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<document id="D3B9E0F50448B6D885AFD5510A994081" ID-DOI="10.5281/zenodo.11512244" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512244" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1717719192119" checkinUser="felipe" docAuthor="Zug, George R." docDate="2015" docId="03856357CC74FFD5FFD0585EFC10608F" docLanguage="en" docName="ProcCalifAcadSci.62.5.135-190.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (5)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Kalophrynus minusculus Iskandar 1998" docType="treatment" docVersion="3" lastPageNumber="171" masterDocId="FFBC1B2FCC57FFF1FFAB5C0AFF8C611C" masterDocTitle="Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs" masterLastPageNumber="190" masterPageNumber="135" pageNumber="170" updateTime="1720720661996" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D6F345CBA67248E2CB21D5A96C9CF157">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
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<mods:namePart id="E317B7F9D0C9FEF710517C907BD5BB45">Zug, George R.</mods:namePart>
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<treatment id="03856357CC74FFD5FFD0585EFC10608F" ID-DOI="http://doi.org/10.5281/zenodo.11512396" ID-Zenodo-Dep="11512396" LSID="urn:lsid:plazi:treatment:03856357CC74FFD5FFD0585EFC10608F" httpUri="http://treatment.plazi.org/id/03856357CC74FFD5FFD0585EFC10608F" lastPageId="36" lastPageNumber="171" pageId="35" pageNumber="170">
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<paragraph id="8B93D241CC74FFD2FFD0585EFB026570" blockId="35.[123,1166,1108,1132]" box="[123,1166,1108,1132]" pageId="35" pageNumber="170">
<heading id="D0DB652DCC74FFD2FFD0585EFB026570" box="[123,1166,1108,1132]" centered="true" fontSize="10" level="2" pageId="35" pageNumber="170" reason="4">
<taxonomicName id="4C2CA9C2CC74FFD2FFD0585EFB026570" ID-CoL="6NK4X" authority="Iskandar, 1998 Dwarf Sticky Frog" authorityName="Iskandar" authorityYear="1998" box="[123,1166,1108,1132]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="35" pageNumber="170" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="B9580E53CC74FFD2FFD0585EFB026570" bold="true" box="[123,1166,1108,1132]" pageId="35" pageNumber="170">
<emphasis id="B9580E53CC74FFD2FFD0585EFE036570" bold="true" box="[123,399,1108,1132]" italics="true" pageId="35" pageNumber="170">Kalophrynus minusculus</emphasis>
<bibRefCitation id="EFBDAFB0CC74FFD2FE3C585EFDB36570" author="ISKANDAR, D. J." box="[407,575,1108,1132]" pageId="35" pageNumber="170" refId="ref37710" refString="ISKANDAR, D. J. 1998. The Amphibians of Java and Bali. Research and Development Centre for Biology - LIPI, Indonesia [no city given]." type="book" year="1998">Iskandar, 1998</bibRefCitation>
Dwarf Sticky Frog
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC74FFD2FFD05876FD8565AE" pageId="35" pageNumber="170" type="reference_group">
<paragraph id="8B93D241CC74FFD2FFD05876FD8565AE" blockId="35.[123,1173,1148,1202]" pageId="35" pageNumber="170">
<taxonomicName id="4C2CA9C2CC74FFD2FFD05876FD8C658D" ID-CoL="6NK4X" authority="Iskandar, 1998" authorityName="Iskandar" authorityYear="1998" box="[123,512,1148,1170]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="35" pageNumber="170" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="B9580E53CC74FFD2FFD05876FEE6658D" box="[123,362,1148,1169]" italics="true" pageId="35" pageNumber="170">Kalophrynus minusculus</emphasis>
<bibRefCitation id="EFBDAFB0CC74FFD2FEDA5876FD8C658D" author="ISKANDAR, D. J." box="[369,512,1148,1170]" pageId="35" pageNumber="170" refId="ref37710" refString="ISKANDAR, D. J. 1998. The Amphibians of Java and Bali. Research and Development Centre for Biology - LIPI, Indonesia [no city given]." type="book" year="1998">Iskandar, 1998</bibRefCitation>
</taxonomicName>
,
<emphasis id="B9580E53CC74FFD2FDA65876FCD8658D" box="[525,852,1148,1170]" italics="true" pageId="35" pageNumber="170">
The Amphibians of
<collectingRegion id="49E81CA3CC74FFD2FD675877FCD8658D" box="[716,852,1148,1170]" isEnumeration="true" pageId="35" pageNumber="170">Java and Bali</collectingRegion>
</emphasis>
<materialsCitation id="3B44D81CCC74FFD2FCF05877FD8F65AD" country="Indonesia" location="Cigeunteur" municipality="Ujung Kulon" pageId="35" pageNumber="170" specimenCount="1" stateProvince="Jawa Barat">
p.53 [
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locality: “
<location id="8EF3849ACC74FFD2FB8D5876FB1C658E" LSID="urn:lsid:plazi:treatment:03856357CC74FFD5FFD0585EFC10608F:8EF3849ACC74FFD2FB8D5876FB1C658E" box="[1062,1168,1148,1170]" country="Indonesia" municipality="Ujung Kulon" name="Cigeunteur" pageId="35" pageNumber="170" stateProvince="Jawa Barat">Cigeunteur</location>
,
<collectingMunicipality id="6BF7483BCC74FFD2FF305896FE9665AE" box="[155,282,1180,1202]" pageId="35" pageNumber="170">Ujung Kulon</collectingMunicipality>
,
<collectingRegion id="49E81CA3CC74FFD2FE8D5897FE0B65AE" box="[294,391,1181,1202]" country="Indonesia" name="Jawa Barat" pageId="35" pageNumber="170">West Java</collectingRegion>
”,
<collectingCountry id="F33B92D1CC74FFD2FE355896FE7765AE" box="[414,507,1180,1202]" name="Indonesia" pageId="35" pageNumber="170">Indonesia</collectingCountry>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC74FFD2FF0058CFFCB36761" pageId="35" pageNumber="170" type="materials_examined">
<paragraph id="8B93D241CC74FFD2FF0058CFFD5B6401" blockId="35.[123,1174,1221,1725]" pageId="35" pageNumber="170">
<materialsCitation id="3B44D81CCC74FFD2FF0058CFFC3165C1" box="[171,957,1221,1245]" collectionCode="TYPE, MATERIAL" pageId="35" pageNumber="170" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED3D4A84CC74FFD2FF0058CFFF6865C0" box="[171,228,1221,1245]" pageId="35" pageNumber="170">TYPE</collectionCode>
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.—
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<typeStatus id="54976CE3CC74FFD2FE2358CCFE7B65C0" box="[392,503,1222,1245]" pageId="35" pageNumber="170" type="holotype">HOLOTYPE</typeStatus>
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: Museum Zoologicum Bogoriense 367
</materialsCitation>
.
<materialsCitation id="3B44D81CCC74FFD2FC7858CCFEBF65E1" collectionCode="MZB" pageId="35" pageNumber="170" specimenCode="MZB 265-66, 375-76" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54976CE3CC74FFD2FC7858CCFBC965C0" box="[979,1093,1222,1245]" pageId="35" pageNumber="170" type="paratype">PARATYPES</typeStatus>
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:
<specimenCode id="DB8A7A3ACC74FFD2FBF758CCFF4765E1" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="170" type="Museum">MZB 26566</specimenCode>
,
<specimenCode id="DB8A7A3ACC74FFD2FF7758EFFEA165E1" box="[220,301,1253,1277]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="170" type="Museum">37576</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B44D81CCC74FFD2FE9558ECFE3365E1" box="[318,447,1253,1277]" collectionCode="MZB" pageId="35" pageNumber="170" specimenCode="MZB 2339" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8A7A3ACC74FFD2FE9558ECFE3B65E1" box="[318,439,1253,1277]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="170" type="Museum">MZB 2339</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B44D81CCC74FFD2FE6258ECFDCF65E1" box="[457,579,1253,1277]" collectionCode="MZB" pageId="35" pageNumber="170" specimenCode="MZB 2924" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8A7A3ACC74FFD2FE6258ECFDCF65E1" box="[457,579,1253,1277]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="170" type="Museum">MZB 2924</specimenCode>
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, and possibly ten
<typeStatus id="54976CE3CC74FFD2FCBE58EFFC4A65E1" box="[789,966,1253,1277]" isOtherTypeStatus="true" pageId="35" pageNumber="170">other specimens</typeStatus>
not designated by museum number. All specimens by original designation.
</paragraph>
<paragraph id="8B93D241CC74FFD2FF00592FFC2F64E1" blockId="35.[123,1174,1221,1725]" pageId="35" pageNumber="170">
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<emphasis id="B9580E53CC74FFD2FF00592FFEA56420" bold="true" box="[171,297,1317,1341]" pageId="35" pageNumber="170">DEFINITION</emphasis>
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.— Small, adult females to
<quantity id="4CD47FA4CC74FFD2FDFE592FFD136421" box="[597,671,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="170" unit="mm" value="35.0">35 mm</quantity>
<collectionCode id="ED3D4A84CC74FFD2FD03592FFD566421" box="[680,730,1317,1341]" pageId="35" pageNumber="170">SVL</collectionCode>
(
<emphasis id="B9580E53CC74FFD2FD40592CFD746421" box="[747,760,1318,1341]" italics="true" pageId="35" pageNumber="170">n</emphasis>
= 1), adult males to
<quantity id="4CD47FA4CC74FFD2FC77592FFBAA6421" box="[988,1062,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="35" pageNumber="170" unit="mm" value="25.0">25 mm</quantity>
<collectionCode id="ED3D4A84CC74FFD2FB84592FFBED6421" box="[1071,1121,1317,1341]" pageId="35" pageNumber="170">SVL</collectionCode>
(
<emphasis id="B9580E53CC74FFD2FBD9592CFBF36421" box="[1138,1151,1318,1341]" italics="true" pageId="35" pageNumber="170">n</emphasis>
= 10); head modest length NA % HeadL/
<collectionCode id="ED3D4A84CC74FFD2FD82594FFDD76441" box="[553,603,1349,1373]" pageId="35" pageNumber="170">SVL</collectionCode>
; head wider than long NA % HeadW/HeadL; snout moderately broad NA % SnW/HeadL; naris closer to snout than to eye NA % NarEye/SnEye; eye moderately large NA % EyeD/HeadL; tympanum visible and smaller than eye NA % Tymp/EyeD; moderate length forelimb NA % Forarm/
<collectionCode id="ED3D4A84CC74FFD2FD9B59AFFDEE64A1" box="[560,610,1445,1469]" pageId="35" pageNumber="170">SVL</collectionCode>
and forearm equal crus length NA % Forarm/CrusL; hindlimb moderately long NA % HndlL/
<collectionCode id="ED3D4A84CC74FFD2FD8459CFFDED64C1" box="[559,609,1477,1501]" pageId="35" pageNumber="170">SVL</collectionCode>
, NA % CrusL/
<collectionCode id="ED3D4A84CC74FFD2FCAF59CFFCBA64C1" box="[772,822,1477,1501]" pageId="35" pageNumber="170">SVL</collectionCode>
, and NA % CrusL/ThghL; hindfoot well developed NA % HndfL/CrusL; limb measurements not reported.
</paragraph>
<paragraph id="8B93D241CC74FFD2FF005A0FFE026721" blockId="35.[123,1174,1221,1725]" pageId="35" pageNumber="170">Vomerine teeth presence or absence not reported; palatal fold morphology not reported; tongue morphology not reported.</paragraph>
<paragraph id="8B93D241CC74FFD2FF005A4FFDD36741" blockId="35.[123,1174,1221,1725]" box="[171,607,1605,1629]" pageId="35" pageNumber="170">Hand and foot morphology not reported.</paragraph>
<paragraph id="8B93D241CC74FFD2FF005A6CFCB36761" blockId="35.[123,1174,1221,1725]" box="[171,831,1637,1661]" pageId="35" pageNumber="170">In life, dorsum brownish black with darker bands and stripes.</paragraph>
</subSubSection>
<subSubSection id="C33681CACC74FFD2FF005A8CFDCC67A1" pageId="35" pageNumber="170" type="etymology">
<paragraph id="8B93D241CC74FFD2FF005A8CFDCC67A1" blockId="35.[123,1174,1221,1725]" pageId="35" pageNumber="170">
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<emphasis id="B9580E53CC74FFD2FF005A8CFEB96780" bold="true" box="[171,309,1670,1694]" pageId="35" pageNumber="170">ETYMOLOGY</emphasis>
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.— The specific epithet
<taxonomicName id="4C2CA9C2CC74FFD2FD965A8FFD396781" authorityName="Iskandar" authorityYear="1998" box="[573,693,1669,1693]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="35" pageNumber="170" phylum="Chordata" rank="species" species="minusculus">
<emphasis id="B9580E53CC74FFD2FD965A8FFD396781" box="[573,693,1669,1693]" italics="true" pageId="35" pageNumber="170">minusculus</emphasis>
</taxonomicName>
refers to small size of adults relative to the larger
<taxonomicName id="4C2CA9C2CC74FFD2FF6B5AACFE7667A1" box="[192,506,1701,1725]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="35" pageNumber="170" phylum="Chordata" rank="species" species="interlineatus-pleurostigma">
<emphasis id="B9580E53CC74FFD2FF6B5AACFE7667A1" box="[192,506,1701,1725]" italics="true" pageId="35" pageNumber="170">K. interlineatus-pleurostigma</emphasis>
</taxonomicName>
frogs.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC73FFD5FF005CD1FCF3606F" pageId="36" pageNumber="171" type="distribution">
<paragraph id="8B93D241CC73FFD5FF005CD1FEEC602F" blockId="36.[123,1174,219,403]" pageId="36" pageNumber="171">
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<emphasis id="B9580E53CC73FFD5FF005CD1FECF61ED" bold="true" box="[171,323,219,243]" pageId="36" pageNumber="171">DISTRIBUTION</emphasis>
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.—
<materialsCitation id="3B44D81CCC73FFD5FEC05CD1FE1B600F" collectorName="This" location="Ujung" municipality="Peucang Island" pageId="36" pageNumber="171" specimenCount="1" stateProvince="West">
<collectorName id="26D9B797CC73FFD5FEC05CD1FE1661EF" box="[363,410,219,243]" pageId="36" pageNumber="171">This</collectorName>
species appears to be fairly common on
<collectingMunicipality id="6BF7483BCC73FFD5FCC55CD1FB9861EF" box="[878,1044,219,243]" pageId="36" pageNumber="171">Peucang Island</collectingMunicipality>
and on the mainland of
<location id="8EF3849ACC73FFD5FEA35CF1FEC7600F" LSID="urn:lsid:plazi:treatment:03856357CC74FFD5FFD0585EFC10608F:8EF3849ACC73FFD5FEA35CF1FEC7600F" box="[264,331,251,275]" municipality="Peucang Island" name="Ujung" pageId="36" pageNumber="171" stateProvince="West">Ujung</location>
<collectingRegion id="49E81CA3CC73FFD5FEFF5CF1FE1B600F" box="[340,407,251,275]" country="Cameroon" name="West" pageId="36" pageNumber="171">Kulon</collectingRegion>
</materialsCitation>
<materialsCitation id="3B44D81CCC73FFD5FE0A5CF1FDFB600F" box="[417,631,251,275]" collectorName="National Park" location="Java" pageId="36" pageNumber="171" specimenCount="1" stateProvince="Jawa Timur">
<collectorName id="26D9B797CC73FFD5FE0A5CF1FDBB600F" box="[417,567,251,275]" pageId="36" pageNumber="171">National Park</collectorName>
,
<collectingRegion id="49E81CA3CC73FFD5FDE35CF6FDFB600F" box="[584,631,252,275]" country="Indonesia" name="Jawa Timur" pageId="36" pageNumber="171">Java</collectingRegion>
</materialsCitation>
.
<materialsCitation id="3B44D81CCC73FFD5FD235CF6FB19600F" box="[648,1173,251,275]" collectorName="It" location="Sumatra" pageId="36" pageNumber="171" specimenCount="1" stateProvince="Sumatera Utara">
<collectorName id="26D9B797CC73FFD5FD235CF6FD14600F" box="[648,664,252,275]" pageId="36" pageNumber="171">It</collectorName>
is also reported to occur in southern
<collectingRegion id="49E81CA3CC73FFD5FB975CF1FB19600F" box="[1084,1173,251,275]" country="Indonesia" name="Sumatera Utara" pageId="36" pageNumber="171">Sumatra</collectingRegion>
</materialsCitation>
<materialsCitation id="3B44D81CCC73FFD5FFD05D11FED5602F" box="[123,345,283,308]" collectorName="It" isTailEnumeration="true" location="Sumatra" pageId="36" pageNumber="171" specimenCount="1" stateProvince="Lampung">
(
<collectingRegion id="49E81CA3CC73FFD5FF2F5D16FEDC602F" box="[132,336,283,308]" country="Indonesia" name="Lampung" pageId="36" pageNumber="171">Lampung Province</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B93D241CC73FFD5FF005D31FCF3606F" blockId="36.[123,1174,219,403]" pageId="36" pageNumber="171">
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<emphasis id="B9580E53CC73FFD5FF005D31FE9D604E" bold="true" box="[171,273,315,339]" pageId="36" pageNumber="171">NATURAL</emphasis>
</smallCapsWord>
<smallCapsWord id="8D75449DCC73FFD5FEB35D31FEF5604E" baselines="333,333" box="[280,377,315,339]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="History" pageId="36" pageNumber="171">
<emphasis id="B9580E53CC73FFD5FEB35D31FEF5604E" bold="true" box="[280,377,315,339]" pageId="36" pageNumber="171">HISTORY</emphasis>
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.—
<taxonomicName id="4C2CA9C2CC73FFD5FE0A5D31FD29604F" authorityName="Iskandar" authorityYear="1998" box="[417,677,315,339]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="36" pageNumber="171" phylum="Chordata" rank="species" species="minisculus">
<emphasis id="B9580E53CC73FFD5FE0A5D31FD29604F" box="[417,677,315,339]" italics="true" pageId="36" pageNumber="171">Kalophrynus minisculus</emphasis>
</taxonomicName>
<materialsCitation id="3B44D81CCC73FFD5FD065D36FCF4606F" location="Females" pageId="36" pageNumber="171" specimenCount="50" specimenCount-egg="50" stateProvince="Jawa Barat">
occurs only in forest at low altitudes in
<collectingRegion id="49E81CA3CC73FFD5FBCA5D36FF26606F" country="Indonesia" name="Jawa Barat" pageId="36" pageNumber="171">West Java</collectingRegion>
.
<location id="8EF3849ACC73FFD5FF135D51FE9D606F" LSID="urn:lsid:plazi:treatment:03856357CC74FFD5FFD0585EFC10608F:8EF3849ACC73FFD5FF135D51FE9D606F" box="[184,273,347,371]" name="Females" pageId="36" pageNumber="171" stateProvince="Jawa Barat">Females</location>
bear 30 to
<specimenCount id="9D2A19C8CC73FFD5FE275D51FE536068" box="[396,479,347,372]" count="50" pageId="36" pageNumber="171" type="egg">50 eggs</specimenCount>
which hatch into nonfeeding tadpoles
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC73FFD5FF005D71FC10608F" box="[171,924,379,403]" pageId="36" pageNumber="171" type="discussion">
<paragraph id="8B93D241CC73FFD5FF005D71FC10608F" blockId="36.[123,1174,219,403]" box="[171,924,379,403]" pageId="36" pageNumber="171">
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<emphasis id="B9580E53CC73FFD5FF005D71FEA4608E" bold="true" box="[171,296,379,403]" pageId="36" pageNumber="171">COMMENTS</emphasis>
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.— Preceding information extracted from Iskander (1998).
</paragraph>
</subSubSection>
</treatment>
</document>