416 lines
51 KiB
XML
416 lines
51 KiB
XML
<document id="28C9893F731F4A7AE119A17B1F94D1B0" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago & Vaz-De, Fernando Z." docDate="2021" docId="03850669FF96FFC9FF10274D3E2FCDE2" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius agenor" docType="key" docVersion="7" lastPageNumber="8" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="7" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C5B2D52542CA0608991B5584A9225E2B">
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<mods:title id="94678E31AD8E289F18729EED58D3A404">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="A7091BD13EF1D357E1A3D25032844167" type="personal">
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<mods:roleTerm id="FC5499EFE59C527E1BEF5B1DAA2105FB">Author</mods:roleTerm>
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<mods:namePart id="2DD3E295421AF1E317801DEA36AC63F7">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="A9382D25DDE3094ACECAD7BCF7E3DE96">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="51E7ADC9DA38BB1F4F2535BF1402A241" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="200FED7904CBE379E2C48AFA01CE42D0">Vaz-De, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="AD8C72C2C5B7A14F9C4A6386E6499ADD">text</mods:typeOfResource>
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<mods:title id="7D019123D4E6091B260EBA25C2AD718F">European Journal of Taxonomy</mods:title>
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<mods:part id="1350F9EA583020EA1EEBE51B0C00A01B">
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<mods:date id="38109E59D2CE5B50551BBF7CE8EA41E0">2021</mods:date>
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<mods:number id="CF8E3089619B56A7922DA2C25DA564AE">2021-02-11</mods:number>
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<mods:number id="6CF81BEAB789474A0D9AC5B4D5C57AB1">734</mods:number>
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<treatment id="03850669FF96FFC9FF10274D3E2FCDE2" ID-DOI="http://doi.org/10.5281/zenodo.4545704" ID-GBIF-Taxon="177420813" ID-Zenodo-Dep="4545704" LSID="urn:lsid:plazi:treatment:03850669FF96FFC9FF10274D3E2FCDE2" httpUri="http://treatment.plazi.org/id/03850669FF96FFC9FF10274D3E2FCDE2" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="C336E4F4FF96FFC8FF10274D3E2FCA75" box="[189,1398,264,291]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8B93B77FFF96FFC8FF10274D3E2FCA75" blockId="6.[189,1398,264,291]" box="[189,1398,264,291]" pageId="6" pageNumber="7">
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<heading id="D0DB0013FF96FFC8FF10274D3E2FCA75" box="[189,1398,264,291]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
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<emphasis id="B9586B6DFF96FFC8FF10274D3E2FCA75" bold="true" box="[189,1398,264,291]" pageId="6" pageNumber="7">
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Identification key to the species of the
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<taxonomicName id="4C2CCCFCFF96FFC8FD39274D38D1CA75" ID-CoL="6CS36" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[660,904,264,291]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="agenor">
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<emphasis id="B9586B6DFF96FFC8FD39274D38D1CA75" bold="true" box="[660,904,264,291]" italics="true" pageId="6" pageNumber="7">Dichotomius agenor</emphasis>
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</taxonomicName>
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species group based on male characters
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C336E4F4FF96FFC9FF1027723E2FCDE2" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="key">
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<key id="FFBDE04EFF96FFC9FF1027723E2FCDE2" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<keyStep id="30D8AFDAFF96FFC8FF1027723E2ECAED" pageId="6" pageNumber="7">
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<paragraph id="8B93B77FFF96FFC8FF1027723E2FCA22" blockId="6.[189,1399,311,443]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF1027723E2FCA22" pageId="6" pageNumber="7">
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1. Dorsally shining coppery, sometimes with green reflections. Large males with cephalic carina narrow, wider than high (
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<collectingCountry id="F33BF7EFFF96FFC8FDBF271F3907CA22" box="[530,606,346,372]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
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,
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<collectingCountry id="F33BF7EFFF96FFC8FDC4271F3994CA22" box="[617,717,346,372]" name="Trinidad and Tobago" pageId="6" pageNumber="7">Trinidad</collectingCountry>
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and
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<collectingCountry id="F33BF7EFFF96FFC8FCAB271F383ACA22" box="[774,867,346,372]" name="Guyana" pageId="6" pageNumber="7">Guyana</collectingCountry>
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)..
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<taxonomicName id="4C2CCCFCFF96FFC8FCD0271F3E2FCA22" ID-CoL="7ZN5G" authority="(Castelnau, 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[893,1398,346,373]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="subaeneus">
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<emphasis id="B9586B6DFF96FFC8FCD0271F3FC7CA23" bold="true" box="[893,1182,346,373]" italics="true" pageId="6" pageNumber="7">Dichotomius subaeneus</emphasis>
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(
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<bibRefCitation id="EFBDCA8EFF96FFC8FB03271F3E37CA22" author="Castelnau M." box="[1198,1390,346,372]" pageId="6" pageNumber="7" pagination="1 - 564" refId="ref29645" refString="Castelnau M. 1840. Histoire naturelle des coleopteres. In: Blanchard E. (eds) Histoire naturelle des animaux articules. Vol. 2: 1 - 564. Dumenil, Paris." type="journal article" year="1840">Castelnau, 1840</bibRefCitation>
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)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8B93B77FFF96FFC8FF6C273B3E2ECAED" blockId="6.[189,1399,311,443]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF6C273B3E2ECAED" pageId="6" pageNumber="7">
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– Dorsally black, brown or dull dark blue, or if metallic green then size less than
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<quantity id="4CD41A9AFF96FFC8FB2A27383F85CACE" box="[1159,1244,381,408]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
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. Large males with cephalic carina conspicuous ...................................................................................................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30D8AFDAFF96FFC8FF1027A73E2EC999" pageId="6" pageNumber="7">
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<paragraph id="8B93B77FFF96FFC8FF1027A73E2EC930" blockId="6.[189,1400,482,720]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF1027A73E2EC930" pageId="6" pageNumber="7">
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2. Dorsally shining green except for the elytra which are opaque. Cephalic carina undeveloped, narrow. (Anterior edge of the metasternum rugose.) Sixth abdominal sternite and pygidium lower, more depressed than the other sternites. Body length less than
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<quantity id="4CD41A9AFF96FFC8FBCF246D3FE6C915" box="[1122,1215,552,579]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
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. (
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<collectingCountry id="F33BF7EFFF96FFC8FB74246D3E7FC914" box="[1241,1318,552,578]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
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:
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<collectingRegion id="49E8799DFF96FFC8FA94246C3A66C930" country="Brazil" name="Mato Grosso" pageId="6" pageNumber="7">Mato Grosso</collectingRegion>
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)............................................................................................
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<taxonomicName id="4C2CCCFCFF96FFC8FC42240E3E4CC933" ID-CoL="7VQMT" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[1007,1301,587,614]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rafanunezi" status="sp. nov.">
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||
<emphasis id="B9586B6DFF96FFC8FC42240E3E4CC933" bold="true" box="[1007,1301,587,614]" italics="true" pageId="6" pageNumber="7">Dichotomius rafanunezi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26BD616FF96FFC8FAB124093E2EC930" box="[1308,1399,588,614]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
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||
</paragraph>
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||
<paragraph id="8B93B77FFF96FFC8FF6C242A3E2EC999" blockId="6.[189,1400,482,720]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF6C242A3E2EC999" pageId="6" pageNumber="7">
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– Dorsally generally black or at most dull dark blue. Cephalic carina well developed, conspicuous. Sixth abdominal and pygidium as high as the other sternites. Body length greater than
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<quantity id="4CD41A9AFF96FFC8FAB624D73E2CC9FB" box="[1307,1397,658,685]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
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, except for
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<taxonomicName id="4C2CCCFCFF96FFC8FEC524F3395AC999" ID-CoL="7VQMR" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[360,515,693,719]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fornicatus">
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<emphasis id="B9586B6DFF96FFC8FEC524F3395AC999" box="[360,515,693,719]" italics="true" pageId="6" pageNumber="7">D. fornicatus</emphasis>
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</taxonomicName>
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(dark blue)
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<emphasis id="B9586B6DFF96FFC8FD3F24F33E38C999" box="[658,1377,694,719]" italics="true" pageId="6" pageNumber="7">..................................................................................................</emphasis>
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3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30D8AFDAFF96FFC8FF1024B23E2EC801" pageId="6" pageNumber="7">
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<paragraph id="8B93B77FFF96FFC8FF1024B23E2FC863" blockId="6.[189,1399,759,856]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF1024B23E2FC863" pageId="6" pageNumber="7">
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3. Pronotum surface subrugose. Elytral striae crenulate (
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<collectingCountry id="F33BF7EFFF96FFC8FCCD24B238BCC847" box="[864,997,759,785]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
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,
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<collectingCountry id="F33BF7EFFF96FFC8FC5D24B23F36C847" box="[1008,1135,759,785]" name="Guatemala" pageId="6" pageNumber="7">Guatemala</collectingCountry>
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and
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<collectingCountry id="F33BF7EFFF96FFC8FB0524B23E7AC844" box="[1192,1315,759,786]" name="Nicaragua" pageId="6" pageNumber="7">Nicaragua</collectingCountry>
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)......... ..................................................................................................................
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<taxonomicName id="4C2CCCFCFF96FFC8FB98255F3E2FC863" ID-CoL="6CRQV" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1077,1398,793,821]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="centralis">
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<emphasis id="B9586B6DFF96FFC8FB98255F3F99C862" bold="true" box="[1077,1216,793,820]" italics="true" pageId="6" pageNumber="7">D. centralis</emphasis>
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(
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<bibRefCitation id="EFBDCA8EFF96FFC8FB7D255F3E37C863" author="Harold E." box="[1232,1390,794,821]" pageId="6" pageNumber="7" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
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)
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</taxonomicName>
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||
</keyLead>
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</paragraph>
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||
<paragraph id="8B93B77FFF96FFC8FF6C257B3E2EC801" blockId="6.[189,1399,759,856]" box="[193,1399,829,856]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF6C257B3E2EC801" box="[193,1399,829,856]" pageId="6" pageNumber="7">– Pronotum with punctures, never rugose. Elytral striae not crenulate................................................ 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30D8AFDAFF96FFC8FF10253A3E2ECF3B" pageId="6" pageNumber="7">
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<paragraph id="8B93B77FFF96FFC8FF10253A3E2FCF54" blockId="6.[189,1399,894,1133]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF10253A3E2FCF54" pageId="6" pageNumber="7">
|
||
4. Metasternum lateral edges smooth, except for minute setae arising from the centre of each ocellate puncture. Cephalic carina antero-posteriorly flattened, medial tubercle acute distally, never apically expanded or emarginate medially. Pronotum gibbous, smooth and shiny. Body pilosity spare or absent. (
|
||
<collectingCountry id="F33BF7EFFF96FFC8FEE125AD3A9ACF55" box="[332,451,1000,1027]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
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).........................................................................................
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<taxonomicName id="4C2CCCFCFF96FFC8FBF025AD3E2FCF54" ID-CoL="35NB8" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[1117,1398,1000,1027]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="belus">
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<emphasis id="B9586B6DFF96FFC8FBF025AD3F98CF55" bold="true" box="[1117,1217,1000,1027]" italics="true" pageId="6" pageNumber="7">D. belus</emphasis>
|
||
(
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||
<bibRefCitation id="EFBDCA8EFF96FFC8FB7D25AD3E37CF54" author="Harold E." box="[1232,1390,1000,1026]" pageId="6" pageNumber="7" pagination="13 - 46" refId="ref29882" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Stettiner entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880</bibRefCitation>
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)
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</taxonomicName>
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||
</keyLead>
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</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C22493E2ECF3B" blockId="6.[189,1399,894,1133]" pageId="6" pageNumber="7">
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||
<keyLead id="30DD144AFF96FFC8FF6C22493E2ECF3B" pageId="6" pageNumber="7">
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||
– Metasternum lateral edges usually covered by long/large setae, if smooth then body length less than
|
||
<quantity id="4CD41A9AFF96FFC8FF47226B3A66CF1F" box="[234,319,1070,1097]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
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. Cephalic carina not flattened, medial tubercle not acute distally and in some species apically emarginate. Pronotum with small knob, with well-defined punctures. Body pilosity dense............. 5
|
||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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<keyStep id="30D8AFDAFF96FFC8FF1022D63E2ECF87" pageId="6" pageNumber="7">
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<paragraph id="8B93B77FFF96FFC8FF1022D63E2ECFFB" blockId="6.[189,1399,1171,1233]" box="[189,1399,1171,1198]" pageId="6" pageNumber="7">
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<keyLead id="30DD144AFF96FFC8FF1022D63E2ECFFB" box="[189,1399,1171,1198]" pageId="6" pageNumber="7">5. Ventral clypeal process cordate, truncate or bifurcate ....................................................................... 6</keyLead>
|
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</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C22F23E2ECF87" blockId="6.[189,1399,1171,1233]" box="[193,1399,1206,1233]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF6C22F23E2ECF87" box="[193,1399,1206,1233]" pageId="6" pageNumber="7">– Ventral clypeal process cylindrical, never cordate, truncate or bifurcate .......................................... 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF96FFC8FF1022BD3E2FCE63" pageId="6" pageNumber="7">
|
||
<paragraph id="8B93B77FFF96FFC8FF1022BD3E2ECE44" blockId="6.[189,1399,1272,1334]" box="[189,1399,1272,1298]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF1022BD3E2ECE44" box="[189,1399,1272,1298]" pageId="6" pageNumber="7">6. Metasternum with dense pilosity that conceals the metepisternal suture.......................................... 8</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C23593E2FCE63" blockId="6.[189,1399,1272,1334]" box="[193,1398,1307,1334]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF6C23593E2FCE63" box="[193,1398,1307,1334]" pageId="6" pageNumber="7">– Metasternum without dense pilosity, metepisternal suture clearly visible .......................................11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF96FFC8FF1023183E2ECD1C" pageId="6" pageNumber="7">
|
||
<paragraph id="8B93B77FFF96FFC8FF1023183E2ECEB6" blockId="6.[189,1399,1372,1611]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF1023183E2ECEB6" pageId="6" pageNumber="7">
|
||
7. Cephalic carina distinctly arcuate in dorsal view. Pronotum with abrupt anterior declivity when viewed laterally, middle surface deeper than lateral, displaying two well-developed lobes on the anteromedial edge. (Interocular space with scattered simple punctures.) Dark blue to black. Body length less than
|
||
<quantity id="4CD41A9AFF96FFC8FE0723833959CEB7" box="[426,512,1478,1505]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
|
||
. (
|
||
<collectingCountry id="F33BF7EFFF96FFC8FDBE238339D4CEB6" box="[531,653,1478,1504]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33BF7EFFF96FFC8FD35238339B7CEB6" box="[664,750,1478,1504]" name="Bolivia" pageId="6" pageNumber="7">Bolivia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF96FFC8FC85238238CECEB7" box="[808,919,1479,1505]" name="Paraguay" pageId="6" pageNumber="7">Paraguay</collectingCountry>
|
||
)........
|
||
<taxonomicName id="4C2CCCFCFF96FFC8FC7323833E2ECEB6" ID-CoL="7VQMR" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[990,1399,1478,1504]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fornicatus">
|
||
<emphasis id="B9586B6DFF96FFC8FC7323833F27CEB6" bold="true" box="[990,1150,1478,1504]" italics="true" pageId="6" pageNumber="7">D. fornicatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF96FFC8FB2023833E37CEB6" author="Luederwaldt H." box="[1165,1390,1478,1504]" pageId="6" pageNumber="7" pagination="353 - 361" refId="ref30150" refString="Luederwaldt H. 1931. Tres especies novas de Pinotus, supplementos a Memoria " As especies Brasileiras do genero Pinotus ". na " Revista do Museu Paulista " Tomo XVI, 1929, P 603 SS. Revista do Museu Paulista 17: 353 - 361." type="journal article" year="1931">Luederwaldt, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C23AF3E2ECD1C" blockId="6.[189,1399,1372,1611]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF6C23AF3E2ECD1C" pageId="6" pageNumber="7">
|
||
– Cephalic carina transverse. Pronotum with smooth declivity, not modified anteriorly. Dense ocellate punctures covering the surface next to the eyes. Black to dark brown. Body length greater than
|
||
<quantity id="4CD41A9AFF96FFC8FF47206A3A19CD1C" box="[234,320,1583,1610]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="15.0">15 mm</quantity>
|
||
. (
|
||
<collectingCountry id="F33BF7EFFF96FFC8FEFE20753A89CD1D" box="[339,464,1584,1611]" name="Venezuela" pageId="6" pageNumber="7">Venezuela</collectingCountry>
|
||
)......................................................................
|
||
<taxonomicName id="4C2CCCFCFF96FFC8FC4F206A3E4CCD1C" ID-CoL="7VQMS" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[994,1301,1583,1610]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="henripittieri" status="sp. nov.">
|
||
<emphasis id="B9586B6DFF96FFC8FC4F206A3E4CCD1C" bold="true" box="[994,1301,1583,1610]" italics="true" pageId="6" pageNumber="7">Dichotomius henripittieri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF96FFC8FAB120753E2ECD1C" box="[1308,1399,1584,1610]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF96FFC8FF1020343E2ECC4E" pageId="6" pageNumber="7">
|
||
<paragraph id="8B93B77FFF96FFC8FF1020343E2FCD84" blockId="6.[189,1399,1649,1816]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF1020343E2FCD84" pageId="6" pageNumber="7">
|
||
8. Cephalic carina antero-posteriorly flattened, trituberculate, inclined anteriorly, apically expanded and emarginate medially. Anterior edge of the declivity medial area deeper and more convex than lateral ones (
|
||
<collectingCountry id="F33BF7EFFF96FFC8FE2C20F23AAECD87" box="[385,503,1719,1745]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF96FFC8FD8220F239F3CD84" box="[559,682,1719,1746]" name="Venezuela" pageId="6" pageNumber="7">Venezuela</collectingCountry>
|
||
) ....................................................
|
||
<taxonomicName id="4C2CCCFCFF96FFC8FB9820FD3E2FCD84" ID-CoL="35NBJ" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1077,1398,1719,1746]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="deyrollei">
|
||
<emphasis id="B9586B6DFF96FFC8FB9820FD3F98CD84" bold="true" box="[1077,1217,1719,1746]" italics="true" pageId="6" pageNumber="7">D. deyrollei</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF96FFC8FB7D20F23E37CD84" author="Harold E." box="[1232,1390,1719,1746]" pageId="6" pageNumber="7" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C209E3E2ECC4E" blockId="6.[189,1399,1649,1816]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF6C209E3E2ECC4E" pageId="6" pageNumber="7">– Cephalic carina conical, not antero-posteriorly flattened neither inclined anteriorly. Anterior edge of the pronotum modified, not convex, displaying undeveloped lobes ................................................. 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF96FFC8FF10217A3E2FCCB0" pageId="6" pageNumber="7">
|
||
<paragraph id="8B93B77FFF96FFC8FF10217A3E2ECCF6" blockId="6.[189,1399,1855,2023]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF10217A3E2ECCF6" pageId="6" pageNumber="7">
|
||
9. Pronotum with three undeveloped lobes on the anterior edge. Cephalic carina with lateral tubercles indistinct, medial tubercle with an acute apex, never emarginate.(Clypeus with a tooth or protuberance before the clypeogenal junction.) (
|
||
<collectingCountry id="F33BF7EFFF96FFC8FDD721C339A9CCF6" box="[634,752,1926,1952]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF96FFC8FC8521C338FACCF7" box="[808,931,1926,1953]" name="Venezuela" pageId="6" pageNumber="7">Venezuela</collectingCountry>
|
||
) .............
|
||
<taxonomicName id="4C2CCCFCFF96FFC8FBBF21C33E2ECCF6" ID-CoL="35NC3" authority="(Felsche, 1901)" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1042,1399,1925,1952]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="inachoides">
|
||
<emphasis id="B9586B6DFF96FFC8FBBF21C33FE1CCF6" bold="true" box="[1042,1208,1925,1952]" italics="true" pageId="6" pageNumber="7">D. inachoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF96FFC8FB6521C33E37CCF6" author="Felsche C." box="[1224,1390,1926,1952]" pageId="6" pageNumber="7" pagination="135 - 155" refId="ref29807" refString="Felsche C. 1901. Beschreibungen coprophager Scarabaeiden. Deutsche Entomologische Zeitschrift 2: 135 - 155." type="journal article" year="1901">Felsche, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF96FFC8FF6C21EC3E2FCCB0" blockId="6.[189,1399,1855,2023]" pageId="6" pageNumber="7">
|
||
<keyLead id="30DD144AFF96FFC8FF6C21EC3E2FCCB0" pageId="6" pageNumber="7">– Pronotum with two underdeveloped lobes on the anterior edge. Cephalic carina with defined lateral tubercles, medial tubercle emarginate at apex ................................................................................. 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF10274C3E2FCA86" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF10274C3E2ECA3E" blockId="7.[189,1399,265,464]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF10274C3E2ECA3E" pageId="7" pageNumber="8">
|
||
10. Clypeus not rounded. Head with anterior third surface subrugose. Pronotum as wide as the elytra, surface smooth, with ocellate punctation on the posterior margin. (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FC5A276E3F2DCA10" box="[1015,1140,299,326]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
|
||
)................................. ..................................................................................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FC6E270B3E2ECA3E" ID-CoL="35ND9" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[963,1399,333,360]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="validipilosus">
|
||
<emphasis id="B9586B6DFF97FFC9FC6E270B3F27CA3E" bold="true" box="[963,1150,333,360]" italics="true" pageId="7" pageNumber="8">D. validipilosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB20270B3E37CA3E" author="Luederwaldt H." box="[1165,1390,334,360]" pageId="7" pageNumber="8" pagination="353 - 361" refId="ref30150" refString="Luederwaldt H. 1931. Tres especies novas de Pinotus, supplementos a Memoria " As especies Brasileiras do genero Pinotus ". na " Revista do Museu Paulista " Tomo XVI, 1929, P 603 SS. Revista do Museu Paulista 17: 353 - 361." type="journal article" year="1931">Luederwaldt, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C27343E2FCA86" blockId="7.[189,1399,265,464]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C27343E2FCA86" pageId="7" pageNumber="8">
|
||
– Clypeus rounded. Head anterior third transversally rugose. Pronotum wider than the elytra, posterior margin without ocellate punctation, punctation sparse if present. (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FC4727D63F7ACAFB" box="[1002,1059,403,429]" name="Peru" pageId="7" pageNumber="8">Peru</collectingCountry>
|
||
)............................................ ................................................................................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FC1C27F03E2FCA86" ID-CoL="35ND2" authority="(Luederwaldt, 1935)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[945,1398,436,464]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="simplicicornis">
|
||
<emphasis id="B9586B6DFF97FFC9FC1C27F03F27CA99" bold="true" box="[945,1150,436,463]" italics="true" pageId="7" pageNumber="8">D. simplicicornis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB2027F03E37CA86" author="Luederwaldt H." box="[1165,1390,437,464]" pageId="7" pageNumber="8" pagination="334 - 342" refId="ref30199" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Coleoptera: Scarabaeidae). Revista Entomologica 5: 334 - 342." type="journal article" year="1935">Luederwaldt, 1935</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF1027B33E2FC921" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF1027B33E2FC964" blockId="7.[189,1398,501,631]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF1027B33E2FC964" pageId="7" pageNumber="8">
|
||
11. Setigerous punctures of the medial and lateral edges of the metasternum connected via the inner mesocoxal margins (
|
||
<figureCitation id="1317ABFAFF97FFC9FE70245D3965C964" box="[477,572,536,562]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="7" pageNumber="8">Fig. 2G</figureCitation>
|
||
). Pronotal anterior declivity with a medial subemargination ............ 12
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C247E3E2FC921" blockId="7.[189,1398,501,631]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C247E3E2FC921" pageId="7" pageNumber="8">
|
||
– Setigerous punctures of the medial and lateral edges of the metasternum not connected via the inner mesocoxal margins (
|
||
<figureCitation id="1317ABFAFF97FFC9FE7024193962C921" box="[477,571,604,631]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="7" pageNumber="8">Fig. 2H</figureCitation>
|
||
). Pronotal anterior declivity without medial subemargination.......... 14
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF1024D83E2FC835" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF1024D83E2FC9AB" blockId="7.[189,1399,669,868]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF1024D83E2FC9AB" pageId="7" pageNumber="8">
|
||
12. Male median tubercle of the cephalic carina thick, cylindrical and with a pronounced apical emargination. Metacoxae posterior edge bearing row of organized, shallow ocellate setigerous punctures which merge together (
|
||
<figureCitation id="1317ABFAFF97FFC9FDDE24A73996C9AA" box="[627,719,738,764]" captionStart="Fig" captionStartId="13.[189,232,1855,1881]" captionTargetBox="[236,1348,271,1823]" captionTargetId="figure-84@13.[233,1354,265,1824]" captionTargetPageId="13" captionText="Fig. 4. Dichotomius amplicollis (Harold, 1869). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus. D. Anterior view of the cephalic carina, the central tubercle cylindrical and apically emarginate.E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metacoxa, displaying a single row of organized ocellate setigerous punctures which merge together on the posterior margin." figureDoi="http://doi.org/10.5281/zenodo.4542115" httpUri="https://zenodo.org/record/4542115/files/figure.png" pageId="7" pageNumber="8">Fig. 4H</figureCitation>
|
||
) (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FD4A24A7381DC9AB" box="[743,836,738,765]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
|
||
) ...........................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FBB424A73E2FC9AB" ID-CoL="35NB2" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1049,1398,737,765]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="amplicollis">
|
||
<emphasis id="B9586B6DFF97FFC9FBB424A73F98C9AA" bold="true" box="[1049,1217,737,764]" italics="true" pageId="7" pageNumber="8">D. amplicollis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB7D24A73E37C9AB" author="Harold E." box="[1232,1390,738,765]" pageId="7" pageNumber="8" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C25403E2FC835" blockId="7.[189,1399,669,868]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C25403E2FC835" pageId="7" pageNumber="8">
|
||
– Male median tubercle of the cephalic carina not cylindrical, apex feebly emarginate or rounded. Metacoxae posterior edge bearing row of organized, ocellate setigerous punctures not merging together (
|
||
<figureCitation id="1317ABFAFF97FFC9FEF7250C3A9DC835" box="[346,452,841,867]" captionStart="Fig" captionStartId="34.[189,232,1893,1919]" captionTargetBox="[225,1361,274,1867]" captionTargetId="figure-76@34.[223,1364,265,1877]" captionTargetPageId="34" captionText="Fig. 11. Dichotomius inachoides (Felsche, 1901). A–B. Dorsal habitus. A. ♂. B. ♀. C. Frontal view of the head showing the male cephalic carina. D. Sternites smooth, lacking ocellate punctation. E. Female antero-lateral edge of the pronotum rugose. F. Lateral view of the aedeagus. G. Ventral view of the aedeagus. H–I. Lamella copulatrix." figureDoi="http://doi.org/10.5281/zenodo.4542129" httpUri="https://zenodo.org/record/4542129/files/figure.png" pageId="7" pageNumber="8">Figs 11E</figureCitation>
|
||
and
|
||
<figureCitation id="1317ABFAFF97FFC9FE50250C3977C835" box="[509,558,841,867]" captionStart="Fig" captionStartId="37.[189,232,1895,1921]" captionTargetBox="[212,1376,265,1860]" captionTargetId="figure-70@37.[207,1381,259,1866]" captionTargetPageId="37" captionText="Fig. 12. Dichotomius rafanunezi sp. nov. A–B. Dorsal habitus.A. Holotype, ♂ (CEMT). B. ♀. C. Lateral view of the aedeagus. D. Frontal view of the head. E. Dorsal view of the aedeagus. F–G. Lamella copulatrix. H. Ventral view of the metasternum; antero-medial edge near the interior margin of the mesocoxa rugose." figureDoi="http://doi.org/10.5281/zenodo.4542131" httpUri="https://zenodo.org/record/4542131/files/figure.png" pageId="7" pageNumber="8">12E</figureCitation>
|
||
) ............................................................................................................ 13
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF1025CF3E2ECFC3" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF1025CF3E2ECF5D" blockId="7.[189,1399,906,1173]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF1025CF3E2ECF5D" pageId="7" pageNumber="8">
|
||
13. Interocular space smooth with fine rounded punctures. Male median tubercle of the cephalic carina not emarginate apically (
|
||
<figureCitation id="1317ABFAFF97FFC9FDB125E939D5C890" box="[540,652,940,966]" captionStart="Fig" captionStartId="34.[189,232,1893,1919]" captionTargetBox="[225,1361,274,1867]" captionTargetId="figure-76@34.[223,1364,265,1877]" captionTargetPageId="34" captionText="Fig. 11. Dichotomius inachoides (Felsche, 1901). A–B. Dorsal habitus. A. ♂. B. ♀. C. Frontal view of the head showing the male cephalic carina. D. Sternites smooth, lacking ocellate punctation. E. Female antero-lateral edge of the pronotum rugose. F. Lateral view of the aedeagus. G. Ventral view of the aedeagus. H–I. Lamella copulatrix." figureDoi="http://doi.org/10.5281/zenodo.4542129" httpUri="https://zenodo.org/record/4542129/files/figure.png" pageId="7" pageNumber="8">Fig. 11D</figureCitation>
|
||
). Pronotum surface covered by fine punctures, margins with row of dense interrupted ocellate punctures. Female pronotum surface smooth with fine punctures (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FF5F25B43A16CF5A" box="[242,335,1009,1036]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
|
||
) .................................................................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FC1925B43E55CF5A" ID-CoL="7VQMV" authority="(Harold, 1869)" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[948,1292,1009,1036]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sagittarius" status="stat. nov.">
|
||
<emphasis id="B9586B6DFF97FFC9FC1925B43F0ECF5D" bold="true" box="[948,1111,1009,1035]" italics="true" pageId="7" pageNumber="8">D. sagittarius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FBCA25B43E5DCF5A" author="Harold E." box="[1127,1284,1009,1036]" pageId="7" pageNumber="8" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF97FFC9FAB925B43E2ECF5D" box="[1300,1399,1009,1035]" pageId="7" pageNumber="8">stat. rev.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C22513E2ECFC3" blockId="7.[189,1399,906,1173]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C22513E2ECFC3" pageId="7" pageNumber="8">
|
||
– Interocular space covered by shagreened microsculpture and flat punctation. Male median tubercle of the cephalic carina feebly emarginate apically. Pronotum surface covered by well-impressed simple punctures, margins with row of dense uninterrupted ocellate punctures. Female pronotum covered with dense and deeply impressed punctures (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FD55223F3824CFC2" box="[760,893,1146,1172]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33BF7EFFF97FFC9FC24223F3F51CFC2" box="[905,1032,1146,1172]" name="Guatemala" pageId="7" pageNumber="8">Guatemala</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF97FFC9FBEC223F3FC2CFC3" box="[1089,1179,1146,1173]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
|
||
)..
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FB18223E3E4CCFC2" ID-CoL="7VQMQ" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" box="[1205,1301,1146,1173]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="enioi" status="sp. nov.">
|
||
<emphasis id="B9586B6DFF97FFC9FB18223E3E4CCFC2" bold="true" box="[1205,1301,1146,1173]" italics="true" pageId="7" pageNumber="8">D. enioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26BD616FF97FFC9FAB1223E3E2ECFC3" box="[1308,1399,1147,1173]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF1022FE3E2FCE09" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF1022FE3E2FCE4C" blockId="7.[189,1400,1211,1375]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF1022FE3E2FCE4C" pageId="7" pageNumber="8">
|
||
14. Median tubercle of the cephalic carina apically emarginate. Lateral margin of the hypomera with multiple rows of setae (
|
||
<figureCitation id="1317ABFAFF97FFC9FDA52298393CCFA1" box="[520,613,1245,1271]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="7" pageNumber="8">Fig. 2E</figureCitation>
|
||
). Metacoxa with more than one row of punctures on the posterior margin (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FEE322BA3A9FCE4F" box="[334,454,1279,1305]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33BF7EFFF97FFC9FE7F23453976CE4C" box="[466,559,1280,1306]" name="Panama" pageId="7" pageNumber="8">Panamá</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF97FFC9FDCA22BA39BBCE4C" box="[615,738,1279,1306]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
|
||
)...............................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FBE523453E2FCE4C" ID-CoL="6CS36" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1096,1398,1279,1306]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="agenor">
|
||
<emphasis id="B9586B6DFF97FFC9FBE523453F98CE4C" bold="true" box="[1096,1217,1280,1306]" italics="true" pageId="7" pageNumber="8">D. agenor</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB7D22BA3E37CE4C" author="Harold E." box="[1232,1390,1279,1306]" pageId="7" pageNumber="8" pagination="124 - 144" refId="ref29859" refString="Harold E. 1869. Revision des especes qui rentrent dans le genre Pinotus Erichs. L'Abeille 6: 124 - 144." type="journal article" year="1869">Harold, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C23673E2FCE09" blockId="7.[189,1400,1211,1375]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C23673E2FCE09" pageId="7" pageNumber="8">– Median tubercle of the cephalic carina not apically emarginate. Lateral margin of the hypomera with single row of setae. Metacoxa with single row of punctures on the posterior margin .................... 15</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30D8AFDAFF97FFC9FF1023C03E2FCDE2" pageId="7" pageNumber="8">
|
||
<paragraph id="8B93B77FFF97FFC9FF1023C03E2FCD7F" blockId="7.[189,1399,1413,1717]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF1023C03E2FCD7F" pageId="7" pageNumber="8">
|
||
15. Dorsal and ventral surface black. Clypeus with weak central emargination. Anterior portion of the pronotum with a strong, abrupt declivity. Median tubercle of the cephalic carina approximately two and a half times as high as lateral tubercles. Punctures of the lateral edges of the metasternum separated by three times the puncture diameter (
|
||
<figureCitation id="1317ABFAFF97FFC9FCB223A9383ECD50" box="[799,871,1516,1542]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
|
||
). (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FC2923A938A3CD50" box="[900,1018,1516,1542]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33BF7EFFF97FFC9FB9E23A93FF7CD51" box="[1075,1198,1516,1543]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
|
||
)......................... ..............................................................................................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FBBA204A3E2FCD7F" ID-CoL="35ND8" authority="(Luederwaldt, 1923)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[1047,1398,1550,1577]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tristis">
|
||
<emphasis id="B9586B6DFF97FFC9FBBA204A3F27CD7E" bold="true" box="[1047,1150,1550,1577]" italics="true" pageId="7" pageNumber="8">D. tristis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB20204B3E37CD7F" author="Luederwaldt H." box="[1165,1390,1550,1577]" pageId="7" pageNumber="8" pagination="1 - 13" refId="ref30108" refString="Luederwaldt H. 1923. Neue Pinotus - Arten. Revista do Museu Paulista 14: 1 - 13." type="journal article" year="1923">Luederwaldt, 1923</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFF97FFC9FF6C20743E2FCDE2" blockId="7.[189,1399,1413,1717]" pageId="7" pageNumber="8">
|
||
<keyLead id="30DD144AFF97FFC9FF6C20743E2FCDE2" pageId="7" pageNumber="8">
|
||
– Dorsal and ventral surface dark brown. Clypeus with deep central emargination separating two obtuse teeth. Median tubercle of the cephalic carina twice as high as lateral tubercles. Punctures of the lateral edges of the metasternum separated by one puncture diameter (
|
||
<collectingCountry id="F33BF7EFFF97FFC9FF5F20DF3A03CDE3" box="[242,346,1690,1717]" name="Trinidad and Tobago" pageId="7" pageNumber="8">Trinidad</collectingCountry>
|
||
) ......................................................................................
|
||
<taxonomicName id="4C2CCCFCFF97FFC9FC4E20DF3E2FCDE2" ID-CoL="35ND7" authority="(Luederwaldt, 1923)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1923" box="[995,1398,1690,1716]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="triquetrus">
|
||
<emphasis id="B9586B6DFF97FFC9FC4E20DF3F27CDE2" bold="true" box="[995,1150,1690,1716]" italics="true" pageId="7" pageNumber="8">D. triquetrus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFF97FFC9FB2020DF3E37CDE2" author="Luederwaldt H." box="[1165,1390,1690,1716]" pageId="7" pageNumber="8" pagination="1 - 13" refId="ref30108" refString="Luederwaldt H. 1923. Neue Pinotus - Arten. Revista do Museu Paulista 14: 1 - 13." type="journal article" year="1923">Luederwaldt, 1923</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |