treatments-xml/data/85/54/87/855487ADFFFBFFA1FF7AF1321B7DD1E2.xml

<document id="2552B9370C26D2AD7DA386A1A5AF0097" ID-DOI="10.11646/zootaxa.3664.3.1" ID-GBIF-Dataset="0fbaac9f-afa4-4ddb-a656-7d97151f81af" ID-ISSN="1175-5326" ID-Zenodo-Dep="217551" ID-ZooBank="8B5606C6-5C88-49BF-A71F-ACEA8946D76F" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460617368255" checkinUser="plazi" docAuthor="Uyeno, Daisuke" docDate="2013" docId="855487ADFFFBFFA1FF7AF1321B7DD1E2" docLanguage="en" docName="zt03664p311.pdf" docOrigin="Zootaxa 3664 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cardiodectes shini Uyeno, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="309" masterDocId="796DFFD5FFFEFFA9FFEDF35E1F06D245" masterDocTitle="Two new species of Cardiodectes Wilson, 1917 (Copepoda: Siphonostomatoida: Pennellidae) from gobiid fishes (Actinopterygii: Perciformes) in the western Pacific Ocean" masterLastPageNumber="311" masterPageNumber="301" pageNumber="306" updateTime="1698332183538" updateUser="plazi">
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<mods:title id="9740B1AF21213EDA788E8EE3D5FBEF7E">Two new species of Cardiodectes Wilson, 1917 (Copepoda: Siphonostomatoida: Pennellidae) from gobiid fishes (Actinopterygii: Perciformes) in the western Pacific Ocean</mods:title>
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<mods:namePart id="91F1716CE0633D328E564A47F82A48D3">Uyeno, Daisuke</mods:namePart>
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<mods:title id="01282E2A600B88E2D43D8F15DCCF6650">Zootaxa</mods:title>
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<mods:date id="0FDB912C32FF43E96922E29EF7D7723A">2013</mods:date>
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<treatment id="855487ADFFFBFFA1FF7AF1321B7DD1E2" ID-DOI="http://doi.org/10.5281/zenodo.6149026" ID-GBIF-Taxon="119568713" ID-Zenodo-Dep="6149026" LSID="urn:lsid:plazi:treatment:855487ADFFFBFFA1FF7AF1321B7DD1E2" httpUri="http://treatment.plazi.org/id/855487ADFFFBFFA1FF7AF1321B7DD1E2" lastPageId="8" lastPageNumber="309" pageId="5" pageNumber="306">
<subSubSection id="45E76530FFFBFFACFF7AF1321E5AD0E3" pageId="5" pageNumber="306" type="nomenclature">
<paragraph id="0D4236BBFFFBFFACFF7AF1321EC6D0C0" blockId="5.[151,448,619,678]" box="[151,448,619,646]" pageId="5" pageNumber="306">
<heading id="560A81D7FFFBFFACFF7AF1321EC6D0C0" bold="true" box="[151,448,619,646]" fontSize="11" level="1" pageId="5" pageNumber="306" reason="1">
<emphasis id="3F89EAA9FFFBFFACFF7AF1321EC6D0C0" bold="true" box="[151,448,619,646]" pageId="5" pageNumber="306">
<taxonomicName id="CAFD4D38FFFBFFACFF7AF1321E70D0C3" box="[151,374,620,646]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFFBFFACFF7AF1321E70D0C3" bold="true" box="[151,374,620,646]" italics="true" pageId="5" pageNumber="306">Cardiodectes shini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24BA57D2FFFBFFACFE93F1351EC6D0C0" box="[382,448,619,645]" pageId="5" pageNumber="306" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0D4236BBFFFBFFACFF7AF1D01E5AD0E3" blockId="5.[151,448,619,678]" box="[151,348,654,678]" pageId="5" pageNumber="306">
(
<figureCitation id="95C62A3EFFFBFFACFF72F1D01FE4D0E3" box="[159,226,654,678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1858,1881]" captionTargetBox="[216,1365,193,1816]" captionTargetId="figure@2.[216,1366,193,1818]" captionTargetPageId="2" captionText="FIGURE 1. Postmetamorphic adult females of Cardiodectes bellwoodi n. sp. and C. shini n. sp. attached to their hosts. A, two specimens of Istigobius nigroocellatus (Günther) carrying the type series of C. bellwoodi n. sp.; B, a holotype (AMP. 90285) of C. bellwoodi n. sp. attached to the host; C, a specimen of Pleurosicya micheli Fourmanoir carrying a holotype (NSMT – Cr 22332) of C. shini n. sp.; D, a specimens of Eviota sebreei Jordan &amp; Seale carrying paratypes (NSMT – Cr 22335) of C. shini n. sp. Scale bars: A, 10 mm; B, C, D, 3 mm." httpUri="https://zenodo.org/record/217552/files/figure.png" pageId="5" pageNumber="306">Figs 1</figureCitation>
C, D, 4–5)
</paragraph>
</subSubSection>
<subSubSection id="45E76530FFFBFFACFF7AF1891A76D63F" pageId="5" pageNumber="306" type="materials_examined">
<paragraph id="0D4236BBFFFBFFACFF7AF1891A76D63F" blockId="5.[151,1436,726,1834]" pageId="5" pageNumber="306">
<emphasis id="3F89EAA9FFFBFFACFF7AF1891E42D0AA" bold="true" box="[151,324,726,751]" pageId="5" pageNumber="306">
<typeStatus id="D2468819FFFBFFACFF7AF1891FD4D0AA" box="[151,210,727,751]" pageId="5" pageNumber="306">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D2468819FFFBFFACFEA7F1881EB1D0AB" box="[330,439,726,750]" pageId="5" pageNumber="306" type="holotype">Holotype</typeStatus>
: female (NSMT–Cr 22332), ex 
<taxonomicName id="CAFD4D38FFFBFFACFCF5F1881B86D0AB" authority="Fourmanoir" authorityName="Fourmanoir" box="[792,1152,726,750]" class="Actinopterygii" family="Gobiidae" genus="Pleurosicya" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="micheli">
<emphasis id="3F89EAA9FFFBFFACFCF5F1881CF3D0AB" box="[792,1013,726,750]" italics="true" pageId="5" pageNumber="306">Pleurosicya micheli</emphasis>
Fourmanoir
</taxonomicName>
(
<taxonomicName id="CAFD4D38FFFBFFACFB63F1881A1AD0AB" box="[1166,1308,726,750]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="order">Perciformes</taxonomicName>
: 
<taxonomicName id="CAFD4D38FFFBFFACFAC8F1881A96D0AB" box="[1317,1424,726,750]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
), Oura Bay (
<geoCoordinate id="68C9507CFFFBFFACFEF9F1A41E71D157" box="[276,375,762,786]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.516666">26°31’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFE69F1A41EE2D157" box="[388,484,762,786]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="128.06667">128°4’E</geoCoordinate>
), Okinawa-jima Island, Ryukyu Islands, North Pacific Ocean, 
<collectingCountry id="75EA762BFFFBFFACFB49F1A51BE1D157" box="[1188,1255,763,786]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFB1EF1A41A2AD157" box="[1267,1324,762,786]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="306" unit="m" value="33.0">33 m</quantity>
depth, 
<date id="7943107BFFFBFFACFA6DF1A41E11D173" pageId="5" pageNumber="306" value="2012-04-29">29 April 2012</date>
, reg. D. Uyeno and S. Nishihira. 
<typeStatus id="D2468819FFFBFFACFD70F0411C08D172" box="[669,782,799,823]" pageId="5" pageNumber="306" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="1BFBFD32FFFBFFACFCF1F0401C8CD173" box="[796,906,798,822]" pageId="5" pageNumber="306" type="female">3 females</specimenCount>
(NSMT–Cr 22333), ex 
<taxonomicName id="CAFD4D38FFFBFFACFB4EF0411BEAD173" box="[1187,1260,799,822]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="3F89EAA9FFFBFFACFB4EF0411BEAD173" box="[1187,1260,799,822]" italics="true" pageId="5" pageNumber="306">Eviota</emphasis>
</taxonomicName>
sp., off Zatsun (
<geoCoordinate id="68C9507CFFFBFFACFF72F01C1E05D11F" box="[159,259,834,858]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.816668">26°49’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFEE3F01C1E7BD11F" box="[270,381,834,858]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="128.23334">128°14’E</geoCoordinate>
), Okinawa-jima Island, Ryukyu Islands, East 
<collectingCountry id="75EA762BFFFBFFACFC97F01C1CBBD11F" box="[890,957,834,858]" name="China" pageId="5" pageNumber="306">China</collectingCountry>
Sea, 
<collectingCountry id="75EA762BFFFBFFACFC14F01D1B3AD11F" box="[1017,1084,835,858]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFBABF01C1B78D11F" box="[1094,1150,834,858]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="306" unit="m" value="15.0">15 m</quantity>
depth, 
<date id="7943107BFFFBFFACFB3DF01C1A61D11F" box="[1232,1383,834,858]" pageId="5" pageNumber="306" value="2010-06-27">27 June 2010</date>
, reg. D. Uyeno; 
<specimenCount id="1BFBFD32FFFBFFACFEF1F0381E84D13B" box="[284,386,870,894]" pageId="5" pageNumber="306" type="female">1 female</specimenCount>
(RUMF–ZC– 2391), ex 
<emphasis id="3F89EAA9FFFBFFACFD41F0361C1DD13B" box="[684,795,870,895]" italics="true" pageId="5" pageNumber="306">P. m i c h e l i</emphasis>
, Oura Bay (
<geoCoordinate id="68C9507CFFFBFFACFC5AF0381B1DD13B" box="[951,1051,870,894]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.516666">26°31’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFBC6F0381B8DD13B" box="[1067,1163,870,894]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="128.06667">128°4’E</geoCoordinate>
), Okinawa-jima Island, Ryukyu Islands, North Pacific Ocean, 
<collectingCountry id="75EA762BFFFBFFACFDAAF0D51D8CD1E7" box="[583,650,907,930]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFD78F0D41DC9D1E7" box="[661,719,906,930]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="306" unit="m" value="31.0">31 m</quantity>
depth, 
<date id="7943107BFFFBFFACFCCFF0D41CC5D1E7" box="[802,963,906,930]" pageId="5" pageNumber="306" value="2012-04-17">17 April 2012</date>
, reg. D. Uyeno and S. Nishihira; 
<specimenCount id="1BFBFD32FFFBFFACFAD7F0D41A9AD1E7" box="[1338,1436,906,930]" pageId="5" pageNumber="306" type="female">1 female</specimenCount>
(NSMT–Cr 22334), ex 
<taxonomicName id="CAFD4D38FFFBFFACFE45F0EE1D9CD182" authority="Oura Bay" authorityName="Oura Bay" box="[424,666,942,967]" class="Actinopterygii" family="Gobiidae" genus="Pleurosicya" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="micheli">
<emphasis id="3F89EAA9FFFBFFACFE45F0EE1D1FD183" box="[424,537,942,967]" italics="true" pageId="5" pageNumber="306">P. micheli</emphasis>
, Oura Bay
</taxonomicName>
(
<geoCoordinate id="68C9507CFFFBFFACFD40F0F01C17D183" box="[685,785,942,966]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.516666">26°31’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFCF2F0F01C86D183" box="[799,896,942,966]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="128.06667">128°4’E</geoCoordinate>
), Okinawa-jima Island, Ryukyu Islands, North Pacific Ocean, 
<collectingCountry id="75EA762BFFFBFFACFEAFF08D1E83D1AF" box="[322,389,979,1002]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFE7DF08C1ECFD1AF" box="[400,457,978,1002]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="306" unit="m" value="30.0">30 m</quantity>
depth, 
<date id="7943107BFFFBFFACFDF1F08C1DBBD1AF" box="[540,701,978,1002]" pageId="5" pageNumber="306" value="2012-04-29">29 April 2012</date>
, reg. D. Uyeno and S. Nishihira; 
<specimenCount id="1BFBFD32FFFBFFACFBDEF08C1B92D1AF" box="[1075,1172,978,1002]" pageId="5" pageNumber="306" type="female">2 female</specimenCount>
(NSMT–Cr 22335), ex 
<taxonomicName id="CAFD4D38FFFBFFACFF7AF0A91E3AD64B" box="[151,316,1014,1038]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="species" species="sebreei">
<emphasis id="3F89EAA9FFFBFFACFF7AF0A91E3AD64B" box="[151,316,1014,1038]" italics="true" pageId="5" pageNumber="306">Eviota sebreei</emphasis>
</taxonomicName>
<collectingCountry id="75EA762BFFFBFFACFEAAF0A81E94D64B" box="[327,402,1014,1038]" name="Jordan" pageId="5" pageNumber="306">Jordan</collectingCountry>
&amp; Seale (
<taxonomicName id="CAFD4D38FFFBFFACFDFDF0A81D98D64B" box="[528,670,1014,1038]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="order">Perciformes</taxonomicName>
: 
<taxonomicName id="CAFD4D38FFFBFFACFD40F0A81C1ED64B" box="[685,792,1014,1038]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
), Oura Bay (
<geoCoordinate id="68C9507CFFFBFFACFC5BF0A81B1CD64B" box="[950,1050,1014,1038]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.516666">26°31’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFBC7F0A81B8DD64B" box="[1066,1163,1014,1038]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="128.06667">128°4’E</geoCoordinate>
), Okinawa-jima Island, Ryukyu Islands, North Pacific Ocean, 
<collectingCountry id="75EA762BFFFBFFACFDAAF7451D8CD677" box="[583,650,1051,1074]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFD78F7441DC9D677" box="[661,719,1050,1074]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="306" unit="m" value="28.0">28 m</quantity>
depth, 
<date id="7943107BFFFBFFACFCCFF7441CC5D677" box="[802,963,1050,1074]" pageId="5" pageNumber="306" value="2012-04-29">29 April 2012</date>
, reg. D. Uyeno and S. Nishihira; 
<specimenCount id="1BFBFD32FFFBFFACFAD7F7441A9AD677" box="[1338,1436,1050,1074]" pageId="5" pageNumber="306" type="female">1 female</specimenCount>
(RUMF–ZC– 2392), ex 
<taxonomicName id="CAFD4D38FFFBFFACFE50F7601D25D613" box="[445,547,1086,1110]" class="Actinopterygii" family="Gobiidae" genus="Priolepis" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="genus">
<emphasis id="3F89EAA9FFFBFFACFE50F7601D25D613" box="[445,547,1086,1110]" italics="true" pageId="5" pageNumber="306">Priolepis</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="CAFD4D38FFFBFFACFD85F7601DF0D613" box="[616,758,1086,1110]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="order">Perciformes</taxonomicName>
: 
<taxonomicName id="CAFD4D38FFFBFFACFCE5F7601C75D613" box="[776,883,1086,1110]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="306" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
), off Kyoda (
<geoCoordinate id="68C9507CFFFBFFACFBF0F7601B87D613" box="[1053,1153,1086,1110]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="26.533333">26°32’N</geoCoordinate>
, 
<geoCoordinate id="68C9507CFFFBFFACFB79F7601A05D613" box="[1172,1283,1086,1110]" direction="east" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="127.95">127°57’E</geoCoordinate>
), Nago Bay, Okinawa-jima Island, Ryukyu Islands, East 
<collectingCountry id="75EA762BFFFBFFACFD6EF73C1DC0D63F" box="[643,710,1122,1146]" name="China" pageId="5" pageNumber="306">China</collectingCountry>
Sea, 
<collectingCountry id="75EA762BFFFBFFACFCE9F73D1C41D63F" box="[772,839,1123,1146]" name="Japan" pageId="5" pageNumber="306">Japan</collectingCountry>
, 
<quantity id="CA059B5EFFFBFFACFCBFF73C1C8DD63F" box="[850,907,1122,1146]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="306" unit="m" value="33.0">33 m</quantity>
depth, 
<date id="7943107BFFFBFFACFC30F73C1B95D63F" box="[989,1171,1122,1147]" pageId="5" pageNumber="306" value="2012-08-16">16 August 2012</date>
, reg. N. Shirakawa.
</paragraph>
</subSubSection>
<subSubSection id="45E76530FFFBFFA1FF24F7D81B05D3C2" lastPageId="8" lastPageNumber="309" pageId="5" pageNumber="306" type="description">
<paragraph id="0D4236BBFFFBFFACFF24F7D81C91D73C" blockId="5.[151,1436,726,1834]" pageId="5" pageNumber="306">
<emphasis id="3F89EAA9FFFBFFACFF24F7D81DFAD6DA" bold="true" box="[201,764,1158,1183]" pageId="5" pageNumber="306">Description of postmetamorphic adult female.</emphasis>
Body (
<figureCitation id="95C62A3EFFFBFFACFCB8F7D81C9AD6DB" box="[853,924,1158,1182]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
A, B) 2705 long, comprising cephalothorax, neck and trunk. Cephalothorax (
<figureCitation id="95C62A3EFFFBFFACFDEFF7F41D40D687" box="[514,582,1194,1218]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
C, D, E) wider than long 725 × 917, bearing nodular and branching anterior processes on distal part of ventral surface and a pair of anterior digitiform lobes, as well as an expanded posterolateral pair of round lobes; anterior processes covering anterior half of cephalothorax (
<figureCitation id="95C62A3EFFFBFFACFB5CF7AC1BF2D74F" box="[1201,1268,1266,1290]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
A, B, C). Neck region (
<figureCitation id="95C62A3EFFFBFFACFF00F6471E36D774" box="[237,304,1305,1329]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
C, D, E) narrow, bearing pair of lobes, and bending through 90o. Trunk (
<figureCitation id="95C62A3EFFFBFFACFBBEF6471B90D774" box="[1107,1174,1305,1329]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
A, B) less than twice as long as wide 1900 × 980, oval with greatest width at posterior ¾ and with convex posterior margin. Egg sac spiral (
<figureCitation id="95C62A3EFFFBFFACFF72F63F1FE7D73C" box="[159,225,1377,1401]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1930]" captionTargetBox="[241,1344,230,1808]" captionTargetId="figure@6.[198,1379,195,1865]" captionTargetPageId="6" captionText="FIGURE 4. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx 2 = maxilla, p 1 = leg 1, p 2 = leg 2, p 3 = leg 3; E, same, lateral, a 1 = antennule, a 2 = antenna. Scale bars: A, B, 1000 μm; C, D, 500 μm; E, 400 μm." httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="5" pageNumber="306">Fig. 4</figureCitation>
A) uniseriate, originating at posterior lateral genital apertures.
</paragraph>
<paragraph id="0D4236BBFFFBFFACFF24F6D81B1ED4A4" blockId="5.[151,1436,726,1834]" pageId="5" pageNumber="306">
Rostrum, antennules, and antennae situated closely on anterodorsal surface of cephalothorax (
<figureCitation id="95C62A3EFFFBFFACFAC3F6DB1A71D7D8" box="[1326,1399,1413,1437]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
A). Rostrum (
<figureCitation id="95C62A3EFFFBFFACFEE6F6F71E56D784" box="[267,336,1449,1473]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
A) hemispherical, situated between bases of antennules. Antennule (
<figureCitation id="95C62A3EFFFBFFACFB83F6F71BB2D784" box="[1134,1204,1449,1473]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
A, C) unsegmented, bearing 8 setae mainly on anterior margin; distal tip bearing 8 setae, 2 bifurcate setae, and 1 aesthetasc. Antenna (
<figureCitation id="95C62A3EFFFBFFACFF72F6AF1FE5D44C" box="[159,227,1521,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
A, D) 3-segmented, chelate, typical pennellid in form; proximal segment with highly sclerotized ridge on surface; middle segment bearing inner medial pointed projection; terminal claw with 1 small basal seta on posterior surface. Mouth tube, maxillule, and maxilla located on anterior part of ventral surface of cephalothorax (
<figureCitation id="95C62A3EFFFBFFACFADBF5671A7CD414" box="[1334,1402,1593,1617]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
B). Maxillule (
<figureCitation id="95C62A3EFFFBFFACFEFBF5031E5CD430" box="[278,346,1629,1653]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
E) in form of bilobate knob, located lateral to base of mouth tube; each knob bearing 2 and 1 simple blunt processes. Maxilla (
<figureCitation id="95C62A3EFFFBFFACFDFCF5DF1D53D4DC" box="[529,597,1665,1689]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
B, F) 2-segmented; proximal segment with pointed anteromedial process; terminal segment indistinctly 2-segmented by constriction, covered with fine spinules at middle. Conspicuous bilobed process present between bases of maxillae (
<figureCitation id="95C62A3EFFFBFFACFD36F5971C18D4A4" box="[731,798,1737,1761]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
B). Maxilliped absent.
</paragraph>
<paragraph id="0D4236BBFFFBFFACFF24F5B31B6CD56C" blockId="5.[151,1436,726,1834]" pageId="5" pageNumber="306">
Both legs 1 and 2 (
<figureCitation id="95C62A3EFFFBFFACFE4BF5B31EEFD540" box="[422,489,1773,1797]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
G, H) biramous, present centrally on cephalothorax (
<figureCitation id="95C62A3EFFFBFFACFBAEF5B31B81D540" box="[1091,1159,1773,1797]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">Fig. 5</figureCitation>
D, E). Leg 3 (
<figureCitation id="95C62A3EFFFBFFACFAC7F5B31A69D540" box="[1322,1391,1773,1797]" captionStart="FIGURE 5" captionStartId="7.[151,250,1876,1899]" captionTargetBox="[204,1380,217,1856]" captionTargetId="figure@7.[204,1385,193,1863]" captionTargetPageId="7" captionText="FIGURE 5. Cardiodectes shini n. sp., postmetamorphic adult female, holotype, NSMT – Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50 μm; C, D, J, 10 μm; E – I, 20 μm." httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="5" pageNumber="306">
Fig. 
<date id="7943107BFFFBFFACFA8DF5B31A69D540" box="[1376,1391,1773,1797]" pageId="5" pageNumber="306">5</date>
</figureCitation>
I, J) uniramous, situated behind lobe on neck. Armature formula of all three legs as follows:
</paragraph>
<paragraph id="0D4236BBFFFBFFACFE7CF41B1B8FD587" pageId="5" pageNumber="306">
<table id="7FFDC41BFFFB0056FF71F41B1B9AD587" box="[156,1180,1861,1986]" colsContinueFrom="1.[159,1178,1600,1725]" gridcols="4" gridrows="4" pageId="5" pageNumber="306">
<tr id="B3CD34F9FFFB0056FF71F41B1B9AD51E" box="[156,1180,1861,1883]" gridrow="0" pageId="5" pageNumber="306" rowspan-0="1">
<th id="F01C5D85FFFB0056FE7CF41B1EE8D51E" box="[401,494,1861,1883]" gridcol="1" gridrow="0" pageId="5" pageNumber="306">Protopod</th>
<th id="F01C5D85FFFB0056FD3FF41B1C26D51E" box="[722,800,1861,1883]" gridcol="2" gridrow="0" pageId="5" pageNumber="306">Exopod</th>
<th id="F01C5D85FFFB0056FBACF41B1B9AD51E" box="[1089,1180,1861,1883]" gridcol="3" gridrow="0" pageId="5" pageNumber="306">Endopod</th>
</tr>
<tr id="B3CD34F9FFFB0056FF71F4311B9AD5C1" box="[156,1180,1903,1924]" gridrow="1" pageId="5" pageNumber="306">
<th id="F01C5D85FFFB0056FF71F4311FD1D5C1" box="[156,215,1903,1924]" gridcol="0" gridrow="1" pageId="5" pageNumber="306">Leg 1</th>
<td id="F01C5D85FFFB0056FE7CF4311EE8D5C1" box="[401,494,1903,1924]" gridcol="1" gridrow="1" pageId="5" pageNumber="306">1–1</td>
<td id="F01C5D85FFFB0056FD3FF4311C26D5C1" box="[722,800,1903,1924]" gridcol="2" gridrow="1" pageId="5" pageNumber="306">1–1; 7</td>
<td id="F01C5D85FFFB0056FBACF4311B9AD5C1" box="[1089,1180,1903,1924]" gridcol="3" gridrow="1" pageId="5" pageNumber="306">0–0; 7</td>
</tr>
<tr id="B3CD34F9FFFB0056FF71F4D01B9AD5E6" box="[156,1180,1934,1955]" gridrow="2" pageId="5" pageNumber="306">
<th id="F01C5D85FFFB0056FF71F4D01FD1D5E6" box="[156,215,1934,1955]" gridcol="0" gridrow="2" pageId="5" pageNumber="306">Leg 2</th>
<td id="F01C5D85FFFB0056FE7CF4D01EE8D5E6" box="[401,494,1934,1955]" gridcol="1" gridrow="2" pageId="5" pageNumber="306">1–0</td>
<td id="F01C5D85FFFB0056FD3FF4D01C26D5E6" box="[722,800,1934,1955]" gridcol="2" gridrow="2" pageId="5" pageNumber="306">1–1; 7</td>
<td id="F01C5D85FFFB0056FBACF4D01B9AD5E6" box="[1089,1180,1934,1955]" gridcol="3" gridrow="2" pageId="5" pageNumber="306">0–0; 7</td>
</tr>
<tr id="B3CD34F9FFFB0056FF71F4F21B9AD587" box="[156,1180,1964,1986]" gridrow="3" pageId="5" pageNumber="306">
<th id="F01C5D85FFFB0056FF71F4F21FD1D587" box="[156,215,1964,1986]" gridcol="0" gridrow="3" pageId="5" pageNumber="306">Leg 3</th>
<td id="F01C5D85FFFB0056FE7CF4F21EE8D587" box="[401,494,1964,1986]" gridcol="1" gridrow="3" pageId="5" pageNumber="306">0–0</td>
<td id="F01C5D85FFFB0056FD3FF4F21C26D587" box="[722,800,1964,1986]" gridcol="2" gridrow="3" pageId="5" pageNumber="306">0–0; 6</td>
<td id="F01C5D85FFFB0056FBACF4F21B9AD587" box="[1089,1180,1964,1986]" gridcol="3" gridrow="3" pageId="5" pageNumber="306">Absent</td>
</tr>
</table>
</paragraph>
<caption id="59826633FFF8FFAFFF7AF42D1EF2D5A6" httpUri="https://zenodo.org/record/217555/files/figure.png" pageId="6" pageNumber="307" targetBox="[241,1344,230,1808]" targetPageId="6">
<paragraph id="0D4236BBFFF8FFAFFF7AF42D1EF2D5A6" blockId="6.[151,1436,1907,2021]" pageId="6" pageNumber="307">
<emphasis id="3F89EAA9FFF8FFAFFF7AF42D1E14D5CC" bold="true" box="[151,274,1907,1930]" pageId="6" pageNumber="307">FIGURE 4.</emphasis>
<taxonomicName id="CAFD4D38FFF8FFAFFEF5F42A1ED0D5CF" box="[280,470,1908,1930]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF8FFAFFEF5F42A1ED0D5CF" box="[280,470,1908,1930]" italics="true" pageId="6" pageNumber="307">Cardiodectes shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF8FFAFFE31F42A1D13D5CC" bold="true" box="[476,533,1908,1929]" pageId="6" pageNumber="307">
<taxonomicNameLabel id="24BA57D2FFF8FFAFFE31F42A1D13D5CC" box="[476,533,1908,1929]" pageId="6" pageNumber="307" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, postmetamorphic adult female, holotype, NSMT–Cr 22332. A, habitus, dorsal; B, same, lateral; C, cephalothorax and neck region, ventral; D, cephalothorax and neck region with digitiform processes removed, ventral, mx2 = maxilla, p1 = leg 1, p2 = leg 2, p3 = leg 3; E, same, lateral, a1 = antennule, a2 = antenna. Scale bars: A, B, 1000μm; C, D, 500μm; E, 400μm.
</paragraph>
</caption>
<caption id="59826633FFF9FFAEFF7AF40A1E0CD5B5" httpUri="https://zenodo.org/record/217556/files/figure.png" pageId="7" pageNumber="308" targetBox="[204,1380,217,1856]" targetPageId="7">
<paragraph id="0D4236BBFFF9FFAEFF7AF40A1E0CD5B5" blockId="7.[151,1436,1876,2034]" pageId="7" pageNumber="308">
<emphasis id="3F89EAA9FFF9FFAEFF7AF40A1E1FD52F" bold="true" box="[151,281,1876,1899]" pageId="7" pageNumber="308">FIGURE 5.</emphasis>
<taxonomicName id="CAFD4D38FFF9FFAEFEC8F40B1EEFD52E" box="[293,489,1877,1899]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF9FFAEFEC8F40B1EEFD52E" box="[293,489,1877,1899]" italics="true" pageId="7" pageNumber="308">Cardiodectes shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF9FFAEFE1BF40B1D33D52F" bold="true" box="[502,565,1877,1898]" pageId="7" pageNumber="308">
<taxonomicNameLabel id="24BA57D2FFF9FFAEFE1BF40B1D33D52F" box="[502,565,1877,1898]" pageId="7" pageNumber="308" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, postmetamorphic adult female, holotype, NSMT–Cr 22332. A, anterior part of cephalothorax with antennule, antennae and rostrum; B, oral cone, maxillae and bilobed process; C, right antennule, anterior; D, right antenna, anterior; E, left maxillule; F, right maxilla, anterior; G, left leg 1 with intercoxal sclerite; H, right leg 2 with intercoxal sclerite; I, right leg 3 protopod with intercoxal sclerite; J, right leg 3 exopod. Scale bars: A, B, 50μm; C, D, J, 10μm; E–I, 20μm.
</paragraph>
</caption>
<paragraph id="0D4236BBFFF6FFA1FF24F3C61C53D2EA" blockId="8.[151,1436,151,935]" box="[201,853,151,176]" pageId="8" pageNumber="309">Leg 3 bearing protopod separated from intercoxal sclerite.</paragraph>
<paragraph id="0D4236BBFFF6FFA1FF24F3E51EA0D37A" blockId="8.[151,1436,151,935]" pageId="8" pageNumber="309">
<emphasis id="3F89EAA9FFF6FFA1FF24F3E51D6DD291" bold="true" box="[201,619,187,212]" pageId="8" pageNumber="309">Variability of female morphology.</emphasis>
The morphology of the female 
<typeStatus id="D2468819FFF6FFA1FC1DF3E31B5DD290" box="[1008,1115,189,213]" pageId="8" pageNumber="309" type="paratype">paratypes</typeStatus>
is as in the 
<typeStatus id="D2468819FFF6FFA1FB16F3E21A67D291" box="[1275,1377,188,212]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
. The measurements of the 
<typeStatus id="D2468819FFF6FFA1FE68F3BE1EB3D2BD" box="[389,437,224,248]" pageId="8" pageNumber="309">type</typeStatus>
series (n = 9) are as follows: body length 
<date id="7943107BFFF6FFA1FC66F3811B0CD2B2" box="[907,1034,223,247]" pageId="8" pageNumber="309" value="2038" valueMax="2767">2038–2767</date>
(2494 ± 253); cephalothorax length 495–840 (643 ± 98); cephalothorax width 616–957 (808 ± 115); trunk length 
<date id="7943107BFFF6FFA1FC1BF25A1B72D359" box="[1014,1140,260,284]" pageId="8" pageNumber="309" value="1462" valueMax="1986">1462–1986</date>
(1813 ± 199); trunk width 870–1250 (1050 ± 116).
</paragraph>
<paragraph id="0D4236BBFFF6FFA1FF24F2151B05D3C2" blockId="8.[151,1436,151,935]" pageId="8" pageNumber="309">
<emphasis id="3F89EAA9FFF6FFA1FF24F2151E94D321" bold="true" box="[201,402,331,356]" pageId="8" pageNumber="309">Attachment site.</emphasis>
The cephalothorax and neck region of the copepod were embedded in the host’s head musculature, while its trunk and egg sacs stick out into the water (
<figureCitation id="95C62A3EFFF6FFA1FC96F2311CBCD3C2" box="[891,954,367,391]" captionStart="FIGURE 1" captionStartId="2.[151,250,1858,1881]" captionTargetBox="[216,1365,193,1816]" captionTargetId="figure@2.[216,1366,193,1818]" captionTargetPageId="2" captionText="FIGURE 1. Postmetamorphic adult females of Cardiodectes bellwoodi n. sp. and C. shini n. sp. attached to their hosts. A, two specimens of Istigobius nigroocellatus (Günther) carrying the type series of C. bellwoodi n. sp.; B, a holotype (AMP. 90285) of C. bellwoodi n. sp. attached to the host; C, a specimen of Pleurosicya micheli Fourmanoir carrying a holotype (NSMT – Cr 22332) of C. shini n. sp.; D, a specimens of Eviota sebreei Jordan &amp; Seale carrying paratypes (NSMT – Cr 22335) of C. shini n. sp. Scale bars: A, 10 mm; B, C, D, 3 mm." httpUri="https://zenodo.org/record/217552/files/figure.png" pageId="8" pageNumber="309">Fig. 1</figureCitation>
C, D).
</paragraph>
</subSubSection>
<subSubSection id="45E76530FFF6FFA1FF24F2CA1BEDD17F" pageId="8" pageNumber="309" type="discussion">
<paragraph id="0D4236BBFFF6FFA1FF24F2CA1BEDD17F" blockId="8.[151,1436,151,935]" pageId="8" pageNumber="309">
<emphasis id="3F89EAA9FFF6FFA1FF24F2CA1E3BD3E9" bold="true" box="[201,317,404,428]" pageId="8" pageNumber="309">Remarks.</emphasis>
As 
<taxonomicName id="CAFD4D38FFF6FFA1FE84F2CB1EFED3E9" box="[361,504,404,428]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bellwoodi" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FE84F2CB1EFED3E9" box="[361,504,404,428]" italics="true" pageId="8" pageNumber="309">C. bellwoodi</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FE13F2CA1D3BD3E9" bold="true" box="[510,573,404,428]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FE13F2CA1D3BD3E9" box="[510,573,404,428]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="CAFD4D38FFF6FFA1FDA6F2CB1DA4D3E9" box="[587,674,404,428]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FDA6F2CB1D58D3E9" box="[587,606,405,428]" italics="true" pageId="8" pageNumber="309">C</emphasis>
. 
<emphasis id="3F89EAA9FFF6FFA1FD86F2CA1DA4D3E9" box="[619,674,404,428]" italics="true" pageId="8" pageNumber="309">shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FD45F2CA1DE1D3E9" bold="true" box="[680,743,404,428]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FD45F2CA1DE1D3E9" box="[680,743,404,428]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is assigned to the ‘rubosus’ group because of the absence of a defined abdomen (see Izawa 1970; Bellwood 1981). 
<taxonomicName id="CAFD4D38FFF6FFA1FCECF2E61CD0D395" box="[769,982,440,464]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FCECF2E61CD0D395" box="[769,982,440,464]" italics="true" pageId="8" pageNumber="309">Cardiodectes shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FC0FF2E61B22D395" bold="true" box="[994,1060,440,464]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FC0FF2E61B22D395" box="[994,1060,440,464]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="CAFD4D38FFF6FFA1FB29F2E61A77D395" box="[1220,1393,440,464]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="hardenbergi">
<emphasis id="3F89EAA9FFF6FFA1FB29F2E61A77D395" box="[1220,1393,440,464]" italics="true" pageId="8" pageNumber="309">C. hardenbergi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CAFD4D38FFF6FFA1FA6FF2E61FF5D3B1" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="krishnai">
<emphasis id="3F89EAA9FFF6FFA1FA6FF2E61FF5D3B1" italics="true" pageId="8" pageNumber="309">C. krishnai</emphasis>
</taxonomicName>
, 
<taxonomicName id="CAFD4D38FFF6FFA1FEEDF2831ED0D3B1" box="[256,470,476,500]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="rotundicaudatus">
<emphasis id="3F89EAA9FFF6FFA1FEEDF2831ED0D3B1" box="[256,470,476,500]" italics="true" pageId="8" pageNumber="309">C. rotundicaudatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CAFD4D38FFF6FFA1FDF9F2831D8BD3B1" box="[532,653,476,500]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="rubosus">
<emphasis id="3F89EAA9FFF6FFA1FDF9F2831D8BD3B1" box="[532,653,476,500]" italics="true" pageId="8" pageNumber="309">C. rubosus</emphasis>
</taxonomicName>
by having a trunk less than twice as long as wide (vs. more than twice as long as wide, Leigh-Sharpe 1934; Markevich 1936; Sebastian 1968; Izawa 1970; Bellwood 1981). Of the remaining five species, 
<taxonomicName id="CAFD4D38FFF6FFA1FE46F17B1D0AD079" box="[427,524,549,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="3F89EAA9FFF6FFA1FE46F17B1D0AD079" box="[427,524,549,572]" italics="true" pageId="8" pageNumber="309">C. asper</emphasis>
</taxonomicName>
, 
<taxonomicName id="CAFD4D38FFF6FFA1FDF3F17B1DB6D079" box="[542,688,548,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bellwoodi">
<emphasis id="3F89EAA9FFF6FFA1FDF3F17B1DB6D079" box="[542,688,548,572]" italics="true" pageId="8" pageNumber="309">C. bellwoodi</emphasis>
</taxonomicName>
n. sp, 
<taxonomicName id="CAFD4D38FFF6FFA1FCE8F17B1C95D079" box="[773,915,548,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bertrandi">
<emphasis id="3F89EAA9FFF6FFA1FCE8F17B1C95D079" box="[773,915,548,572]" italics="true" pageId="8" pageNumber="309">C. bertrandi</emphasis>
</taxonomicName>
, 
<taxonomicName id="CAFD4D38FFF6FFA1FC49F17B1B28D079" box="[932,1070,548,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="boxshalli">
<emphasis id="3F89EAA9FFF6FFA1FC49F17B1B28D079" box="[932,1070,548,572]" italics="true" pageId="8" pageNumber="309">C. boxshalli</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CAFD4D38FFF6FFA1FB99F17B1BEAD079" box="[1140,1260,548,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="spiralis">
<emphasis id="3F89EAA9FFF6FFA1FB99F17B1BEAD079" box="[1140,1260,548,572]" italics="true" pageId="8" pageNumber="309">C. spiralis</emphasis>
</taxonomicName>
, only 
<taxonomicName id="CAFD4D38FFF6FFA1FAD7F17B1A9AD079" box="[1338,1436,549,572]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="3F89EAA9FFF6FFA1FAD7F17B1A9AD079" box="[1338,1436,549,572]" italics="true" pageId="8" pageNumber="309">C. asper</emphasis>
</taxonomicName>
shares the following characters with 
<taxonomicName id="CAFD4D38FFF6FFA1FDD7F1151D95D026" box="[570,659,587,611]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FDD7F1151D4BD027" box="[570,589,587,610]" italics="true" pageId="8" pageNumber="309">C</emphasis>
. 
<emphasis id="3F89EAA9FFF6FFA1FDB1F1151D95D026" box="[604,659,587,611]" italics="true" pageId="8" pageNumber="309">shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FD71F1151DDAD026" bold="true" box="[668,732,587,611]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FD71F1151DDAD026" box="[668,732,587,611]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: the neck region bends through 90o in the middle and bears a well-developed pair of lobes, and leg 3 is located on the posterior part of the base of the neck lobes (Bellwood 1981; Uyeno &amp; Nagasawa 2010). These two species resemble each other, but 
<taxonomicName id="CAFD4D38FFF6FFA1FBE1F1CD1B62D0EE" box="[1036,1124,659,683]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FBE1F1CD1B19D0EF" box="[1036,1055,659,682]" italics="true" pageId="8" pageNumber="309">C</emphasis>
. 
<emphasis id="3F89EAA9FFF6FFA1FBC0F1CD1B62D0EE" box="[1069,1124,659,683]" italics="true" pageId="8" pageNumber="309">shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FB81F1CD1BAAD0EE" bold="true" box="[1132,1196,659,683]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FB81F1CD1BAAD0EE" box="[1132,1196,659,683]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinguishable by the presence of the well-developed, bilobed process between the bases of the maxillae and by the cephalothorax bearing two pairs of lobes (vs. without the well-developed bilobed process and the cephalothorax with 3 pairs of lobes (see Uyeno &amp; Nagasawa 2010)). In addition, legs 1 and 2 of 
<taxonomicName id="CAFD4D38FFF6FFA1FC71F05E1CF0D152" box="[924,1014,767,791]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FC71F05E1CA9D152" box="[924,943,768,791]" italics="true" pageId="8" pageNumber="309">C</emphasis>
. 
<emphasis id="3F89EAA9FFF6FFA1FC52F1A11CF0D152" box="[959,1014,767,791]" italics="true" pageId="8" pageNumber="309">shini</emphasis>
</taxonomicName>
<emphasis id="3F89EAA9FFF6FFA1FC13F1A11B39D152" bold="true" box="[1022,1087,767,791]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="24BA57D2FFF6FFA1FC13F1A11B39D152" box="[1022,1087,767,791]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
have protopods which are not separated from their intercoxal sclerites (vs. separated in 
<taxonomicName id="CAFD4D38FFF6FFA1FCFAF07D1C73D17E" box="[791,885,803,827]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="3F89EAA9FFF6FFA1FCFAF07D1C2CD17F" box="[791,810,803,826]" italics="true" pageId="8" pageNumber="309">C</emphasis>
. 
<emphasis id="3F89EAA9FFF6FFA1FCDAF07A1C73D17E" box="[823,885,804,827]" italics="true" pageId="8" pageNumber="309">asper</emphasis>
</taxonomicName>
(see Uyeno &amp; Nagasawa 2010)).
</paragraph>
</subSubSection>
<subSubSection id="45E76530FFF6FFA1FF24F0191B7DD1E2" pageId="8" pageNumber="309" type="etymology">
<paragraph id="0D4236BBFFF6FFA1FF24F0191BACD1C7" blockId="8.[151,1436,151,935]" pageId="8" pageNumber="309">
<emphasis id="3F89EAA9FFF6FFA1FF24F0191E56D11A" bold="true" box="[201,336,839,863]" pageId="8" pageNumber="309">Etymology.</emphasis>
The specific name of the new species, 
<taxonomicName id="CAFD4D38FFF6FFA1FCC9F0191C5CD11A" box="[804,858,839,863]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini">
<emphasis id="3F89EAA9FFF6FFA1FCC9F0191C5CD11A" box="[804,858,839,863]" italics="true" pageId="8" pageNumber="309">shini</emphasis>
</taxonomicName>
, is after 
<collectingCountry id="75EA762BFFF6FFA1FC2AF0191CFDD11A" box="[967,1019,839,863]" name="Morocco" pageId="8" pageNumber="309">Shin</collectingCountry>
“Snufkin” Nishihira, a President of Diving Team Snuck Snufkin, who is the best explorer of the Oura Bay, Okinawa-jima Island.
</paragraph>
<paragraph id="0D4236BBFFF6FFA1FF24F0D11B7DD1E2" blockId="8.[151,1436,151,935]" box="[201,1147,910,935]" pageId="8" pageNumber="309">
<emphasis id="3F89EAA9FFF6FFA1FF24F0D11CAED1E2" bold="true" box="[201,936,910,935]" pageId="8" pageNumber="309">
Newly established Japanese name for 
<taxonomicName id="CAFD4D38FFF6FFA1FD61F0D01C65D1E2" box="[652,867,910,935]" class="Maxillopoda" family="Pennellidae" genus="Cardiodectes" kingdom="Animalia" order="Siphonostomatoida" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="shini" status="sp. nov.">
<emphasis id="3F89EAA9FFF6FFA1FD61F0D01C65D1E2" bold="true" box="[652,867,910,935]" italics="true" pageId="8" pageNumber="309">Cardiodectes shini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24BA57D2FFF6FFA1FC84F0D11CAED1E2" box="[873,936,911,935]" pageId="8" pageNumber="309" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Shin-no-kanzashi.
</paragraph>
</subSubSection>
</treatment>
</document>