241 - 245" refId="ref9378" refString="Segawa R. 2000. Molecular phylogenetic study of potamoid crabs in Ryukyu Islands. Kaiyo Monthly 32: 241 - 245. (Jpn)." type="journal article" year="2000">Segawa 2000</bibRefCitation>
|
||
, p 243, Table 1,
|
||
<figureCitation id="06D8D0CA1D7FFFA82051FE63713E20E6" box="[779,883,443,467]" captionStart="Figure 1" captionStartId="3.[408,468,1663,1682]" captionTargetBox="[125,1172,776,1627]" captionTargetId="figure-199@3.[125,1174,773,1627]" captionTargetPageId="3" captionText="Figure 1. Map of the Okinawa Group in East Asia." figureDoi="http://doi.org/10.5281/zenodo.4581762" httpUri="https://zenodo.org/record/4581762/files/figure.png" pageId="9" pageNumber="766">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="06D8D0CA1D7FFFA820DAFE6371C220E6" box="[896,911,443,467]" captionStart="Figure 2" captionStartId="5.[125,185,1383,1402]" captionTargetBox="[9,1174,133,1345]" captionTargetId="figure-71@5.[125,1174,133,1347]" captionTargetPageId="5" captionText="Figure 2. Geothelphusa leυicerυix (Rathbun, 1905), holotype, USNM 20642, male, CL 48.3 mm. (a) Dorsal view; (b) frontal view of carapace; (c) ventral side of left G1; (d) dorsal side of left G1. Scale bars: 10 mm (b); 5 mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.4581766" httpUri="https://zenodo.org/record/4581766/files/figure.png" pageId="9" pageNumber="766">2</figureCitation>
|
||
(partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA82758FE63729820C6" author="Shokita S & Naruse T & Fujii H" pageId="9" pageNumber="766" pagination="443 - 448" refId="ref9704" refString="Shokita S, Naruse T, Fujii H. 2002. Geothelphusa miyakoensis, a new species of freshwater crab (Crustacea: Decapoda: Brachyura: Potamidae) from Miyako Island, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 50: 443 - 448." type="journal article" year="2002">Shokita et al. 2002</bibRefCitation>
|
||
, p 446 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA822F3FE03708120C6" author="Kasai H & Naruse T" box="[425,716,475,499]" pageId="9" pageNumber="766" pagination="282 - 288" refId="ref8507" refString="Kasai H, Naruse T. 2003. Potamidae. In: Nishida M, Shikatani N, Shokita S, editors. The flora and fauna of inland waters in the Ryukyu Islands. Tokyo: Tokai University Press. p 282 - 288. (Jpn)." type="book chapter" year="2003">Kasai and Naruse 2003</bibRefCitation>
|
||
, p 283, Figures 67B, 71, 72 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA823C2FE2273702327" author="Naruse T" box="[152,317,506,530]" pageId="9" pageNumber="766" refId="ref8841" refString="Naruse T. 2005 a. Geothelphusa leυicerυix. In: Nature Conservation Division (Department of Cultural and Environmental Affairs, Okinawa Prefectural Government), editor: Threatened wildlife in Okinawa (animals) - Red Data Okinawa. 2 nd ed, Naha: Nature Conservation Division (Department of Cultural and Environmental Affairs, Okinawa Prefectural Government). 190 p. Plate (Crustacea: upper left). (Jpn)." type="book" year="2005">Naruse 2005a</bibRefCitation>
|
||
, p 190 (partim);
|
||
<bibRefCitation id="FA72B1BE1D7FFFA8215AFE2270E62327" author="Shokita S" box="[512,683,506,530]" pageId="9" pageNumber="766" refId="ref9642" refString="Shokita S. 2005 b. Geothelphusa leυicerυix. In: Ministry of the Environment, editor: Threatened wildlife of Japan- Red Data Book. 2 nd ed. Volume 7, Invertebrate (except Insecta and Mollusca), Tokyo: Japan Wildlife Research Center. 56 p. Plate 3 (middle right). (Jpn)." type="book" year="2005">Shokita 2005b</bibRefCitation>
|
||
, p 56 (partim), Plate 3 (middle right).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D6F99FC41D7FFFAD2327FD93708C27A0" lastPageId="12" lastPageNumber="769" pageId="9" pageNumber="766" type="materials_examined">
|
||
<paragraph id="9E5CCC4F1D7FFFA82327FD9373012356" blockId="9.[125,332,587,611]" box="[125,332,587,611]" pageId="9" pageNumber="766">
|
||
<emphasis id="AC97105D1D7FFFA82327FD9373012356" box="[125,332,587,611]" italics="true" pageId="9" pageNumber="766">Material examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7FFFA82327FDA473B12447" blockId="9.[125,1172,636,1394]" pageId="9" pageNumber="766">
|
||
<materialsCitation id="2E8BC6121D7FFFA82327FDA473312381" collectingDate="1972-06-25" collectionCode="ZLKU" collectorName="H. Minei" location="Tamagusuku Village" pageId="9" pageNumber="766" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="415872ED1D7FFFA82327FDA472A023A1" box="[125,237,636,660]" pageId="9" pageNumber="766" type="holotype">Holotype</typeStatus>
|
||
: one male, CL
|
||
<quantity id="591B61AA1D7FFFA822FBFDA4704723A1" box="[417,522,636,660]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="9" pageNumber="766" unit="mm" value="40.5">40.5 mm</quantity>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA8214DFDA4702B23A1" box="[535,614,636,660]" pageId="9" pageNumber="766">ZLKU</collectionCode>
|
||
m. 1094, Fusato,
|
||
<location id="9B3C9A941D7FFFA82064FDA4767D23A1" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82064FDA4767D23A1" box="[830,1072,636,660]" name="Tamagusuku Village" pageId="9" pageNumber="766">Tamagusuku Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA8272CFDA472882381" pageId="9" pageNumber="766">H. Minei</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA8238FFD4473312381" box="[213,380,668,692]" pageId="9" pageNumber="766" value="1972-06-25">
|
||
<collectingDate id="FA1913671D7FFFA8238FFD4473312381" box="[213,380,668,692]" pageId="9" pageNumber="766" value="1972-06-25">25 June 1972</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="2E8BC6121D7FFFA822D0FD44761023E1" collectingDate="1972-06-25" collectionCode="ZLKU" collectorName="H. Minei" location="Tamagusuku Village" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="415872ED1D7FFFA822D0FD4473B32381" box="[394,510,668,692]" pageId="9" pageNumber="766" type="paratype">Paratypes</typeStatus>
|
||
: eight males, CL
|
||
<quantity id="591B61AA1D7FFFA8218DFD44713E2381" box="[727,883,668,692]" metricMagnitude="-2" metricUnit="m" metricValue="2.5700000000000003" metricValueMax="4.2700000000000005" metricValueMin="0.8699999999999999" pageId="9" pageNumber="766" unit="mm" value="25.7" valueMax="42.7" valueMin="8.7">8.7–42.7 mm</quantity>
|
||
, two females, CL
|
||
<quantity id="591B61AA1D7FFFA82709FD4472AA23E1" metricMagnitude="-2" metricUnit="m" metricValue="2.705" metricValueMax="3.42" metricValueMin="1.9899999999999998" pageId="9" pageNumber="766" unit="mm" value="27.05" valueMax="34.2" valueMin="19.9">19.9– 34.2 mm</quantity>
|
||
, three juveniles, CL
|
||
<quantity id="591B61AA1D7FFFA82284FD64703723E1" box="[478,634,700,724]" metricMagnitude="-2" metricUnit="m" metricValue="1.095" metricValueMax="1.23" metricValueMin="0.96" pageId="9" pageNumber="766" unit="mm" value="10.95" valueMax="12.3" valueMin="9.6">9.6–12.3 mm</quantity>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA821D2FD64709523E1" box="[648,728,700,724]" pageId="9" pageNumber="766">ZLKU</collectionCode>
|
||
m. 1094, data same as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82730FD6473712221" collectingDate="1994-12-03" collectionCode="ZRC" collectorName="S. Shokita" location="Ohgimi Village" municipality="Taiho River" pageId="9" pageNumber="766" specimenCode="ZRC 2006.0075" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA823A8FD04731623C1" box="[242,347,732,756]" metricMagnitude="-2" metricUnit="m" metricValue="4.39" pageId="9" pageNumber="766" unit="mm" value="43.9">43.9 mm</quantity>
|
||
,
|
||
<specimenCode id="CE4564341D7FFFA8223DFD04706423C1" box="[359,553,732,756]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="766">ZRC 2006.0075</specimenCode>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA8216EFD04708C23C1" box="[564,705,732,756]" pageId="9" pageNumber="766">Taiho River</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA82197FD04713323C1" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82197FD04713323C1" box="[717,894,732,756]" municipality="Taiho River" name="Ohgimi Village" pageId="9" pageNumber="766">Ohgimi Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA8209BFD04767A23C1" box="[961,1079,732,756]" pageId="9" pageNumber="766">S. Shokita</collectorName>
|
||
et al.,
|
||
<date id="EA5DEA8F1D7FFFA827DFFD0473712221" pageId="9" pageNumber="766" value="1994-12-03">
|
||
<collectingDate id="FA1913671D7FFFA827DFFD0473712221" pageId="9" pageNumber="766" value="1994-12-03">3 December 1994</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82213FD2473B22201" collectingDate="2001-07-05" collectionCode="T" collectorName="Naruse" location="Iji" municipality="Ohkuni Forest" pageId="9" pageNumber="766" specimenCode="ZRC 2006.0076" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA822ABFD2470172221" box="[497,602,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="4.4799999999999995" pageId="9" pageNumber="766" unit="mm" value="44.8">44.8 mm</quantity>
|
||
,
|
||
<specimenCode id="CE4564341D7FFFA8213DFD2471672221" box="[615,810,764,788]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="766">ZRC 2006.0076</specimenCode>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA8206CFD2471AC2221" box="[822,993,764,788]" pageId="9" pageNumber="766">Ohkuni Forest</collectingMunicipality>
|
||
Road, north of
|
||
<location id="9B3C9A941D7FFFA82327FCC472D42201" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82327FCC472D42201" box="[125,153,796,820]" municipality="Ohkuni Forest" name="Iji" pageId="9" pageNumber="766">Iji</location>
|
||
, coll.
|
||
<collectionCode id="F8F2548A1D7FFFA823BEFCC472B62201" box="[228,251,796,820]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="766">T</collectionCode>
|
||
.
|
||
<collectorName id="3316A9991D7FFFA82251FCC4732F2201" box="[267,354,796,820]" pageId="9" pageNumber="766">Naruse</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA8222BFCC473B22201" box="[369,511,796,820]" pageId="9" pageNumber="766" value="2001-07-05">
|
||
<collectingDate id="FA1913671D7FFFA8222BFCC473B22201" box="[369,511,796,820]" pageId="9" pageNumber="766" value="2001-07-05">5 July 2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82155FCC470FA2266" collectingDate="1983-08" collectorName="S. Shokita" location="Kunigami Village" municipality="Yona River" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA821E5FCC471652201" box="[703,808,796,820]" metricMagnitude="-2" metricUnit="m" metricValue="4.38" pageId="9" pageNumber="766" unit="mm" value="43.8">43.8 mm</quantity>
|
||
, RUMF-ZC-140,
|
||
<collectingMunicipality id="7E3856351D7FFFA8275CFCC476DD2201" box="[1030,1168,796,820]" pageId="9" pageNumber="766">Yona River</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA82327FCE373012266" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82327FCE373012266" box="[125,332,827,851]" municipality="Yona River" name="Kunigami Village" pageId="9" pageNumber="766">Kunigami Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA822CEFCE3705D2266" box="[404,528,827,851]" pageId="9" pageNumber="766">S. Shokita</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82144FCE370FA2266" box="[542,695,827,851]" pageId="9" pageNumber="766" value="1983-08">
|
||
<collectingDate id="FA1913671D7FFFA82144FCE370FA2266" box="[542,695,827,851]" pageId="9" pageNumber="766" value="1983-08">August 1983</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA8219FFCE370E922A6" collectingDate="1998-12" collectionCode="T" collectorName="S. Shokita & Y. Fujita & Nagai & H. Kawaguchi & Md. & Islam" county="Ishiyama" location="Kunigami Village" municipality="Hedo" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
two males, CL
|
||
<quantity id="591B61AA1D7FFFA82026FCE3766A2266" box="[892,1063,827,851]" metricMagnitude="-2" metricUnit="m" metricValue="2.945" metricValueMax="2.9699999999999998" metricValueMin="2.92" pageId="9" pageNumber="766" unit="mm" value="29.45" valueMax="29.7" valueMin="29.2">29.2–29.7 mm</quantity>
|
||
, RUMF- ZC-141, near
|
||
<collectingCounty id="773DB4C31D7FFFA8227EFC8373C22246" box="[292,399,859,883]" pageId="9" pageNumber="766">Ishiyama</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA822C6FC8373AD2246" box="[412,480,859,883]" pageId="9" pageNumber="766">Hedo</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA822B6FC8370F62246" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA822B6FC8370F62246" box="[492,699,859,883]" county="Ishiyama" municipality="Hedo" name="Kunigami Village" pageId="9" pageNumber="766">Kunigami Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA8205AFC8371372246" box="[768,890,859,883]" pageId="9" pageNumber="766">S. Shokita</collectorName>
|
||
,
|
||
<collectorName id="3316A9991D7FFFA820DCFC8371BC2246" box="[902,1009,859,883]" pageId="9" pageNumber="766">Y. Fujita</collectorName>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA820A4FC8376582246" box="[1022,1045,859,883]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="766">T</collectionCode>
|
||
.
|
||
<collectorName id="3316A9991D7FFFA8277BFC8376242246" box="[1057,1129,859,883]" pageId="9" pageNumber="766">Nagai</collectorName>
|
||
,
|
||
<collectorName id="3316A9991D7FFFA8272CFC8372B322A6" pageId="9" pageNumber="766">H. Kawaguchi</collectorName>
|
||
and
|
||
<collectorName id="3316A9991D7FFFA82261FCA3732022A6" box="[315,365,891,915]" pageId="9" pageNumber="766">Md.</collectorName>
|
||
S.
|
||
<collectorName id="3316A9991D7FFFA822CFFCA3739522A6" box="[405,472,891,915]" pageId="9" pageNumber="766">Islam</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA822BFFCA370E922A6" box="[485,676,891,915]" pageId="9" pageNumber="766" value="1998-12">
|
||
<collectingDate id="FA1913671D7FFFA822BFFCA370E922A6" box="[485,676,891,915]" pageId="9" pageNumber="766" value="1998-12">December 1998</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA821EAFCA3739A22E6" collectingDate="1998-12-27" collectionCode="T" collectorName="S. Shokita & Y. Fujita & Nagai & H. Kawaguchi & Md. & Islam" location="Ohgimi Village" municipality="Oshi River" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA82000FCA3718E22A6" box="[858,963,891,915]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="9" pageNumber="766" unit="mm" value="41.5">41.5 mm</quantity>
|
||
, RUMF-ZC-142,
|
||
<collectingMunicipality id="7E3856351D7FFFA82327FC4372B02286" box="[125,253,923,947]" pageId="9" pageNumber="766">Oshi River</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA82253FC4373F62286" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82253FC4373F62286" box="[265,443,923,947]" municipality="Oshi River" name="Ohgimi Village" pageId="9" pageNumber="766">Ohgimi Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA8215AFC4370362286" box="[512,635,923,947]" pageId="9" pageNumber="766">S. Shokita</collectorName>
|
||
,
|
||
<collectorName id="3316A9991D7FFFA821DDFC4370BF2286" box="[647,754,923,947]" pageId="9" pageNumber="766">Y. Fujita</collectorName>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA821A5FC43715B2286" box="[767,790,923,947]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="766">T</collectionCode>
|
||
.
|
||
<collectorName id="3316A9991D7FFFA82079FC4371262286" box="[803,875,923,947]" pageId="9" pageNumber="766">Nagai</collectorName>
|
||
,
|
||
<collectorName id="3316A9991D7FFFA8202DFC43766E2286" box="[887,1059,923,947]" pageId="9" pageNumber="766">H. Kawaguchi</collectorName>
|
||
, and
|
||
<collectorName id="3316A9991D7FFFA82738FC4376D92286" box="[1122,1172,923,947]" pageId="9" pageNumber="766">Md.</collectorName>
|
||
S.
|
||
<collectorName id="3316A9991D7FFFA823C5FC6372AF22E6" box="[159,226,955,979]" pageId="9" pageNumber="766">Islam</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA823AAFC63739A22E6" box="[240,471,955,979]" pageId="9" pageNumber="766" value="1998-12-27">
|
||
<collectingDate id="FA1913671D7FFFA823AAFC63739A22E6" box="[240,471,955,979]" pageId="9" pageNumber="766" value="1998-12-27">27 December 1998</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA822BFFC63712322C6" collectingDate="2001-11-27" collectionCode="T" collectorName="Naruse" location="Kunigami Village" municipality="Benoki Dam reserve" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one female, CL
|
||
<quantity id="591B61AA1D7FFFA821FDFC63715D22E6" box="[679,784,955,979]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="9" pageNumber="766" unit="mm" value="27.1">27.1 mm</quantity>
|
||
, RUMF-ZC-143, east of
|
||
<collectingMunicipality id="7E3856351D7FFFA82719FC63735922C6" pageId="9" pageNumber="766">Benoki Dam reserve</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA82278FC0373BC22C6" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82278FC0373BC22C6" box="[290,497,987,1011]" municipality="Benoki Dam reserve" name="Kunigami Village" pageId="9" pageNumber="766">Kunigami Village</location>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA822A7FC03705922C6" box="[509,532,987,1011]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="766">T</collectionCode>
|
||
.
|
||
<collectorName id="3316A9991D7FFFA82178FC03703422C6" box="[546,633,987,1011]" pageId="9" pageNumber="766">Naruse</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA821DCFC03712322C6" box="[646,878,987,1011]" pageId="9" pageNumber="766" value="2001-11-27">
|
||
<collectingDate id="FA1913671D7FFFA821DCFC03712322C6" box="[646,878,987,1011]" pageId="9" pageNumber="766" value="2001-11-27">27 November 2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82021FC0376142526" collectingDate="2001-07-05" collectionCode="NSMT, T" collectorName="Ohkuni Forest Road & Naruse" location="between Hiji and Iji" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA8277CFC0376C222C6" box="[1062,1167,987,1011]" metricMagnitude="-2" metricUnit="m" metricValue="3.76" pageId="9" pageNumber="766" unit="mm" value="37.6">37.6 mm</quantity>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA82327FC23729B2526" box="[125,214,1019,1043]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="9" pageNumber="766">NSMT</collectionCode>
|
||
,
|
||
<collectorName id="3316A9991D7FFFA823BDFC2373AF2526" box="[231,482,1019,1043]" pageId="9" pageNumber="766">Ohkuni Forest Road</collectorName>
|
||
,
|
||
<location id="9B3C9A941D7FFFA822A8FC2370A72526" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA822A8FC2370A72526" box="[498,746,1019,1043]" name="between Hiji and Iji" pageId="9" pageNumber="766">between Hiji and Iji</location>
|
||
, coll.
|
||
<collectionCode id="F8F2548A1D7FFFA82062FC2371022526" box="[824,847,1019,1043]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="766">T</collectionCode>
|
||
.
|
||
<collectorName id="3316A9991D7FFFA8203AFC2371FA2526" box="[864,951,1019,1043]" pageId="9" pageNumber="766">Naruse</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA8209DFC2376142526" box="[967,1113,1019,1043]" pageId="9" pageNumber="766" value="2001-07-05">
|
||
<collectingDate id="FA1913671D7FFFA8209DFC2376142526" box="[967,1113,1019,1043]" pageId="9" pageNumber="766" value="2001-07-05">5 July 2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82730FC23735D2566" collectingDate="2000-07-08" collectionCode="NSMT" collectorName="H. Kawauchi" location="Kunigami Village" municipality="Ohkuni Forest" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one female, CL 34.0 mm,
|
||
<collectionCode id="F8F2548A1D7FFFA82227FBC3739B2506" box="[381,470,1051,1075]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="9" pageNumber="766">NSMT</collectionCode>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA822B8FBC370C12506" box="[482,652,1051,1075]" pageId="9" pageNumber="766">Ohkuni Forest</collectingMunicipality>
|
||
Road,
|
||
<location id="9B3C9A941D7FFFA821BAFBC371E32506" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821BAFBC371E32506" box="[736,942,1051,1075]" municipality="Ohkuni Forest" name="Kunigami Village" pageId="9" pageNumber="766">Kunigami Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA820A8FBC376DD2506" box="[1010,1168,1051,1075]" pageId="9" pageNumber="766">H. Kawauchi</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82327FBE3735D2566" box="[125,272,1083,1107]" pageId="9" pageNumber="766" value="2000-07-08">
|
||
<collectingDate id="FA1913671D7FFFA82327FBE3735D2566" box="[125,272,1083,1107]" pageId="9" pageNumber="766" value="2000-07-08">8 July 2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="2E8BC6121D7FFFA82245FBE370182546" accessionNumber="C20007" collectingDate="1972-06-25" collectionCode="ZLKU" collectorName="H. Minei" location="Others" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
<location id="9B3C9A941D7FFFA82245FBE3733F2566" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA82245FBE3733F2566" box="[287,370,1083,1107]" name="Others" pageId="9" pageNumber="766">Others</location>
|
||
: four males, CL 30.0–
|
||
<quantity id="591B61AA1D7FFFA821D3FBE370BD2566" box="[649,752,1083,1107]" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="9" pageNumber="766" unit="mm" value="37.1">37.1 mm</quantity>
|
||
, three females, CL
|
||
<quantity id="591B61AA1D7FFFA820BFFBE376C22566" box="[997,1167,1083,1107]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" metricValueMax="4.15" metricValueMin="3.21" pageId="9" pageNumber="766" unit="mm" value="36.8" valueMax="41.5" valueMin="32.1">32.1–41.5 mm</quantity>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA82327FB8372802546" box="[125,205,1115,1139]" pageId="9" pageNumber="766">ZLKU</collectionCode>
|
||
C20007, data same as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA8213EFB8372F62587" collectingDate="1962-06-26" collectionCode="ZLKU" collectorName="H. Minei" location="Tamagusuku Village" municipality="Mt Yanbaru-Yama" pageId="9" pageNumber="766" specimenCode="ZLKU 10987" specimenCount="1" typeStatus="paratype">
|
||
two males, CL
|
||
<quantity id="591B61AA1D7FFFA82045FB8371872546" box="[799,970,1115,1139]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.06" metricValueMin="3.84" pageId="9" pageNumber="766" unit="mm" value="39.5" valueMax="40.6" valueMin="38.4">38.4–40.6 mm</quantity>
|
||
, one female, CL 44.0 mm,
|
||
<specimenCode id="CE4564341D7FFFA823A8FBA273DB25A7" box="[242,406,1146,1170]" collectionCode="ZLKU" pageId="9" pageNumber="766">ZLKU 10987</specimenCode>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA822F8FBA270CC25A7" box="[418,641,1146,1170]" pageId="9" pageNumber="766">Mt Yanbaru-Yama</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA821D7FBA2713225A7" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821D7FBA2713225A7" box="[653,895,1146,1170]" municipality="Mt Yanbaru-Yama" name="Tamagusuku Village" pageId="9" pageNumber="766">Tamagusuku Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA82098FBA2766225A7" box="[962,1071,1146,1170]" pageId="9" pageNumber="766">H. Minei</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82761FBA272F62587" pageId="9" pageNumber="766" value="1962-06-26">
|
||
<collectingDate id="FA1913671D7FFFA82761FBA272F62587" pageId="9" pageNumber="766" value="1962-06-26">26 June 1962</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82391FB42761125E7" collectingDate="1961-08-31" collectingDateMax="1962-06-26" collectingDateMin="1961-08-31" collectionCode="ZLKU" collectorName="H. Minei" location="Tamagusuku Village" municipality="Mt Yanbaru-Yama" pageId="9" pageNumber="766" specimenCode="ZLKU 11020" specimenCount="1" typeStatus="paratype">
|
||
one female, CL
|
||
<quantity id="591B61AA1D7FFFA822CAFB4273BA2587" box="[400,503,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="9" pageNumber="766" unit="mm" value="35.8">35.8 mm</quantity>
|
||
(
|
||
<date id="EA5DEA8F1D7FFFA82151FB42709C2587" box="[523,721,1178,1202]" pageId="9" pageNumber="766" value="1961-08-31">
|
||
<collectingDate id="FA1913671D7FFFA82151FB42709C2587" box="[523,721,1178,1202]" pageId="9" pageNumber="766" value="1961-08-31">31 August 1961</collectingDate>
|
||
</date>
|
||
), two females, CL
|
||
<quantity id="591B61AA1D7FFFA820E0FB42762C2587" box="[954,1121,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="4.234999999999999" metricValueMax="4.38" metricValueMin="4.09" pageId="9" pageNumber="766" unit="mm" value="42.349999999999994" valueMax="43.8" valueMin="40.9">40.9–43.8 mm</quantity>
|
||
(
|
||
<date id="EA5DEA8F1D7FFFA8272FFB4272B725E7" pageId="9" pageNumber="766" value="1962-06-26">
|
||
<collectingDate id="FA1913671D7FFFA8272FFB4272B725E7" pageId="9" pageNumber="766" value="1962-06-26">26 June 1962</collectingDate>
|
||
</date>
|
||
),
|
||
<specimenCode id="CE4564341D7FFFA8224BFB6273FB25E7" box="[273,438,1210,1234]" collectionCode="ZLKU" pageId="9" pageNumber="766">ZLKU 11020</specimenCode>
|
||
,
|
||
<collectingMunicipality id="7E3856351D7FFFA82299FB6270E825E7" box="[451,677,1210,1234]" pageId="9" pageNumber="766">Mt Yanbaru-Yama</collectingMunicipality>
|
||
,
|
||
<location id="9B3C9A941D7FFFA821E8FB6271EB25E7" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821E8FB6271EB25E7" box="[690,934,1210,1234]" municipality="Mt Yanbaru-Yama" name="Tamagusuku Village" pageId="9" pageNumber="766">Tamagusuku Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA820B7FB62761125E7" box="[1005,1116,1210,1234]" pageId="9" pageNumber="766">H. Minei</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82730FB6272F62427" accessionNumber="C20004" collectingDate="1972-06-25" collectionCode="ZLKU" collectorName="Fusato & H. Minei" location="Tamagusuku Village" pageId="9" pageNumber="766" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA823A8FB02731125C7" box="[242,348,1242,1266]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="9" pageNumber="766" unit="mm" value="35.2">35.2 mm</quantity>
|
||
,
|
||
<collectionCode id="F8F2548A1D7FFFA82232FB0273FA25C7" box="[360,439,1242,1266]" pageId="9" pageNumber="766">ZLKU</collectionCode>
|
||
C20004,
|
||
<collectorName id="3316A9991D7FFFA82176FB02703225C7" box="[556,639,1242,1266]" pageId="9" pageNumber="766">Fusato</collectorName>
|
||
,
|
||
<location id="9B3C9A941D7FFFA821D1FB02713025C7" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821D1FB02713025C7" box="[651,893,1242,1266]" name="Tamagusuku Village" pageId="9" pageNumber="766">Tamagusuku Village</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA8209BFB02766225C7" box="[961,1071,1242,1266]" pageId="9" pageNumber="766">H. Minei</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82761FB0272F62427" pageId="9" pageNumber="766" value="1972-06-25">
|
||
<collectingDate id="FA1913671D7FFFA82761FB0272F62427" pageId="9" pageNumber="766" value="1972-06-25">25 June 1972</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82392FB2272F62407" collectingDate="1956-08-05" collectionCode="ZLKU" collectorName="Kazuo Koba" location="Mt Meiji-Yama" pageId="9" pageNumber="766" specimenCode="ZLKU 9666" specimenCount="1" typeStatus="paratype">
|
||
one male, CL
|
||
<quantity id="591B61AA1D7FFFA8222BFB2273972427" box="[369,474,1274,1298]" metricMagnitude="-2" metricUnit="m" metricValue="4.58" pageId="9" pageNumber="766" unit="mm" value="45.8">45.8 mm</quantity>
|
||
,
|
||
<specimenCode id="CE4564341D7FFFA822BDFB2270312427" box="[487,636,1274,1298]" collectionCode="ZLKU" pageId="9" pageNumber="766">ZLKU 9666</specimenCode>
|
||
,
|
||
<location id="9B3C9A941D7FFFA821D3FB22710E2427" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821D3FB22710E2427" box="[649,835,1274,1298]" name="Mt Meiji-Yama" pageId="9" pageNumber="766">Mt Meiji-Yama</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA820D2FB2276522427" box="[904,1055,1274,1298]" pageId="9" pageNumber="766">Kazuo Koba</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82771FB2272F62407" pageId="9" pageNumber="766" value="1956-08-05">
|
||
<collectingDate id="FA1913671D7FFFA82771FB2272F62407" pageId="9" pageNumber="766" value="1956-08-05">5 August 1956</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="2E8BC6121D7FFFA82390FAC272F12467" collectingDate="1956-08-05" collectionCode="ZLKU" collectorName="K. Koba" location="Mt Meiji-Yama" pageId="9" pageNumber="766" specimenCode="ZLKU 9667" specimenCount="1" typeStatus="paratype">
|
||
one female, CL
|
||
<quantity id="591B61AA1D7FFFA822D7FAC273BA2407" box="[397,503,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="9" pageNumber="766" unit="mm" value="39.4">39.4 mm</quantity>
|
||
,
|
||
<specimenCode id="CE4564341D7FFFA8215FFAC270D02407" box="[517,669,1306,1330]" collectionCode="ZLKU" pageId="9" pageNumber="766">ZLKU 9667</specimenCode>
|
||
,
|
||
<location id="9B3C9A941D7FFFA821F6FAC2712A2407" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA821F6FAC2712A2407" box="[684,871,1306,1330]" name="Mt Meiji-Yama" pageId="9" pageNumber="766">Mt Meiji-Yama</location>
|
||
, coll.
|
||
<collectorName id="3316A9991D7FFFA820EBFAC276562407" box="[945,1051,1306,1330]" pageId="9" pageNumber="766">K. Koba</collectorName>
|
||
,
|
||
<date id="EA5DEA8F1D7FFFA82773FAC272F12467" pageId="9" pageNumber="766" value="1956-08-05">
|
||
<collectingDate id="FA1913671D7FFFA82773FAC272F12467" pageId="9" pageNumber="766" value="1956-08-05">5 August 1956</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="2E8BC6121D7FFFA82390FAE273B52447" collectionCode="RUMF" collectorName="M. Moriguchi. All & Island" location="Chinen Village" municipality="Fossil" pageId="9" pageNumber="766" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
|
||
<collectingMunicipality id="7E3856351D7FFFA82390FAE273402467" box="[202,269,1338,1362]" pageId="9" pageNumber="766">Fossil</collectingMunicipality>
|
||
material: one palm,
|
||
<collectionCode id="F8F2548A1D7FFFA82153FAE2702F2467" box="[521,610,1338,1362]" pageId="9" pageNumber="766">RUMF</collectionCode>
|
||
,
|
||
<location id="9B3C9A941D7FFFA8212AFAE271692467" LSID="urn:lsid:plazi:treatment:164A7D591D7FFFAD22F8FF51708C27A0:9B3C9A941D7FFFA8212AFAE271692467" box="[624,804,1338,1362]" municipality="Fossil" name="Chinen Village" pageId="9" pageNumber="766" stateProvince="Okinawa">Chinen Village</location>
|
||
, 2001, coll.
|
||
<collectorName id="3316A9991D7FFFA820E0FAE276D92467" box="[954,1172,1338,1362]" pageId="9" pageNumber="766">M. Moriguchi. All</collectorName>
|
||
specimens from
|
||
<collectingRegion id="5C2702AD1D7FFFA82264FA8273E82447" box="[318,421,1370,1394]" country="Japan" name="Okinawa" pageId="9" pageNumber="766">Okinawa</collectingRegion>
|
||
<collectorName id="3316A9991D7FFFA822F7FA8273B52447" box="[429,504,1370,1394]" pageId="9" pageNumber="766">Island</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7FFFA82327FA72732124F7" blockId="9.[125,364,1450,1474]" box="[125,364,1450,1474]" pageId="9" pageNumber="766">
|
||
<emphasis id="AC97105D1D7FFFA82327FA72732124F7" box="[125,364,1450,1474]" italics="true" pageId="9" pageNumber="766">Comparatiυe material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7FFFA82327FA0371A924C6" blockId="9.[125,996,1499,1523]" box="[125,996,1499,1523]" pageId="9" pageNumber="766">
|
||
<taxonomicName id="59E3B7CC1D7FFFA82327FA0373D824C6" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[125,405,1499,1523]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="766" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7FFFA82327FA0373D824C6" box="[125,405,1499,1523]" italics="true" pageId="9" pageNumber="766">Geothelphusa tenuimanus</emphasis>
|
||
</taxonomicName>
|
||
: all specimens examined in
|
||
<bibRefCitation id="FA72B1BE1D7FFFA821BBFA0371AD24C6" author="Naruse T & Karasawa H & Shokita S & Tanaka T & Moriguchi M" box="[737,992,1499,1523]" pageId="9" pageNumber="766" pagination="1211 - 1218" refId="ref8991" refString="Naruse T, Karasawa H, Shokita S, Tanaka T, Moriguchi M. 2004 a. A first fossil record of the terrestrial crab, Geothelphusa tenuimanus (Miyake and Minei, 1965) (Decapoda, Brachyura, Potamidae) from Okinawa Island, Central Ryukyus, Japan. Crustaceana 76: 1211 - 1218." type="journal article" year="2004">Naruse et al. (2004a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7FFFA82327F9F4731F2771" blockId="9.[125,338,1580,1604]" box="[125,338,1580,1604]" pageId="9" pageNumber="766">
|
||
<emphasis id="AC97105D1D7FFFA82327F9F4731F2771" box="[125,338,1580,1604]" italics="true" pageId="9" pageNumber="766">Description of adult</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7FFFAA2327F98572F425EA" blockId="9.[125,1172,1629,1685]" lastBlockId="11.[125,1173,931,1670]" lastPageId="11" lastPageNumber="768" pageId="9" pageNumber="766">
|
||
Carapace (
|
||
<figureCitation id="06D8D0CA1D7FFFA823A3F98573C82740" box="[249,389,1629,1653]" captionStart="Figure 4" captionStartId="10.[138,198,1513,1532]" captionTargetBox="[9,1186,133,1471]" captionTargetId="figure-1@10.[138,1187,133,1477]" captionTargetPageId="10" captionText="Figure 4. Geothelphusa grandioυata new species. (a) Dorsal view; (b) dorsal view of carapace; (c) ventral side of G1; (d) dorsal side of G1. (a) Paratype, ZRC 2006.0076, CL 44.8 mm; (b–d) holotype, ZLKU m. 1094, CL 40.5 mm. Scale bars: 10 mm (b); 5mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.4581770" httpUri="https://zenodo.org/record/4581770/files/figure.png" pageId="9" pageNumber="766">Figure 4a, b</figureCitation>
|
||
) oval, strongly convex longitudinally; surface including anterolateral margin smooth, cervical groove very shallow, H-shaped gastric groove distinct, postorbital and epigastric cristae absent, postorbital and epigastric regions raised posteriorly gradually, small pit present on posterior part of external orbital angle. Front directed downwards, frontal margin cristate, not granulated; orbital margins cristate, not granulated or granulated on outer third, supraorbital margin L-shaped in dorsal view; epibranchial tooth low, demarcated by interrupted crista, low crista confluent with epibranchial tooth but separate from external orbital angle; anterolateral margin barely cristate, not granulated. Suborbital, subhepatic, and pterygostomial regions (
|
||
<figureCitation id="06D8D0CA1D7DFFAA2740FBBD76C62548" box="[1050,1163,1125,1149]" captionStart="Figure 8" captionStartId="19.[125,185,1284,1303]" captionTargetBox="[9,1174,135,1281]" captionTargetId="figure-87@19.[125,1174,133,1248]" captionTargetPageId="19" captionText="Figure 8. Frontal view of Geothelphusa leυicerυix (Rathbun, 1898) and G. grandioυata new species. (a) Geothelphusa leυicerυix, ZRC 2006.0074, male, CL 35.3 mm; (b) G. grandioυata, paratype, ZRC 2006.0076, male, CL 44.8 mm." figureDoi="http://doi.org/10.5281/zenodo.4581781" httpUri="https://zenodo.org/record/4581781/files/figure.png" pageId="11" pageNumber="768">Figure 8b</figureCitation>
|
||
) smooth, no granulation visible on vertical groove and epimeral sulcus. Eye with relatively small cornea, subdistal width of cornea as wide as base of peduncle in frontal view.
|
||
</paragraph>
|
||
<caption id="CA9C9CC71D7CFFAB23D0FA31739A2704" ID-DOI="http://doi.org/10.5281/zenodo.4581770" ID-Zenodo-Dep="4581770" httpUri="https://zenodo.org/record/4581770/files/figure.png" pageId="10" pageNumber="767" startId="10.[138,198,1513,1532]" targetBox="[9,1186,133,1471]" targetPageId="10">
|
||
<paragraph id="9E5CCC4F1D7CFFAB23D0FA31739A2704" blockId="10.[138,1186,1513,1585]" pageId="10" pageNumber="767">
|
||
Figure 4.
|
||
<emphasis id="AC97105D1D7CFFAB23B2FA31738424C9" box="[232,457,1513,1532]" italics="true" pageId="10" pageNumber="767">
|
||
<taxonomicName id="59E3B7CC1D7CFFAB23B2FA31731124C9" authorityName="Stimpson" authorityYear="1858" box="[232,348,1513,1532]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="767" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
grandioυata
|
||
</emphasis>
|
||
new species. (a) Dorsal view; (b) dorsal view of carapace; (c) ventral side of G1; (d) dorsal side of G1. (a) Paratype, ZRC 2006.0076, CL 44.8 mm; (b–d) holotype, ZLKU m. 1094, CL 40.5 mm. Scale bars: 10 mm (b); 5mm (c, d).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CA9C9CC71D7DFFAA2327FD2B706E220E" ID-DOI="http://doi.org/10.5281/zenodo.4581774" ID-Zenodo-Dep="4581774" httpUri="https://zenodo.org/record/4581774/files/figure.png" pageId="11" pageNumber="768" startId="11.[125,185,755,774]" targetBox="[125,1172,133,719]" targetPageId="11">
|
||
<paragraph id="9E5CCC4F1D7DFFAA2327FD2B706E220E" blockId="11.[125,1173,755,827]" pageId="11" pageNumber="768">
|
||
Figure 5. Fossils of
|
||
<emphasis id="AC97105D1D7DFFAA226DFD2B70542233" box="[311,537,755,774]" italics="true" pageId="11" pageNumber="768">
|
||
<taxonomicName id="59E3B7CC1D7DFFAA226DFD2B73E62233" authorityName="Stimpson" authorityYear="1858" box="[311,427,755,774]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="768" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
grandioυata
|
||
</emphasis>
|
||
new species and
|
||
<taxonomicName id="59E3B7CC1D7DFFAA21E2FD2B76782233" authority="(Miyake and Minei, 1965)" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[696,1077,755,774]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="768" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7DFFAA21E2FD2B71742233" box="[696,825,755,774]" italics="true" pageId="11" pageNumber="768">G. tenuimanus</emphasis>
|
||
(
|
||
<bibRefCitation id="FA72B1BE1D7DFFAA201EFD2B76632233" author="Miyake S & Minei H" box="[836,1070,755,774]" pageId="11" pageNumber="768" pagination="377 - 382" refId="ref8786" refString="Miyake S, Minei H. 1965. A new fresh-water crab, Potamon (Geothelphusa) tenuimanus sp. nov. from Okinawajima, the Ryukyu Islands. Science Bulletin of the Faculty of Agriculture, Kyushu University 4: 377 - 382. Plate 21. (Jpn with Eng summary)." type="journal article" year="1965">Miyake and Minei, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. (a) Chela of
|
||
<emphasis id="AC97105D1D7DFFAA23CCFCD673562214" box="[150,283,782,801]" italics="true" pageId="11" pageNumber="768">G. grandioυata</emphasis>
|
||
(RUMF, palm length 28.6+ mm, width 23.7+ mm); (b) chela of
|
||
<taxonomicName id="59E3B7CC1D7DFFAA202EFCD671B82214" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[884,1013,782,801]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="768" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7DFFAA202EFCD671B82214" box="[884,1013,782,801]" italics="true" pageId="11" pageNumber="768">G. tenuimanus</emphasis>
|
||
</taxonomicName>
|
||
(RUMF-GF-11, palm length 18.6 mm, palm width 17.3 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9E5CCC4F1D7DFFAA23C2FB3F76132415" blockId="11.[125,1173,931,1670]" pageId="11" pageNumber="768">Chelae of large male asymmetrical in size and shape, major chela with wide, thick, and reticulated manus, movable finger long, curved, with large gape when fingers closed.</paragraph>
|
||
<paragraph id="9E5CCC4F1D7DFFAA23C2FAF1707A24F7" blockId="11.[125,1173,931,1670]" pageId="11" pageNumber="768">Ambulatory legs glabrous, propodi with elliptical cross-section, two longitudinal rows of spines on inner margins; dactyli with rectangular cross-section, each margin with row of spines, inner margins with short, stout, sharp spines, outer margins with short, stout, rounded spine, subdistal spine of outer dorsal margin of fourth ambulatory dactylus stout, rounded, smaller than terminal spine.</paragraph>
|
||
<paragraph id="9E5CCC4F1D7DFFAA23C2FA13715E2731" blockId="11.[125,1173,931,1670]" pageId="11" pageNumber="768">Male abdominal segments and telson triangular, telson slightly bell-shaped, reaching imaginary line joining proximal fifth of chelipedal coxae.</paragraph>
|
||
<paragraph id="9E5CCC4F1D7DFFAA23C2F9D5730727B3" blockId="11.[125,1173,931,1670]" pageId="11" pageNumber="768">
|
||
G1 (
|
||
<figureCitation id="06D8D0CA1D7DFFAA2394F9D573132710" box="[206,350,1549,1573]" captionStart="Figure 4" captionStartId="10.[138,198,1513,1532]" captionTargetBox="[9,1186,133,1471]" captionTargetId="figure-1@10.[138,1187,133,1477]" captionTargetPageId="10" captionText="Figure 4. Geothelphusa grandioυata new species. (a) Dorsal view; (b) dorsal view of carapace; (c) ventral side of G1; (d) dorsal side of G1. (a) Paratype, ZRC 2006.0076, CL 44.8 mm; (b–d) holotype, ZLKU m. 1094, CL 40.5 mm. Scale bars: 10 mm (b); 5mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.4581770" httpUri="https://zenodo.org/record/4581770/files/figure.png" pageId="11" pageNumber="768">Figure 4c, d</figureCitation>
|
||
) stout; subterminal segment curving outwards, distal part wider than proximal part, with knob on outer proximal angle; synovial membrane wide; terminal segment directed outwards, subdistal margin narrower than distal margin of subterminal segment, uneven.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FF5072BF2195" blockId="12.[138,242,136,160]" box="[138,242,136,160]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FF5072BF2195" box="[138,242,136,160]" italics="true" pageId="12" pageNumber="769">Variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FF6170F62045" blockId="12.[138,1185,185,368]" pageId="12" pageNumber="769">
|
||
In one female specimen (RUMF-ZC-143, CL
|
||
<quantity id="591B61AA1D7AFFAD21F5FF61715721E4" box="[687,794,185,209]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="12" pageNumber="769" unit="mm" value="27.1">27.1 mm</quantity>
|
||
), the carapace is relatively flatter; with frontal margin convex medially and along both outer angles; the external orbital angle and epibranchial tooth are connected by a crista; the anterolateral margin has very low granules; the anterolateral region possesses very shallow grooves, appearing rugose; the pit behind the external orbital angle is relatively shallower; and there is a row of spines present on the outer margin of the ambulatory propodi.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FE71729F20F4" blockId="12.[138,210,425,449]" box="[138,210,425,449]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FE71729F20F4" box="[138,210,425,449]" italics="true" pageId="12" pageNumber="769">Colour</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FE02735B2304" blockId="12.[138,1185,474,561]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FE0273E220C7" box="[138,431,474,498]" italics="true" pageId="12" pageNumber="769">
|
||
<taxonomicName id="59E3B7CC1D7AFFAD23D0FE02735620C7" authorityName="Stimpson" authorityYear="1858" box="[138,283,474,498]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
grandioυata
|
||
</emphasis>
|
||
is maroon to olive in colour in life. Some individuals are coloured a bluish slate-grey on the cardiac region, sides of the carapace, and third maxillipeds.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FDB2729223B7" blockId="12.[138,223,618,642]" box="[138,223,618,642]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FDB2729223B7" box="[138,223,618,642]" italics="true" pageId="12" pageNumber="769">Habitat</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FD4376052267" blockId="12.[138,1185,667,850]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FD4373E92386" box="[138,420,667,691]" italics="true" pageId="12" pageNumber="769">
|
||
<taxonomicName id="59E3B7CC1D7AFFAD23D0FD4373572386" authorityName="Stimpson" authorityYear="1858" box="[138,282,667,691]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
grandioυata
|
||
</emphasis>
|
||
occurs near middle to upper reaches of the banks or along small swamps in submontane areas. It also digs deep burrows adjacent to areas with water seeping out. On the basis of our collections, large individuals seem to prefer burrows which have at least some water in the bottom, but medium-sized individuals (e.g. a female, RUMF-ZC- 143, CL
|
||
<quantity id="591B61AA1D7AFFAD23AEFCC273112207" box="[244,348,794,818]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="12" pageNumber="769" unit="mm" value="27.1">27.1 mm</quantity>
|
||
; a male, RUMF-ZC-141, CL
|
||
<quantity id="591B61AA1D7AFFAD21E6FCC271692207" box="[700,804,794,818]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="12" pageNumber="769" unit="mm" value="29.7">29.7 mm</quantity>
|
||
) have been obtained from totally dried burrows (
|
||
<collectionCode id="F8F2548A1D7AFFAD2265FCE2731B2267" box="[319,342,826,850]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="769">T</collectionCode>
|
||
. Naruse, unpublished data;
|
||
<collectionCode id="F8F2548A1D7AFFAD21C5FCE270FB2267" box="[671,694,826,850]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="769">T</collectionCode>
|
||
. Nagai, personal communication).
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FC5373462296" blockId="12.[138,267,907,931]" box="[138,267,907,931]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FC5373462296" box="[138,267,907,931]" italics="true" pageId="12" pageNumber="769">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FC6370A122E6" blockId="12.[138,748,955,979]" box="[138,748,955,979]" pageId="12" pageNumber="769">
|
||
<collectingRegion id="5C2702AD1D7AFFAD23D0FC6372BC22E6" box="[138,241,955,979]" country="Japan" name="Okinawa" pageId="12" pageNumber="769">Okinawa</collectingRegion>
|
||
Island, the Central Ryukyu Islands,
|
||
<collectingCountry id="E6F48CDF1D7AFFAD21F8FC6370AA22E6" box="[674,743,955,979]" name="Japan" pageId="12" pageNumber="769">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FBD472B62511" blockId="12.[138,251,1036,1060]" box="[138,251,1036,1060]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FBD472B62511" box="[138,251,1036,1060]" italics="true" pageId="12" pageNumber="769">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FBE471A12541" blockId="12.[138,1185,1084,1140]" pageId="12" pageNumber="769">
|
||
The species name is derived from the Latin ‘‘
|
||
<emphasis id="AC97105D1D7AFFAD21F2FBE470BB2561" box="[680,758,1084,1108]" italics="true" pageId="12" pageNumber="769">grandis</emphasis>
|
||
’’ (for large) and ‘‘
|
||
<emphasis id="AC97105D1D7AFFAD208EFBE576572561" box="[980,1050,1085,1108]" italics="true" pageId="12" pageNumber="769">oυatus</emphasis>
|
||
’’ (for ovalshaped), suggesting the large body size and the strongly swollen carapace.
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FB7572A425F0" blockId="12.[138,233,1197,1221]" box="[138,233,1197,1221]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FB7572A425F0" box="[138,233,1197,1221]" italics="true" pageId="12" pageNumber="769">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23D0FB0671F02440" blockId="12.[138,1185,1246,1685]" pageId="12" pageNumber="769">
|
||
<emphasis id="AC97105D1D7AFFAD23D0FB0673E825C3" box="[138,421,1246,1270]" italics="true" pageId="12" pageNumber="769">
|
||
<taxonomicName id="59E3B7CC1D7AFFAD23D0FB06735625C3" authorityName="Stimpson" authorityYear="1858" box="[138,283,1246,1270]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
|
||
grandioυata
|
||
</emphasis>
|
||
has been regarded as
|
||
<emphasis id="AC97105D1D7AFFAD21FAFB06716425C3" box="[672,809,1246,1270]" italics="true" pageId="12" pageNumber="769">G. leυicerυix</emphasis>
|
||
since
|
||
<bibRefCitation id="FA72B1BE1D7AFFAD202AFB0676D025C3" author="Miyake S & Minei H" box="[880,1181,1246,1270]" pageId="12" pageNumber="769" pagination="377 - 382" refId="ref8786" refString="Miyake S, Minei H. 1965. A new fresh-water crab, Potamon (Geothelphusa) tenuimanus sp. nov. from Okinawajima, the Ryukyu Islands. Science Bulletin of the Faculty of Agriculture, Kyushu University 4: 377 - 382. Plate 21. (Jpn with Eng summary)." type="journal article" year="1965">Miyake and Minei (1965)</bibRefCitation>
|
||
. However,
|
||
<emphasis id="AC97105D1D7AFFAD2252FB2673FB2423" box="[264,438,1278,1302]" italics="true" pageId="12" pageNumber="769">G. grandioυata</emphasis>
|
||
can clearly be differentiated from
|
||
<emphasis id="AC97105D1D7AFFAD2031FB2671B62423" box="[875,1019,1278,1302]" italics="true" pageId="12" pageNumber="769">G. leυicerυix</emphasis>
|
||
s. str. by its proportionately wider G1 (versus more slender in
|
||
<emphasis id="AC97105D1D7AFFAD2050FAC571D12400" box="[778,924,1309,1333]" italics="true" pageId="12" pageNumber="769">G. leυicerυix</emphasis>
|
||
), the wider synovial membrane of the G1 (versus more slender), and the shape of the inner margin between terminal and subterminal segment of the G1 (uneven versus smooth).
|
||
</paragraph>
|
||
<paragraph id="9E5CCC4F1D7AFFAD23FFFAA6708C27A0" blockId="12.[138,1185,1246,1685]" pageId="12" pageNumber="769">
|
||
A fossil of a large brachyuran manus (
|
||
<figureCitation id="06D8D0CA1D7AFFAD213CFAA6709A24A3" box="[614,727,1406,1430]" captionStart="Figure 5" captionStartId="11.[125,185,755,774]" captionTargetBox="[125,1172,133,719]" captionTargetId="figure-243@11.[125,1172,133,719]" captionTargetPageId="11" captionText="Figure 5. Fossils of Geothelphusa grandioυata new species and G. tenuimanus (Miyake and Minei, 1965). (a) Chela of G. grandioυata (RUMF, palm length 28.6+ mm, width 23.7+ mm); (b) chela of G. tenuimanus (RUMF-GF-11, palm length 18.6 mm, palm width 17.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.4581774" httpUri="https://zenodo.org/record/4581774/files/figure.png" pageId="12" pageNumber="769">Figure 5a</figureCitation>
|
||
,
|
||
<collectionCode id="F8F2548A1D7AFFAD21BFFAA6717324A3" box="[741,830,1406,1430]" pageId="12" pageNumber="769">RUMF</collectionCode>
|
||
, palm length 28.6+ mm, palm width 23.7+ mm) was discovered in the limestone fissure sediments of the southern part of
|
||
<collectingRegion id="5C2702AD1D7AFFAD23D0FA6672BC24E3" box="[138,241,1470,1494]" country="Japan" name="Okinawa" pageId="12" pageNumber="769">Okinawa</collectingRegion>
|
||
Island, where fossils of
|
||
<taxonomicName id="59E3B7CC1D7AFFAD2153FA6670E124E3" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[521,684,1470,1494]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7AFFAD2153FA6670E124E3" box="[521,684,1470,1494]" italics="true" pageId="12" pageNumber="769">G. tenuimanus</emphasis>
|
||
</taxonomicName>
|
||
have been collected (
|
||
<bibRefCitation id="FA72B1BE1D7AFFAD20F2FA6676D924E3" author="Naruse T & Karasawa H & Shokita S & Tanaka T & Moriguchi M" box="[936,1172,1470,1494]" pageId="12" pageNumber="769" pagination="1211 - 1218" refId="ref8991" refString="Naruse T, Karasawa H, Shokita S, Tanaka T, Moriguchi M. 2004 a. A first fossil record of the terrestrial crab, Geothelphusa tenuimanus (Miyake and Minei, 1965) (Decapoda, Brachyura, Potamidae) from Okinawa Island, Central Ryukyus, Japan. Crustaceana 76: 1211 - 1218." type="journal article" year="2004">Naruse et al. 2004a</bibRefCitation>
|
||
). The discovered manus can be identified as that of
|
||
<emphasis id="AC97105D1D7AFFAD2057FA0671D024C3" box="[781,925,1502,1526]" italics="true" pageId="12" pageNumber="769">G. leυicerυix</emphasis>
|
||
species group by its large size [versus palm length,
|
||
<quantity id="591B61AA1D7AFFAD2168FA2570D42720" box="[562,665,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="12" pageNumber="769" unit="mm" value="18.6">18.6 mm</quantity>
|
||
(RUMF-GF-11); palm width,18.0 mm (
|
||
<collectionCode id="F8F2548A1D7AFFAD23C8F9C572912700" box="[146,220,1565,1589]" pageId="12" pageNumber="769">MFM</collectionCode>
|
||
142451) in
|
||
<taxonomicName id="59E3B7CC1D7AFFAD2230F9C570422700" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[362,527,1565,1589]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7AFFAD2230F9C570422700" box="[362,527,1565,1589]" italics="true" pageId="12" pageNumber="769">G. tenuimanus</emphasis>
|
||
</taxonomicName>
|
||
] (
|
||
<figureCitation id="06D8D0CA1D7AFFAD2176F9C570ED2700" box="[556,672,1565,1589]" captionStart="Figure 5" captionStartId="11.[125,185,755,774]" captionTargetBox="[125,1172,133,719]" captionTargetId="figure-243@11.[125,1172,133,719]" captionTargetPageId="11" captionText="Figure 5. Fossils of Geothelphusa grandioυata new species and G. tenuimanus (Miyake and Minei, 1965). (a) Chela of G. grandioυata (RUMF, palm length 28.6+ mm, width 23.7+ mm); (b) chela of G. tenuimanus (RUMF-GF-11, palm length 18.6 mm, palm width 17.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.4581774" httpUri="https://zenodo.org/record/4581774/files/figure.png" pageId="12" pageNumber="769">Figure 5b</figureCitation>
|
||
). Since
|
||
<emphasis id="AC97105D1D7AFFAD2059F9C571E32700" box="[771,942,1565,1589]" italics="true" pageId="12" pageNumber="769">G. grandioυata</emphasis>
|
||
is the only member of
|
||
<emphasis id="AC97105D1D7AFFAD23F5F9E573722760" box="[175,319,1597,1621]" italics="true" pageId="12" pageNumber="769">G. leυicerυix</emphasis>
|
||
species group known from
|
||
<collectingRegion id="5C2702AD1D7AFFAD21FBF9E571452760" box="[673,776,1597,1621]" country="Japan" name="Okinawa" pageId="12" pageNumber="769">Okinawa</collectingRegion>
|
||
Island, the fossil manus is most probably that of
|
||
<emphasis id="AC97105D1D7AFFAD2217F98573BB2740" box="[333,502,1629,1653]" italics="true" pageId="12" pageNumber="769">G. grandioυata</emphasis>
|
||
.
|
||
<bibRefCitation id="FA72B1BE1D7AFFAD215FF98570B32740" author="Naruse T & Karasawa H & Shokita S & Tanaka T & Moriguchi M" box="[517,766,1629,1653]" pageId="12" pageNumber="769" pagination="1211 - 1218" refId="ref8991" refString="Naruse T, Karasawa H, Shokita S, Tanaka T, Moriguchi M. 2004 a. A first fossil record of the terrestrial crab, Geothelphusa tenuimanus (Miyake and Minei, 1965) (Decapoda, Brachyura, Potamidae) from Okinawa Island, Central Ryukyus, Japan. Crustaceana 76: 1211 - 1218." type="journal article" year="2004">Naruse et al. (2004a)</bibRefCitation>
|
||
estimated that the geological age of the fossils of
|
||
<taxonomicName id="59E3B7CC1D7AFFAD227FF9A5738527A0" baseAuthorityName="Miyake and Minei" baseAuthorityYear="1965" box="[293,456,1661,1685]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="769" phylum="Arthropoda" rank="species" species="tenuimanus">
|
||
<emphasis id="AC97105D1D7AFFAD227FF9A5738527A0" box="[293,456,1661,1685]" italics="true" pageId="12" pageNumber="769">G. tenuimanus</emphasis>
|
||
</taxonomicName>
|
||
was late Pleistocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |