treatments-xml/data/03/F4/B0/03F4B022E3425B43FE5A647A81105B10.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="EDF7773655A997EAECD496FE4B440157">Three new species of Satsuma Adams, 1868 (Gastropoda, Camaenidae) from Taiwan</mods:title>
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<mods:namePart id="950EC3C20611756BACE82251AD994D71">Hwang, Chung-Chi</mods:namePart>
<mods:affiliation id="2DCF7A180361838800AA8270BC06B3CF">Department of Life Sciences, National University of Kaohsiung, No. 700, Kaohsiung University Road, Nan-Tzu District, Kaohsiung 81148 (Taiwan)</mods:affiliation>
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<mods:affiliation id="6E27060D284B791EFFC786017F784613">Department of Earth and Life Sciences, University of Taipei, No. 1, Ai-Guo West Road, Taipei 10048 (Taiwan)</mods:affiliation>
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<taxonomicName id="4C5D7AB7E3425B42FCE3647A807D5A65" ID-CoL="7NX8W" authority="Adams, 1868" authorityName="Adams" authorityYear="1868" box="[788,1036,1087,1114]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="genus">
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<emphasis id="B929DD26E3425B42FCE3647A87025A64" box="[788,883,1088,1114]" italics="true" pageId="14" pageNumber="73">SATSUMA</emphasis>
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, 1868
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<keyLead id="30ACA201E3425B42FF25644F81125AB1" box="[210,1379,1140,1165]" pageId="14" pageNumber="73">1 Shell high conic; shell height&gt; shell width .................................................................................................. 2</keyLead>
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<keyLead id="30ACA201E3425B42FF2564A881125A95" box="[210,1379,1169,1193]" pageId="14" pageNumber="73">— Shell depressed conic; shell height &lt;shell width .......................................................................................... 3</keyLead>
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<keyStep id="30A91991E3425B42FF2564FF81125B2A" pageId="14" pageNumber="73">
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2 Shell white or yellowish golden; with/without polymorphic bands ................................................................. ................................................................................................
<taxonomicName id="4C5D7AB7E3425B42FC7A64D887975AC5" baseAuthorityName="Adams" baseAuthorityYear="1870" box="[909,998,1249,1273]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="albida">
<emphasis id="B929DD26E3425B42FC7A64D887975AC5" box="[909,998,1249,1273]" italics="true" pageId="14" pageNumber="73">S. albida</emphasis>
</taxonomicName>
species complex (see
<bibRefCitation id="EFCC7CC5E3425B42FB3464D8812C5AC5" author="WU S. - P. &amp; HWANG C. - C. &amp; LIN Y. - S." box="[1219,1373,1249,1273]" pageId="14" pageNumber="73" pagination="437 - 493" refId="ref9748" refString="WU S. - P., HWANG C. - C. &amp; LIN Y. - S. 2008. - Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan. Zoological Journal of the Linnean Society 154 (3): 437 - 493. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00415. x" type="journal article" year="2008">
Wu
<emphasis id="B929DD26E3425B42FB1964D881515AC5" box="[1262,1312,1249,1273]" italics="true" pageId="14" pageNumber="73">et al.</emphasis>
2008
</bibRefCitation>
)
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<keyLead id="30ACA201E3425B42FF2564C581125B2A" box="[210,1379,1278,1302]" pageId="14" pageNumber="73">
— Shell brown; with a peripheral band ...............................................................
<taxonomicName id="4C5D7AB7E3425B42FBDE64C481125B2A" authority="(Pfeiffer, 1866)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1866" box="[1065,1379,1278,1302]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="sphaeroconus">
<emphasis id="B929DD26E3425B42FBDE64C480B05B2A" box="[1065,1217,1278,1302]" italics="true" pageId="14" pageNumber="73">S. sphaeroconus</emphasis>
(Pfeiffer, 1866)
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3 Shell colour yellow ....................................................................................................
<taxonomicName id="4C5D7AB7E3425B42FB86650881125B75" authority="(Pfeiffer, 1866)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1866" box="[1137,1379,1329,1353]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="bacca">
<emphasis id="B929DD26E3425B42FB86650880B15B75" box="[1137,1216,1329,1353]" italics="true" pageId="14" pageNumber="73">S. bacca</emphasis>
(Pfeiffer, 1866)
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<paragraph id="8BE20134E3425B42FF25657581125B5A" blockId="14.[210,1379,1329,1462]" box="[210,1379,1358,1382]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF25657581125B5A" box="[210,1379,1358,1382]" pageId="14" pageNumber="73">— Shell colour light brown to dark brown ....................................................................................................... 4</keyLead>
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<keyLead id="30ACA201E3425B42FF2565B881125BA5" box="[210,1379,1409,1434]" pageId="14" pageNumber="73">4 Umbilicus totally covered by reflexed columellar lip .................................................................................... 5</keyLead>
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<keyLead id="30ACA201E3425B42FF0865A481125B8A" box="[255,1379,1438,1462]" pageId="14" pageNumber="73">Umbilicus partly open to completely open .................................................................................................. 6</keyLead>
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5 Shell large-sized, shell width&gt;
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...................................................................
<taxonomicName id="4C5D7AB7E3425B42FBAD65E881155BD6" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1114,1380,1490,1514]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="arisana">
<emphasis id="B929DD26E3425B42FBAD65E880CD5BD5" box="[1114,1212,1490,1513]" italics="true" pageId="14" pageNumber="73">S. arisana</emphasis>
(
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)
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</keyLead>
</paragraph>
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<keyLead id="30ACA201E3425B42FF0865D48115583B" box="[255,1380,1518,1543]" pageId="14" pageNumber="73">
Shell medium-sized, shell width &lt;
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...................................................................
<taxonomicName id="4C5D7AB7E3425B42FB8D65D58115583B" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1146,1380,1518,1543]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="tani">
<emphasis id="B929DD26E3425B42FB8D65D580CD583A" box="[1146,1212,1519,1542]" italics="true" pageId="14" pageNumber="73">S. tani</emphasis>
(
<bibRefCitation id="EFCC7CC5E3425B42FB3D65D4812D583B" author="KURODA T." box="[1226,1372,1518,1543]" pageId="14" pageNumber="73" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
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</paragraph>
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<keyLead id="30ACA201E3425B42FF25661881125805" box="[210,1379,1570,1594]" pageId="14" pageNumber="73">
6 Shell width&gt;
<quantity id="4CA5ACD1E3425B42FE78661885AB5805" box="[399,474,1570,1594]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="73" unit="mm" value="35.0">35 mm</quantity>
................................................................................................................................... 7
</keyLead>
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<keyLead id="30ACA201E3425B42FF0866058115586B" box="[255,1380,1598,1623]" pageId="14" pageNumber="73">
Shell width &lt;
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................................................................................................................................. 10
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<keyStep id="30A91991E3425B42FF2566488112589B" pageId="14" pageNumber="73">
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<keyLead id="30ACA201E3425B42FF256648811258B6" box="[210,1379,1650,1674]" pageId="14" pageNumber="73">
7 Upper lip clearly sinuous; fold on basal lip present ...............................................
<taxonomicName id="4C5D7AB7E3425B42FBBD6648811258B6" authority="(H. Adams, 1866)" baseAuthorityName="H. Adams" baseAuthorityYear="1866" box="[1098,1379,1650,1674]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="bairdi">
<emphasis id="B929DD26E3425B42FBBD664880D158B6" box="[1098,1184,1650,1674]" italics="true" pageId="14" pageNumber="73">S. bairdi</emphasis>
(H. Adams, 1866)
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</paragraph>
<paragraph id="8BE20134E3425B42FF0866B58112589B" blockId="14.[210,1380,1650,1783]" box="[255,1379,1679,1703]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF0866B58112589B" box="[255,1379,1679,1703]" pageId="14" pageNumber="73">Upper lip smoothly curved; fold on basal lip absent .................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3425B42FF2566F8811258CB" pageId="14" pageNumber="73">
<paragraph id="8BE20134E3425B42FF2566F8811558E6" blockId="14.[210,1380,1650,1783]" box="[210,1380,1730,1754]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF2566F8811558E6" box="[210,1380,1730,1754]" pageId="14" pageNumber="73">
8 Spire lower; shell height/shell width &lt;0.7 ............................................................
<taxonomicName id="4C5D7AB7E3425B42FBBE66F9811558E6" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1097,1380,1730,1754]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="takahasii">
<emphasis id="B929DD26E3425B42FBBE66F980CC58E6" box="[1097,1213,1730,1754]" italics="true" pageId="14" pageNumber="73">S. takahasii</emphasis>
(
<bibRefCitation id="EFCC7CC5E3425B42FB3D66F8812D58E6" author="KURODA T." box="[1226,1372,1730,1754]" pageId="14" pageNumber="73" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3425B42FF2566E5811258CB" blockId="14.[210,1380,1650,1783]" box="[210,1379,1759,1783]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF2566E5811258CB" box="[210,1379,1759,1783]" pageId="14" pageNumber="73">— Spire higher; shell height/shell width&gt; 0.7 .................................................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3425B42FF2567288115597A" pageId="14" pageNumber="73">
<paragraph id="8BE20134E3425B42FF25672881125916" blockId="14.[210,1380,1810,1943]" box="[210,1379,1810,1835]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF25672881125916" box="[210,1379,1810,1835]" pageId="14" pageNumber="73">
9 Umbilicus completely open; penial caecum weak ........................
<taxonomicName id="4C5D7AB7E3425B42FC5D672981125916" authority="Wu, Lin &amp; Hwang, 2007" authorityName="Wu, Lin &amp; Hwang" authorityYear="2007" box="[938,1379,1810,1835]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="longkiauwensis">
<emphasis id="B929DD26E3425B42FC5D672980275916" box="[938,1110,1810,1834]" italics="true" pageId="14" pageNumber="73">S. longkiauwensis</emphasis>
<bibRefCitation id="EFCC7CC5E3425B42FBAB672981125916" author="WU S. - P. &amp; LIN Y. - S. &amp; HWANG C. - C." box="[1116,1379,1810,1835]" pageId="14" pageNumber="73" pagination="59 - 68" refId="ref9690" refString="WU S. - P., LIN Y. - S. &amp; HWANG C. - C. 2007. - A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan. Zootaxa 1608 (1): 59 - 68. https: // doi. org / 10.11646 / zootaxa. 1608.1.8" type="journal article" year="2007">Wu, Lin &amp; Hwang, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3425B42FF2567158115597A" blockId="14.[210,1380,1810,1943]" box="[210,1380,1839,1863]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF2567158115597A" box="[210,1380,1839,1863]" pageId="14" pageNumber="73">
— Umbilicus partly open; penial caecum low conical ..................................................................
<taxonomicName id="4C5D7AB7E3425B42FB2667158157597B" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[1233,1318,1839,1863]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="nivnu" status="n. sp.">
<emphasis id="B929DD26E3425B42FB2667158157597B" box="[1233,1318,1839,1863]" italics="true" pageId="14" pageNumber="73">S. nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3425B42FADB67158115597A" box="[1324,1380,1839,1862]" pageId="14" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3425B42FF256759811559AB" pageId="14" pageNumber="73">
<paragraph id="8BE20134E3425B42FF25675981155947" blockId="14.[210,1380,1810,1943]" box="[210,1380,1890,1915]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF25675981155947" box="[210,1380,1890,1915]" pageId="14" pageNumber="73">10 Periphery angulate to carinate ................................................................................................................... 11</keyLead>
</paragraph>
<paragraph id="8BE20134E3425B42FF2567BA811559AB" blockId="14.[210,1380,1810,1943]" box="[210,1380,1919,1943]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF2567BA811559AB" box="[210,1380,1919,1943]" pageId="14" pageNumber="73">— Periphery round, at most roundly angulate ................................................................................................ 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3425B42FF256789811559DB" pageId="14" pageNumber="73">
<paragraph id="8BE20134E3425B42FF256789811259F6" blockId="14.[210,1380,1970,2023]" box="[210,1379,1970,1995]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF256789811259F6" box="[210,1379,1970,1995]" pageId="14" pageNumber="73">
11 Umbilicus not covered by reflexed columellar lip .............................................
<taxonomicName id="4C5D7AB7E3425B42FBDF6789811259F6" authority="(Pfeiffer, 1866)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1866" box="[1064,1379,1970,1994]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="73" phylum="Mollusca" rank="species" species="mellea">
<emphasis id="B929DD26E3425B42FBDF678980B059F6" box="[1064,1217,1970,1994]" italics="true" pageId="14" pageNumber="73">Satsuma mellea</emphasis>
(Pfeiffer, 1866)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3425B42FF2567EA811559DB" blockId="14.[210,1380,1970,2023]" box="[210,1380,1999,2023]" pageId="14" pageNumber="73">
<keyLead id="30ACA201E3425B42FF2567EA811559DB" box="[210,1380,1999,2023]" pageId="14" pageNumber="73">— Umbilicus partly covered by reflexed columellar lip ................................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF2760E381135F17" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF2760E381105F32" blockId="15.[208,1378,217,299]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2760E381105F32" pageId="15" pageNumber="74">
12 Periphery sharply carinate; surface immediately below periphery concave; basal lip flat .................................. .............................................
<taxonomicName id="4C5D7AB7E3435B43FDC160CC87CA5F32" authority="(Mollendorff, 1884)" baseAuthorityName="Mollendorff" baseAuthorityYear="1884" box="[566,955,246,270]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="friesiana">
<emphasis id="B929DD26E3435B43FDC160CC86955F32" box="[566,740,246,270]" italics="true" pageId="15" pageNumber="74">Satsuma friesiana</emphasis>
(Möllendorff, 1884)
</taxonomicName>
+ syn.
<taxonomicName id="4C5D7AB7E3435B43FBF260CC81105F32" authority="Chang, 1990" authorityName="Chang" authorityYear="1990" box="[1029,1377,246,270]" class="Gastropoda" family="Camaenidae" genus="Coniglobus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="carinatus">
<emphasis id="B929DD26E3435B43FBF260CC80A25F32" box="[1029,1235,246,270]" italics="true" pageId="15" pageNumber="74">Coniglobus carinatus</emphasis>
Chang, 1990
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27612981135F17" blockId="15.[208,1378,217,299]" box="[208,1378,275,299]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27612981135F17" box="[208,1378,275,299]" pageId="15" pageNumber="74">— Periphery bluntly carinate; surface immediately below periphery convex; basal lip smoothly curved .......... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27617C81135F47" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27617C81135F62" blockId="15.[208,1378,326,459]" box="[208,1378,326,350]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27617C81135F62" box="[208,1378,326,350]" pageId="15" pageNumber="74">13 Periphery angulate in front; whorls flat to weakly convex .......................................................................... 14</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27615981135F47" blockId="15.[208,1378,326,459]" box="[208,1378,355,379]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27615981135F47" box="[208,1378,355,379]" pageId="15" pageNumber="74">— Periphery round in front; whorls convex ................................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF2761AC81135FF7" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF2761AC81105F92" blockId="15.[208,1378,326,459]" box="[208,1377,406,430]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2761AC81105F92" box="[208,1377,406,430]" pageId="15" pageNumber="74">
14 Whorls flat; base convex; axial striae dense ........................................
<taxonomicName id="4C5D7AB7E3435B43FC3861AD81105F92" authority="(H. Adams, 1866)" baseAuthorityName="H. Adams" baseAuthorityYear="1866" box="[975,1377,406,430]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="subSpecies" species="succincta" subSpecies="succincta">
<emphasis id="B929DD26E3435B43FC3861AD80EF5F92" box="[975,1182,407,430]" italics="true" pageId="15" pageNumber="74">S. succincta succincta</emphasis>
(H. Adams, 1866)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27618981135FF7" blockId="15.[208,1378,326,459]" box="[208,1378,435,459]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27618981135FF7" box="[208,1378,435,459]" pageId="15" pageNumber="74">
— Whorls convex; base less convex; axial striae loose ...............................................
<taxonomicName id="4C5D7AB7E3435B43FBBF618981135FF7" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1096,1378,435,459]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="paiwanis">
<emphasis id="B929DD26E3435B43FBBF618980CB5FF7" box="[1096,1210,435,459]" italics="true" pageId="15" pageNumber="74">S. paiwanis</emphasis>
(
<bibRefCitation id="EFCC7CC5E3435B43FB3F6189812B5FF7" author="KURODA T." box="[1224,1370,435,459]" pageId="15" pageNumber="74" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF2761DC81135C69" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF2761DC81135C27" blockId="15.[208,1378,486,597]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2761DC81135C27" pageId="15" pageNumber="74">
15 Spire higher; shell height/width ratio = 0.79; peristome very thick; whorls less convex ................................... ...........................................................................................................
<taxonomicName id="4C5D7AB7E3435B43FC24623981135C27" ID-CoL="7V2VW" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[979,1378,515,539]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="subSpecies" species="succincta" subSpecies="rubrotincta">
<emphasis id="B929DD26E3435B43FC24623980CB5C27" box="[979,1210,515,539]" italics="true" pageId="15" pageNumber="74">S. succincta rubrotincta</emphasis>
(
<bibRefCitation id="EFCC7CC5E3435B43FB3F6239812B5C27" author="KURODA T." box="[1224,1370,515,539]" pageId="15" pageNumber="74" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27621A81135C69" blockId="15.[208,1378,486,597]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27621A81135C69" pageId="15" pageNumber="74">
— Spire lower; shell height/width ratio = 0.69; peristome not particularly thickened; whorls convex .................. ............................................................................................................
<taxonomicName id="4C5D7AB7E3435B43FC2E620781135C69" ID-CoL="7P658" authority="(Pilsbry, 1901)" baseAuthorityName="Pilsbry" baseAuthorityYear="1901" box="[985,1378,572,597]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="subSpecies" species="succincta" subSpecies="amblytropis">
<emphasis id="B929DD26E3435B43FC2E620780B25C68" box="[985,1219,572,596]" italics="true" pageId="15" pageNumber="74">S. succincta amblytropis</emphasis>
(Pilsbry, 1901)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27624A81135C99" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27624A81135CB4" blockId="15.[208,1378,624,757]" box="[208,1378,624,648]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27624A81135CB4" box="[208,1378,624,648]" pageId="15" pageNumber="74">
16 Umbilicus close ................................................................................................
<taxonomicName id="4C5D7AB7E3435B43FBC6624A81135CB4" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1073,1378,624,648]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="takkiriensis">
<emphasis id="B929DD26E3435B43FBC6624A80CB5CB4" box="[1073,1210,624,648]" italics="true" pageId="15" pageNumber="74">S. takkiriensis</emphasis>
(
<bibRefCitation id="EFCC7CC5E3435B43FB3F624A812B5CB4" author="KURODA T." box="[1224,1370,624,648]" pageId="15" pageNumber="74" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF2762B781135C99" blockId="15.[208,1378,624,757]" box="[208,1378,653,677]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2762B781135C99" box="[208,1378,653,677]" pageId="15" pageNumber="74">— Umbilicus partly covered ............................................................................................................................ 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF2762FA81135CC9" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF2762FA81105CE4" blockId="15.[208,1378,624,757]" box="[208,1377,704,728]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2762FA81105CE4" box="[208,1377,704,728]" pageId="15" pageNumber="74">
17 Basal lip curved ...............................................................................................
<taxonomicName id="4C5D7AB7E3435B43FBDC62FA81105CE4" authority="(Chang &amp; Tada, 2000)" baseAuthorityName="Chang &amp; Tada" baseAuthorityYear="2000" box="[1067,1377,704,728]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="litus">
<emphasis id="B929DD26E3435B43FBDC62FA801E5CE4" box="[1067,1135,704,728]" italics="true" pageId="15" pageNumber="74">S. litus</emphasis>
(Chang &amp; Tada, 2000)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF2762E781135CC9" blockId="15.[208,1378,624,757]" box="[208,1378,733,757]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2762E781135CC9" box="[208,1378,733,757]" pageId="15" pageNumber="74">— Basal lip flat ................................................................................................................................................ 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27632A81135D79" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27632A81135D14" blockId="15.[208,1378,784,917]" box="[208,1378,784,808]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27632A81135D14" box="[208,1378,784,808]" pageId="15" pageNumber="74">18 Upper surface roughly and loosely ridged; outer lip sinuate ........................................................................ 19</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27631781135D79" blockId="15.[208,1378,784,917]" box="[208,1378,813,837]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27631781135D79" box="[208,1378,813,837]" pageId="15" pageNumber="74">— Upper surface finely and densely ridged; outer lip smoothly curved ............................................................ 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27635A81135DA8" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27635A81135D44" blockId="15.[208,1378,784,917]" box="[208,1378,864,888]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27635A81135D44" box="[208,1378,864,888]" pageId="15" pageNumber="74">
19 Basal lip horizontal, straight; surface above periphery malleated ...........................
<taxonomicName id="4C5D7AB7E3435B43FBBC635B81135D44" authority="(Mollendorff, 1888)" baseAuthorityName="Mollendorff" baseAuthorityYear="1888" box="[1099,1378,864,888]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="nux">
<emphasis id="B929DD26E3435B43FBBC635B80FA5D44" box="[1099,1163,865,888]" italics="true" pageId="15" pageNumber="74">S. nux</emphasis>
(
<bibRefCitation id="EFCC7CC5E3435B43FB6F635A812B5D44" author="MOLLENDORFF O. F. &amp; VON" box="[1176,1370,864,888]" pageId="15" pageNumber="74" pagination="38 - 44" refId="ref9200" refString="MOLLENDORFF O. F. VON 1888. - Mittheilungen aus dem Gebiete der Malakozoologie. Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 20 (3 / 4): 38 - 44." type="journal article" year="1888">Möllendorff, 1888</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27634781135DA8" blockId="15.[208,1378,784,917]" box="[208,1378,893,917]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27634781135DA8" box="[208,1378,893,917]" pageId="15" pageNumber="74">
— Basal lip straightly downwrds; surface above periphery not malleated ................................
<taxonomicName id="4C5D7AB7E3435B43FB52634781555DA9" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[1189,1316,893,917]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="sunlinksea" status="n. sp.">
<emphasis id="B929DD26E3435B43FB52634781555DA9" box="[1189,1316,893,917]" italics="true" pageId="15" pageNumber="74">S. sunlinksea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3435B43FADD634781135DA8" box="[1322,1378,893,916]" pageId="15" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27638B81135DD9" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27638B81135DF5" blockId="15.[208,1378,944,997]" box="[208,1378,944,969]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27638B81135DF5" box="[208,1378,944,969]" pageId="15" pageNumber="74">20 Spire high, shell height/shell width&gt; 0.8 ................................................................................................... 21</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF2763F481135DD9" blockId="15.[208,1378,944,997]" box="[208,1378,973,997]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2763F481135DD9" box="[208,1378,973,997]" pageId="15" pageNumber="74">— Spire low, shell height/shell width &lt;0.75 ................................................................................................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27643B81135A6D" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27643B81135A09" blockId="15.[208,1378,1024,1106]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27643B81135A09" pageId="15" pageNumber="74">
21 Upper lip smoothly curved; basal lip horizontal with a weak fold; penial pouch absent .................................. ..............................................................................................................................
<taxonomicName id="4C5D7AB7E3435B43FBA0642481135A09" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[1111,1378,1053,1077]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="sericata">
<emphasis id="B929DD26E3435B43FBA0642480CB5A09" box="[1111,1210,1054,1077]" italics="true" pageId="15" pageNumber="74">S. sericata</emphasis>
(
<bibRefCitation id="EFCC7CC5E3435B43FB3F6427812B5A09" author="KURODA T." box="[1224,1370,1053,1077]" pageId="15" pageNumber="74" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF27640081135A6D" blockId="15.[208,1378,1024,1106]" box="[208,1378,1082,1106]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27640081135A6D" box="[208,1378,1082,1106]" pageId="15" pageNumber="74">
— Upper lip sinuous; basal lip oblique without a fold; penial pouch present ..........................
<taxonomicName id="4C5D7AB7E3435B43FB51640081555A6E" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[1190,1316,1082,1106]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="hsuehshan" status="n. sp.">
<emphasis id="B929DD26E3435B43FB51640081555A6E" box="[1190,1316,1082,1106]" italics="true" pageId="15" pageNumber="74">S. hsuehshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3435B43FADD640081135A6D" box="[1322,1378,1082,1105]" pageId="15" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27645781135A9E" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27645781105AB9" blockId="15.[208,1378,1133,1186]" box="[208,1377,1133,1157]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27645781105AB9" box="[208,1377,1133,1157]" pageId="15" pageNumber="74">
22 Tooth on basal lip apparent; flagellum hooked on tip, tapering ............................
<taxonomicName id="4C5D7AB7E3435B43FBA6645481105AB9" ID-CoL="6XQB6" authority="Wu &amp; Tsai, 2015" authorityName="Wu &amp; Tsai" authorityYear="2015" box="[1105,1377,1133,1157]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="chengi">
<emphasis id="B929DD26E3435B43FBA6645480D85AB9" box="[1105,1193,1133,1157]" italics="true" pageId="15" pageNumber="74">S. chengi</emphasis>
<bibRefCitation id="EFCC7CC5E3435B43FB58645481105AB9" author="WU S. - P. &amp; TSAI C. - L." box="[1199,1377,1133,1157]" pageId="15" pageNumber="74" pagination="41 - 48" refId="ref9521" refString="WU S. - P. &amp; TSAI C. - L. 2015. - A new endemic dextral Satsuma land snail (Pulmonata: Camaenidae) from Taiwan. Bulletin of Malacology 38: 41 - 48." type="journal article" year="2015">Wu &amp; Tsai, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF2764B181135A9E" blockId="15.[208,1378,1133,1186]" box="[208,1378,1162,1186]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2764B181135A9E" box="[208,1378,1162,1186]" pageId="15" pageNumber="74">— Tooth on basal lip inapparent; flagellum spirally tapering ........................................................................... 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A91991E3435B43FF27648481105B10" pageId="15" pageNumber="74">
<paragraph id="8BE20134E3435B43FF27648481105ACE" blockId="15.[208,1380,1213,1324]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF27648481105ACE" pageId="15" pageNumber="74">
23 Distinguished by molecular characters only; shell larger; shell colour paler; penis shorter; distributed around Chipen, southeastern
<collectingCountry id="F34A41A4E3435B43FE2964E086585ACE" box="[478,553,1242,1266]" name="Taiwan" pageId="15" pageNumber="74">Taiwan</collectingCountry>
....................................................................................
<taxonomicName id="4C5D7AB7E3435B43FB9864E181105ACE" authority="Wu &amp; Wu, 2017" authorityName="Wu &amp; Wu" authorityYear="2017" box="[1135,1377,1242,1266]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="mii">
<emphasis id="B929DD26E3435B43FB9864E180DA5ACE" box="[1135,1195,1243,1266]" italics="true" pageId="15" pageNumber="74">S. mii</emphasis>
Wu &amp; Wu, 2017
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE20134E3435B43FF2764CD81105B10" blockId="15.[208,1380,1213,1324]" pageId="15" pageNumber="74">
<keyLead id="30ACA201E3435B43FF2764CD81105B10" pageId="15" pageNumber="74">
— Distinguished by molecular characters only; shell smaller; shell colour darker; penis longer; distributed around Antong, eastern
<collectingCountry id="F34A41A4E3435B43FE52652E85815B10" box="[421,496,1300,1324]" name="Taiwan" pageId="15" pageNumber="74">Taiwan</collectingCountry>
....................................................................................
<taxonomicName id="4C5D7AB7E3435B43FBC3652E81105B10" authority="Wu &amp; Tsai, 2016" authorityName="Wu &amp; Tsai" authorityYear="2016" box="[1076,1377,1300,1324]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="74" phylum="Mollusca" rank="species" species="paradoxa">
<emphasis id="B929DD26E3435B43FBC3652E80D95B10" box="[1076,1192,1300,1324]" italics="true" pageId="15" pageNumber="74">S. paradoxa</emphasis>
<bibRefCitation id="EFCC7CC5E3435B43FB58652E81105B10" author="WU S. - P. &amp; TSAI C. - L." box="[1199,1377,1300,1324]" pageId="15" pageNumber="74" pagination="47 - 58" refId="ref9561" refString="WU S. - P. &amp; TSAI C. - L. 2016. - A new dextral species land snail of genus Satsuma (Pulmonata: Camaenidae) endemic to Taiwan. Bulletin of Malacology 39: 47 - 58." type="journal article" year="2016">Wu &amp; Tsai, 2016</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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