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<document id="A7682605C55239DA2B9F01849C697CF4" ID-DOI="10.5281/zenodo.174843" ID-GBIF-Dataset="45bfe6e3-4db8-41a7-822f-be069bb47639" ID-ISSN="1175-5326" ID-Zenodo-Dep="174843" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790107098" checkinUser="plazi" docAuthor="Bal, Durmuş Ali" docDate="2006" docId="039B8798FFA8FFF9FED75E93FEDB10DB" docLanguage="en" docName="zt01368p040.pdf" docOrigin="Zootaxa 1368" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Uroobovella anatolica Bal, 2006, sp. nov." docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFA2FFE0FFA5FFEAFFDF5B2DFFA61415" masterDocTitle="New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae)" masterLastPageNumber="40" masterPageNumber="19" pageNumber="32" updateTime="1698219151320" updateUser="plazi">
<mods:mods id="CF4AE6FD050AB9400C1806AF24B12295" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B4DDA7BE1C41A74EA567B46E9A5BC2D">
<mods:title id="9F7C202D47A5D79984ABB4226D4E2FFF">New species of mites in the genera Uropoda, Crinitodiscus and Uroobovella from Turkey (Acari: Mesostigmata: Uropodidae, Urodinychidae)</mods:title>
</mods:titleInfo>
<mods:name id="441D9173455AEB6BCB231195A468D527" type="personal">
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</mods:role>
<mods:namePart id="7E2FF6BE472E6A4A4CE02E8EFA5B5BDE">Bal, Durmuş Ali</mods:namePart>
</mods:name>
<mods:typeOfResource id="9E6A946D89D82594CF0928EF645EC895">text</mods:typeOfResource>
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<mods:title id="17C073D07548AB347FCED553198DFAD9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="ED824FFB340DE1E82AABF6419E80A1C9">
<mods:date id="7B3585E862C02464306C2D9CA912A52F">2006</mods:date>
<mods:detail id="8C401AB9F506F29EA7706761575F2B49" type="volume">
<mods:number id="1A2CDB84B0062DF6BDDA2B8293EC6534">1368</mods:number>
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</mods:extent>
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<mods:classification id="DC0FE66B964103D93533645AB75A9EDB">journal article</mods:classification>
<mods:identifier id="7A2E9181CA0CD343D74415D39CA5F0C4" type="DOI">10.5281/zenodo.174843</mods:identifier>
<mods:identifier id="7B8B94029C7FBCEC3C097CE3430255BD" type="GBIF-Dataset">45bfe6e3-4db8-41a7-822f-be069bb47639</mods:identifier>
<mods:identifier id="26784F0CABB10822457FF09E23CADFF6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6036A61005CD2FB2050BBEFF4D61E763" type="Zenodo-Dep">174843</mods:identifier>
</mods:mods>
<treatment id="039B8798FFA8FFF9FED75E93FEDB10DB" ID-DOI="http://doi.org/10.5281/zenodo.6253486" ID-GBIF-Taxon="119350214" ID-Zenodo-Dep="6253486" LSID="urn:lsid:plazi:treatment:039B8798FFA8FFF9FED75E93FEDB10DB" httpUri="http://treatment.plazi.org/id/039B8798FFA8FFF9FED75E93FEDB10DB" lastPageId="19" lastPageNumber="38" pageId="13" pageNumber="32">
<subSubSection id="C3286505FFA8FFE7FED75E93FDDF11CD" box="[264,633,1470,1496]" pageId="13" pageNumber="32" type="nomenclature">
<paragraph id="8B8D368EFFA8FFE7FED75E93FDDF11CD" blockId="13.[264,633,1470,1496]" box="[264,633,1470,1496]" pageId="13" pageNumber="32">
<heading id="D0C581E2FFA8FFE7FED75E93FDDF11CD" bold="true" box="[264,633,1470,1496]" fontSize="11" level="1" pageId="13" pageNumber="32" reason="1">
<emphasis id="B946EA9CFFA8FFE7FED75E93FDDF11CD" bold="true" box="[264,633,1470,1496]" pageId="13" pageNumber="32">
<taxonomicName id="4C324D0DFFA8FFE7FED75E93FDB511CD" box="[264,531,1470,1496]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFA8FFE7FED75E93FDB511CD" bold="true" box="[264,531,1470,1496]" italics="true" pageId="13" pageNumber="32">Uroobovella anatolica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557E7FFA8FFE7FDC45E93FDDF11CD" box="[539,633,1470,1496]" pageId="13" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3286505FFA8FFE7FED75D25FC9E1266" pageId="13" pageNumber="32" type="materials_examined">
<paragraph id="8B8D368EFFA8FFE7FED75D25FC9E1266" blockId="13.[264,1323,1544,1891]" pageId="13" pageNumber="32">
<emphasis id="B946EA9CFFA8FFE7FED75D25FE5D1237" bold="true" box="[264,507,1544,1570]" pageId="13" pageNumber="32">Material examined.</emphasis>
<typeStatus id="5489882CFFA8FFE7FDDD5D24FDC91236" box="[514,623,1545,1571]" pageId="13" pageNumber="32" type="holotype">Holotype</typeStatus>
Ψ, Bingöl province, Palulu Mountain (
<geoCoordinate id="EE065049FFA8FFE7FBE95D24FB031236" box="[1078,1189,1545,1571]" direction="north" orientation="latitude" pageId="13" pageNumber="32" precision="925" value="38.95">38°57’ N</geoCoordinate>
,
<geoCoordinate id="EE065049FFA8FFE7FB6E5D24FABA1236" box="[1201,1308,1545,1571]" direction="east" orientation="longitude" pageId="13" pageNumber="32" precision="925" value="40.233334">40°14’ E</geoCoordinate>
),
<date id="FF8C104EFFA8FFE7FED75D1CFE0C125E" box="[264,426,1585,1611]" pageId="13" pageNumber="32" value="2004-05-25">25 May 2004</date>
,
<quantity id="4CCA9B6BFFA8FFE7FE665D1CFDBD125E" box="[441,539,1585,1611]" metricMagnitude="0" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="32" unit="m" value="1.25">1,250 m</quantity>
a.s.l.
<typeStatus id="5489882CFFA8FFE7FDBB5D1CFD7F125E" box="[612,729,1585,1611]" pageId="13" pageNumber="32" type="paratype">Paratypes</typeStatus>
, 3 Ψ, 1 ɗ, same data as
<typeStatus id="5489882CFFA8FFE7FBDE5D1CFBCA125E" box="[1025,1132,1585,1611]" pageId="13" pageNumber="32" type="holotype">holotype</typeStatus>
. Litter and bark from deciduous tree stump samples were moist.
</paragraph>
</subSubSection>
<subSubSection id="C3286505FFA8FFFAFEE75DADFCCA1200" lastPageId="16" lastPageNumber="35" pageId="13" pageNumber="32" type="description">
<paragraph id="8B8D368EFFA8FFE7FEE75DADFC311376" blockId="13.[264,1323,1544,1891]" pageId="13" pageNumber="32">
<emphasis id="B946EA9CFFA8FFE7FEE75DADFE35128F" bold="true" box="[312,403,1664,1690]" pageId="13" pageNumber="32">Female</emphasis>
(
<typeStatus id="5489882CFFA8FFE7FE7D5DACFDB0128E" box="[418,534,1665,1691]" pageId="13" pageNumber="32" type="holotype">Holotype</typeStatus>
).
<emphasis id="B946EA9CFFA8FFE7FDF65DACFD21128F" box="[553,647,1665,1690]" italics="true" pageId="13" pageNumber="32">Dorsum</emphasis>
(
<figureCitation id="13092A0BFFA8FFE7FD485DACFD48128E" box="[663,750,1665,1691]" captionStart="FIGURE 15" captionStartId="15.[264,367,1468,1492]" captionTargetBox="[504,1081,646,1436]" captionTargetId="figure@15.[453,1133,600,1474]" captionTargetPageId="15" captionText="FIGURE 15. Uroobovella anatolica sp. nov., dorsal idiosoma of female; scale: 200 µm." httpUri="https://zenodo.org/record/174858/files/figure.png" pageId="13" pageNumber="32">Fig. 15</figureCitation>
). Idiosoma oval, 570 µm long and 420 µm wide, well sclerotised, brown. Dorsal plate entire, strongly convex, without ornamentation. Plate surrounded by an entire marginal plate. Marginal plate narrow, bearing 12 pairs of setae. Dorsal plate with 40–50 relatively short, simple thorn-like setae, setae not reaching insertion of following setae. Posterior end of dorsum with four knife-like setae, these powerful setae longer and wider than other dorsal setae.
</paragraph>
<caption id="DF4D6606FFABFFE4FED75F8FFB5910AC" box="[264,1279,1186,1210]" httpUri="https://zenodo.org/record/174857/files/figure.png" pageId="14" pageNumber="33" targetBox="[488,1109,330,1154]" targetPageId="14">
<paragraph id="8B8D368EFFABFFE4FED75F8FFB5910AC" blockId="14.[264,1279,1186,1210]" box="[264,1279,1186,1210]" pageId="14" pageNumber="33">
<emphasis id="B946EA9CFFABFFE4FED75F8FFE3110AF" bold="true" box="[264,407,1186,1210]" pageId="14" pageNumber="33">FIGURE 14.</emphasis>
<taxonomicName id="4C324D0DFFABFFE4FE415F8FFD2910AC" box="[414,655,1186,1209]" class="Arachnida" family="Trachyuropodidae" genus="Crinitodiscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="33" phylum="Arthropoda" rank="species" species="ayyildizi" status="sp. nov.">
<emphasis id="B946EA9CFFABFFE4FE415F8FFD2910AC" box="[414,655,1186,1209]" italics="true" pageId="14" pageNumber="33">Crinitodiscus ayyildizi</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFABFFE4FD495F8FFD4D10AF" bold="true" box="[662,747,1186,1210]" pageId="14" pageNumber="33">
<taxonomicNameLabel id="A27557E7FFABFFE4FD495F8FFD4D10AF" box="[662,747,1186,1210]" pageId="14" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: ventral idiosoma of deutonymph; scale: 100 µm.
</paragraph>
</caption>
<paragraph id="8B8D368EFFABFFE4FEE75FC0FEEC12F2" blockId="14.[264,1323,1261,1887]" pageId="14" pageNumber="33">
<emphasis id="B946EA9CFFABFFE4FEE75FC0FED91113" box="[312,383,1261,1286]" italics="true" pageId="14" pageNumber="33">Ven te r</emphasis>
(
<figureCitation id="13092A0BFFABFFE4FE4D5FC0FE4F1112" box="[402,489,1261,1287]" captionStart="FIGURE 16" captionStartId="16.[264,367,1296,1320]" captionTargetBox="[474,1100,392,1295]" captionTargetId="figure@16.[453,1133,392,1295]" captionTargetPageId="16" captionText="FIGURE 16. Uroobovella anatolica, ventral idiosoma of female; scale: 200 µm." httpUri="https://zenodo.org/record/174859/files/figure.png" pageId="14" pageNumber="33">Fig. 16</figureCitation>
). Epigynial plate 150 µm long and 110 µm wide, helmet-like, bearing a crown-like anterior extension; posterior margin behind coxae IV, anterior margin behind coxae I. Surface of plate with punctuations and many shining pores. Ventral region with 4 pairs of sternal (
<emphasis id="B946EA9CFFABFFE4FE175E48FDB0116B" box="[456,534,1381,1406]" italics="true" pageId="14" pageNumber="33">st1-st4</emphasis>
) setae, and 7 pairs of ventri-anal setae. Sternal setae
<emphasis id="B946EA9CFFABFFE4FB585E48FB0F116B" box="[1159,1193,1381,1406]" italics="true" pageId="14" pageNumber="33">st1</emphasis>
inserted at the level coxae II, close to epigynium;
<emphasis id="B946EA9CFFABFFE4FD205EA0FCEB11B3" box="[767,845,1421,1446]" italics="true" pageId="14" pageNumber="33">st2-st3</emphasis>
lateral to genital aperture,
<emphasis id="B946EA9CFFABFFE4FB745EA0FB6B11B3" box="[1195,1229,1421,1446]" italics="true" pageId="14" pageNumber="33">st4</emphasis>
behind epigynial plate. Anterior sternal region with a chitinous wrinkled surface process. Ventrianal plate ornamented with micropunctations and small porous shining markings. All ventral setae simple, smooth, needle-like. Peritremes at level of coxae II and pedofossae III, anterior prolongation directed exteriorly. Stigmatal openings with short posterior prolongations. Endopodal line absent. Exopodal plate ornamented with shining pores. Pedofossal grooves distinct and well developed, fossae IV also with shining pores. Anal opening 13 µm in diameter. Post-anal seta present and similar in form to the other ventral setae.
</paragraph>
<paragraph id="8B8D368EFFABFFE5FEE75DD8FC32165B" blockId="14.[264,1323,1261,1887]" lastBlockId="15.[264,1322,284,590]" lastPageId="15" lastPageNumber="34" pageId="14" pageNumber="33">
<emphasis id="B946EA9CFFABFFE4FEE75DD8FE68131B" box="[312,462,1781,1806]" italics="true" pageId="14" pageNumber="33">Gnathosoma</emphasis>
(
<figureCitation id="13092A0BFFABFFE4FE015DD8FD93131A" box="[478,565,1781,1807]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="14" pageNumber="33">Fig. 17</figureCitation>
). Chelicerae unidentate, with large nodus, movable digit 30 µm, fixed digit 52 µm, and middle section 160 µm long (
<figureCitation id="13092A0BFFABFFE4FCA35C30FC741322" box="[892,978,1821,1847]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="14" pageNumber="33">Fig. 17</figureCitation>
A). Setae
<emphasis id="B946EA9CFFABFFE4FB965C30FBC01323" box="[1097,1126,1821,1846]" italics="true" pageId="14" pageNumber="33">h1</emphasis>
simple, smooth, thorn-like, almost reaching end of laciniae. Setae
<emphasis id="B946EA9CFFABFFE4FC965C68FCC0134B" box="[841,870,1861,1886]" italics="true" pageId="14" pageNumber="33">h2</emphasis>
smooth, short and stout, reaching base of
<emphasis id="B946EA9CFFAAFFE5FEF45A31FEEE1520" box="[299,328,284,309]" italics="true" pageId="15" pageNumber="34">h1</emphasis>
. Setae
<emphasis id="B946EA9CFFAAFFE5FE7B5A31FE561520" box="[420,496,284,309]" italics="true" pageId="15" pageNumber="34">h3, h4</emphasis>
branched;
<emphasis id="B946EA9CFFAAFFE5FDA75A31FD301520" box="[632,662,284,309]" italics="true" pageId="15" pageNumber="34">h4</emphasis>
reaching past base of
<emphasis id="B946EA9CFFAAFFE5FC755A31FC6E1520" box="[938,968,284,309]" italics="true" pageId="15" pageNumber="34">h3</emphasis>
, and
<emphasis id="B946EA9CFFAAFFE5FBD15A31FB8D1520" box="[1038,1067,284,309]" italics="true" pageId="15" pageNumber="34">h2</emphasis>
reaching base of
<emphasis id="B946EA9CFFAAFFE5FAD95A31FA851520" box="[1286,1315,284,309]" italics="true" pageId="15" pageNumber="34">h1</emphasis>
. Corniculus horn-like, with blunt end. Hypostomal constrictions present between
<emphasis id="B946EA9CFFAAFFE5FAD95A69FA821548" box="[1286,1316,324,349]" italics="true" pageId="15" pageNumber="34">h3</emphasis>
. Deutosternum with superficial denticulation (
<figureCitation id="13092A0BFFAAFFE5FCC45A41FCD61593" box="[795,880,364,390]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="15" pageNumber="34">Fig. 17</figureCitation>
C). Epistome with triangular base and denticulate lateral margins, denticles gradually lengthening from tip to base (
<figureCitation id="13092A0BFFAAFFE5FB715AB9FAA115BB" box="[1198,1287,404,430]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="15" pageNumber="34">Fig. 17</figureCitation>
B). Tritosternal base vase-shaped, medio-lateral lacinia laterally denticulate and tip bifid, lateral part of laciniae denticulate and shorter than middle part; lateral branches denticulate on one side (
<figureCitation id="13092A0BFFAAFFE5FE735921FDA31633" box="[428,517,524,550]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="15" pageNumber="34">Fig. 17</figureCitation>
D). Palps short and robust, palp tarsus with a set of tactile setae, ventral setae of trochanter finely denticulate (
<figureCitation id="13092A0BFFAAFFE5FCC35919FCD7165B" box="[796,881,564,590]" captionStart="FIGURE 17" captionStartId="17.[264,367,1129,1153]" captionTargetBox="[517,1077,320,1100]" captionTargetId="figure@17.[453,1133,252,1125]" captionTargetPageId="17" captionText="FIGURE 17. Uroobovella anatolica sp. nov., female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm." httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="15" pageNumber="34">Fig. 17</figureCitation>
E).
</paragraph>
<caption id="DF4D6606FFAAFFE5FED75E91FB1C11C6" box="[264,1210,1468,1492]" httpUri="https://zenodo.org/record/174858/files/figure.png" pageId="15" pageNumber="34" targetBox="[504,1081,646,1436]" targetPageId="15">
<paragraph id="8B8D368EFFAAFFE5FED75E91FB1C11C6" blockId="15.[264,1210,1468,1492]" box="[264,1210,1468,1492]" pageId="15" pageNumber="34">
<emphasis id="B946EA9CFFAAFFE5FED75E91FE3711C1" bold="true" box="[264,401,1468,1492]" pageId="15" pageNumber="34">FIGURE 15</emphasis>
.
<taxonomicName id="4C324D0DFFAAFFE5FE415E91FD3611C6" box="[414,656,1468,1491]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="15" pageNumber="34" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFAAFFE5FE415E91FD3611C6" box="[414,656,1468,1491]" italics="true" pageId="15" pageNumber="34">Uroobovella anatolica</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFAAFFE5FD495E91FD4D11C1" bold="true" box="[662,747,1468,1492]" pageId="15" pageNumber="34">
<taxonomicNameLabel id="A27557E7FFAAFFE5FD495E91FD4D11C1" box="[662,747,1468,1492]" pageId="15" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal idiosoma of female; scale: 200 µm.
</paragraph>
</caption>
<paragraph id="8B8D368EFFAAFFE5FEE75D2AFC35128C" blockId="15.[264,1323,1543,1889]" pageId="15" pageNumber="34">
<emphasis id="B946EA9CFFAAFFE5FEE75D2AFEC91235" box="[312,367,1543,1568]" italics="true" pageId="15" pageNumber="34">Legs</emphasis>
(
<figureCitation id="13092A0BFFAAFFE5FEA05D2AFE711234" box="[383,471,1543,1569]" captionStart="FIGURE 18" captionStartId="17.[264,367,1720,1744]" captionTargetBox="[484,1096,1241,1680]" captionTargetId="figure@17.[453,1133,1225,1700]" captionTargetPageId="17" captionText="FIGURE 18. Uroobovella anatolica sp. nov., female. A: leg I; B: leg II; C: leg III; D: leg IV; male. E: femur II with apophysis; scale: 100 µm." httpUri="https://zenodo.org/record/174861/files/figure.png" pageId="15" pageNumber="34">Fig. 18</figureCitation>
). Legs I–II directed anteriorly, and III–IV posteriorly. Tarsi II–IV with a pair of claws at tip of ambulacral prolongation; tarsi I with a lighter claw on a delicate ambulacrum. Whip-like setae longer than tarsi I. Setae on all legs simple and thorn-like. Femurs and trochanters bearing a membraneous flange.
</paragraph>
<paragraph id="8B8D368EFFAAFFE5FEE75D8AFECC12FC" blockId="15.[264,1323,1543,1889]" pageId="15" pageNumber="34">
<emphasis id="B946EA9CFFAAFFE5FEE75D8AFED812D4" bold="true" box="[312,382,1703,1729]" pageId="15" pageNumber="34">Male.</emphasis>
<emphasis id="B946EA9CFFAAFFE5FE585D8AFE4312D5" box="[391,485,1703,1728]" italics="true" pageId="15" pageNumber="34">Dorsum</emphasis>
. All dorsal features as in females. Length of idiosoma 575 µm, width 430 µm.
</paragraph>
<paragraph id="8B8D368EFFAAFFE5FEE75DDAFA8D1374" blockId="15.[264,1323,1543,1889]" pageId="15" pageNumber="34">
<emphasis id="B946EA9CFFAAFFE5FEE75DDAFED91305" box="[312,383,1783,1808]" italics="true" pageId="15" pageNumber="34">Ven te r</emphasis>
(
<figureCitation id="13092A0BFFAAFFE5FE4C5DDAFE4C1304" box="[403,490,1783,1809]" captionStart="FIGURE 19" captionStartId="18.[264,367,1816,1840]" captionTargetBox="[497,1107,1001,1790]" captionTargetId="figure@18.[453,1133,950,1832]" captionTargetPageId="18" captionText="FIGURE 19. Uroobovella anatolica sp. nov., dorsal idiosoma of male; scale: 200 µm." httpUri="https://zenodo.org/record/174862/files/figure.png" pageId="15" pageNumber="34">Fig. 19</figureCitation>
). Surface of sternal plate ornamented with micropunctations and small rounded hollows. Sternal setae straight and thorn-like. Sternal setae
<emphasis id="B946EA9CFFAAFFE5FBF75C32FBEF132D" box="[1064,1097,1823,1848]" italics="true" pageId="15" pageNumber="34">st1</emphasis>
inserted level with coxae II,
<emphasis id="B946EA9CFFAAFFE5FEA25C6AFE6C1375" box="[381,458,1863,1888]" italics="true" pageId="15" pageNumber="34">st2-st4</emphasis>
anterior of genital aperture on line,
<emphasis id="B946EA9CFFAAFFE5FC555C6AFC0A1375" box="[906,940,1863,1888]" italics="true" pageId="15" pageNumber="34">st5</emphasis>
close to posterior end of plate.
</paragraph>
<paragraph id="8B8D368EFFB5FFFAFED75A31FB131593" blockId="16.[264,1323,284,390]" pageId="16" pageNumber="35">Genital aperture circular, between coxae IV, and surrounded thick chitinous wall. Genital plate 45 µm long and 40 µm wide. Surface of genital plate with micropunctation. Sternal plate with a chitinous wrinckled surface ornamentations in front of genital plate.</paragraph>
<caption id="DF4D6606FFB5FFFAFED75E3DFB651132" box="[264,1219,1296,1320]" httpUri="https://zenodo.org/record/174859/files/figure.png" pageId="16" pageNumber="35" targetBox="[474,1100,392,1295]" targetPageId="16">
<paragraph id="8B8D368EFFB5FFFAFED75E3DFB651132" blockId="16.[264,1219,1296,1320]" box="[264,1219,1296,1320]" pageId="16" pageNumber="35">
<emphasis id="B946EA9CFFB5FFFAFED75E3DFE37113D" bold="true" box="[264,401,1296,1320]" pageId="16" pageNumber="35">FIGURE 16</emphasis>
.
<taxonomicName id="4C324D0DFFB5FFFAFE415E3DFD361132" box="[414,656,1296,1319]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="anatolica">
<emphasis id="B946EA9CFFB5FFFAFE415E3DFD361132" box="[414,656,1296,1319]" italics="true" pageId="16" pageNumber="35">Uroobovella anatolica</emphasis>
</taxonomicName>
, ventral idiosoma of female; scale: 200 µm.
</paragraph>
</caption>
<paragraph id="8B8D368EFFB5FFFAFEE75E76FEEE1188" blockId="16.[264,1323,1370,1877]" pageId="16" pageNumber="35">
<emphasis id="B946EA9CFFB5FFFAFEE75E76FEC91161" box="[312,367,1371,1396]" italics="true" pageId="16" pageNumber="35">Legs</emphasis>
. A sexual dimorphism occurs in femur II which bears a powerful apophysis (
<figureCitation id="13092A0BFFB5FFFAFB245E77FE801188" captionStart="FIGURE 18" captionStartId="17.[264,367,1720,1744]" captionTargetBox="[484,1096,1241,1680]" captionTargetId="figure@17.[453,1133,1225,1700]" captionTargetPageId="17" captionText="FIGURE 18. Uroobovella anatolica sp. nov., female. A: leg I; B: leg II; C: leg III; D: leg IV; male. E: femur II with apophysis; scale: 100 µm." httpUri="https://zenodo.org/record/174861/files/figure.png" pageId="16" pageNumber="35">Fig. 18</figureCitation>
E).
</paragraph>
<paragraph id="8B8D368EFFB5FFFAFEE75E86FEF911F8" blockId="16.[264,1323,1370,1877]" pageId="16" pageNumber="35">Other morphological features, setal arrangement, body shape and patterns as for female.</paragraph>
<paragraph id="8B8D368EFFB5FFFAFEE75ED6FCCA1200" blockId="16.[264,1323,1370,1877]" box="[312,876,1531,1557]" pageId="16" pageNumber="35">Deutonymph, protonymph and larva. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3286505FFB5FFF9FEE75D0FFEDB10DB" lastPageId="19" lastPageNumber="38" pageId="16" pageNumber="35" type="discussion">
<paragraph id="8B8D368EFFB5FFFAFEE75D0FFDC312A0" blockId="16.[264,1323,1370,1877]" pageId="16" pageNumber="35">
<emphasis id="B946EA9CFFB5FFFAFEE75D0FFE171229" bold="true" box="[312,433,1570,1596]" pageId="16" pageNumber="35">Remarks.</emphasis>
<bibRefCitation id="EFA34B7FFFB5FFFAFE615D0EFD0F1228" author="Hirschmann" box="[446,681,1571,1597]" pageId="16" pageNumber="39" refString="Hirschmann, W. (1989) Die Ganggattung Uroobovella Berlese 1903. Artengruppen - Bestimmungstabellen - Diagnosen (Dinychini, Uropodinae). Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 36, 84 - 196." type="journal article" year="1989">Hirschmann (1989)</bibRefCitation>
reviewed the cosmopolitan genus
<taxonomicName id="4C324D0DFFB5FFFAFB805D0EFB541229" box="[1119,1266,1571,1596]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B946EA9CFFB5FFFAFB805D0EFB541229" box="[1119,1266,1571,1596]" italics="true" pageId="16" pageNumber="35">Uroobovella</emphasis>
</taxonomicName>
and created 23 species-groups for easy identification. All distinctive characters for the speciesgroups of the genus
<taxonomicName id="4C324D0DFFB5FFFAFDCD5D5EFD031299" box="[530,677,1651,1676]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B946EA9CFFB5FFFAFDCD5D5EFD031299" box="[530,677,1651,1676]" italics="true" pageId="16" pageNumber="35">Uroobovella</emphasis>
</taxonomicName>
are summarised below, emphasising the distinctive characters of the new species.
</paragraph>
<paragraph id="8B8D368EFFB5FFFAFEE75DEEFE581340" blockId="16.[264,1323,1370,1877]" pageId="16" pageNumber="35">
<taxonomicName id="4C324D0DFFB5FFFAFEE75DEEFE6B12C9" box="[312,461,1731,1756]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFB5FFFAFEE75DEEFE6B12C9" box="[312,461,1731,1756]" italics="true" pageId="16" pageNumber="35">U. anatolica</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFB5FFFAFE085DEFFD9112C9" bold="true" box="[471,567,1730,1756]" pageId="16" pageNumber="35">
<taxonomicNameLabel id="A27557E7FFB5FFFAFE085DEFFD9112C9" box="[471,567,1730,1756]" pageId="16" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is differentiated from the
<emphasis id="B946EA9CFFB5FFFAFCA85DEEFEEF1339" italics="true" pageId="16" pageNumber="35">vinicolora, pergibba, costai, schulzi, tricuspis, hilli, marginata, ruhmi, coprophila, flagelliger, fimicola, appendiculata, minima, ipidis</emphasis>
species-groups and all related species by the presence of five pairs of sternal setae in males (
<quantity id="4CCA9B6BFFB5FFFAFE835C16FE2C1340" box="[348,394,1851,1877]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="16" pageNumber="35" unit="in" value="4.0">4 in</quantity>
females).
</paragraph>
<caption id="DF4D6606FFB4FFFBFED75F44FC6410B6" httpUri="https://zenodo.org/record/174860/files/figure.png" pageId="17" pageNumber="36" targetBox="[517,1077,320,1100]" targetPageId="17">
<paragraph id="8B8D368EFFB4FFFBFED75F44FC6410B6" blockId="17.[264,1323,1129,1187]" pageId="17" pageNumber="36">
<emphasis id="B946EA9CFFB4FFFBFED75F44FE3C1094" bold="true" box="[264,410,1129,1153]" pageId="17" pageNumber="36">FIGURE 17</emphasis>
.
<taxonomicName id="4C324D0DFFB4FFFBFE6F5F47FD0C1094" box="[432,682,1130,1153]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFB4FFFBFE6F5F47FD0C1094" box="[432,682,1130,1153]" italics="true" pageId="17" pageNumber="36">Uroobovella anatolica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557E7FFB4FFFBFD665F44FCB11094" box="[697,791,1129,1153]" pageId="17" pageNumber="36" rank="species">
<emphasis id="B946EA9CFFB4FFFBFD665F44FCB11094" bold="true" box="[697,791,1129,1153]" pageId="17" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. A: Ventral view of gnathosoma; B: Tritosternum; C: Epistome; D: Chelicera; E: Palp; scale: 100 µm.
</paragraph>
</caption>
<caption id="DF4D6606FFB4FFFBFED75D95FD7712E7" httpUri="https://zenodo.org/record/174861/files/figure.png" pageId="17" pageNumber="36" targetBox="[484,1096,1241,1680]" targetPageId="17">
<paragraph id="8B8D368EFFB4FFFBFED75D95FD7712E7" blockId="17.[264,1323,1720,1778]" pageId="17" pageNumber="36">
<emphasis id="B946EA9CFFB4FFFBFED75D95FE3712C5" bold="true" box="[264,401,1720,1744]" pageId="17" pageNumber="36">FIGURE 18</emphasis>
.
<taxonomicName id="4C324D0DFFB4FFFBFE415D95FD2912DA" box="[414,655,1720,1743]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFB4FFFBFE415D95FD2912DA" box="[414,655,1720,1743]" italics="true" pageId="17" pageNumber="36">Uroobovella anatolica</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFB4FFFBFD4A5D95FD4C12C5" bold="true" box="[661,746,1720,1744]" pageId="17" pageNumber="36">
<taxonomicNameLabel id="A27557E7FFB4FFFBFD4A5D95FD4C12C5" box="[661,746,1720,1744]" pageId="17" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A: leg I; B: leg II; C: leg III; D: leg IV; male. E: femur II with apophysis; scale: 100 µm.
</paragraph>
</caption>
<paragraph id="8B8D368EFFB4FFF8FEE75C08FE6117A3" blockId="17.[264,1323,1829,1895]" lastBlockId="18.[264,1323,284,950]" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36">
Hovewer, the
<emphasis id="B946EA9CFFB4FFFBFE3B5C0BFE401372" italics="true" pageId="17" pageNumber="36">
foveolata, venusta, elegans, formosana,
<taxonomicName id="4C324D0DFFB4FFFBFC125C0BFBBF132A" box="[973,1049,1830,1855]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="rackei">rackei</taxonomicName>
, micherdzinski, fracta, carinata, pulchella
</emphasis>
and
<emphasis id="B946EA9CFFB4FFFBFDC15C63FD271372" box="[542,641,1870,1895]" italics="true" pageId="17" pageNumber="36">vitzhumi</emphasis>
species-groups and related species share the same number of sternal setae (5 pairs in male 5, 4 pairs in females). The new species is differentiated from the
<emphasis id="B946EA9CFFB7FFF8FEAB5A69FDEC1548" box="[372,586,324,349]" italics="true" pageId="18" pageNumber="37">foveolata, venusta</emphasis>
and
<emphasis id="B946EA9CFFB7FFF8FD5D5A69FD7A1548" box="[642,732,324,349]" italics="true" pageId="18" pageNumber="37">elegans</emphasis>
species-groups, by the presence of a pair of claws on pretarsus I. Members of
<emphasis id="B946EA9CFFB7FFF8FD945A41FD6E1590" box="[587,712,364,389]" italics="true" pageId="18" pageNumber="37">formosana</emphasis>
species-group possess a pair of claws on legs I, but, lack marginal plates and endopodal lines, and their males do not have an apophysis on legs II. On the other hand, in
<emphasis id="B946EA9CFFB7FFF8FDF45A91FD3915C0" box="[555,671,444,469]" italics="true" pageId="18" pageNumber="37">
<taxonomicName id="4C324D0DFFB7FFF8FDF45A91FD3D15C0" box="[555,667,444,469]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="37" phylum="Arthropoda" rank="species" species="rackei">U. rackei</taxonomicName>
,
</emphasis>
there is a well-developed marginal plate, but the dorsal and ventral plates are separated by a soft chitinised area containing platelets. The new species is differentiated from
<taxonomicName id="4C324D0DFFB7FFF8FDA45921FD4B1630" box="[635,749,524,549]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="37" phylum="Arthropoda" rank="species" species="rackei">
<emphasis id="B946EA9CFFB7FFF8FDA45921FD4B1630" box="[635,749,524,549]" italics="true" pageId="18" pageNumber="37">U. rackei</emphasis>
</taxonomicName>
by the relatively hard and heavily sclerotised processes between the dorsal and ventral plates. The species groups
<emphasis id="B946EA9CFFB7FFF8FBF35919FE4C1660" italics="true" pageId="18" pageNumber="37">micherdzinski, fracta, carinata, pulchella</emphasis>
have a marginal plate as a distinctive character in common with the new species. Hovewer, the new species is differentiated from the
<emphasis id="B946EA9CFFB7FFF8FBD659A9FB0D1688" box="[1033,1195,644,669]" italics="true" pageId="18" pageNumber="37">micherdzinski</emphasis>
and
<emphasis id="B946EA9CFFB7FFF8FB3B59A9FA8C1688" box="[1252,1322,644,669]" italics="true" pageId="18" pageNumber="37">fracta</emphasis>
species-groups by the presence of structural ornamentation on these plates. The
<emphasis id="B946EA9CFFB7FFF8FB185981FA8D16D0" box="[1223,1323,684,709]" italics="true" pageId="18" pageNumber="37">carinata</emphasis>
species-group has a dorsal hunch-backed carina, which does not exist in the new species. The new species differs from
<emphasis id="B946EA9CFFB7FFF8FDBE59D1FD631700" box="[609,709,764,789]" italics="true" pageId="18" pageNumber="37">vitzhumi</emphasis>
species-group by the absence of lateral shoulder and vertex, deep circular and polygonal hollows on the plates, marginal plates with densely undulating marginal plate internally, and endopodal line and tasseled setae on marginal plate. Finally, the
<emphasis id="B946EA9CFFB7FFF8FE3F5859FDEB1798" box="[480,589,884,909]" italics="true" pageId="18" pageNumber="37">pectinata</emphasis>
species-group does not have leg pedofossae, in contrast with the new species.
</paragraph>
<caption id="DF4D6606FFB7FFF8FED75C35FB03133A" box="[264,1189,1816,1840]" httpUri="https://zenodo.org/record/174862/files/figure.png" pageId="18" pageNumber="37" targetBox="[497,1107,1001,1790]" targetPageId="18">
<paragraph id="8B8D368EFFB7FFF8FED75C35FB03133A" blockId="18.[264,1189,1816,1840]" box="[264,1189,1816,1840]" pageId="18" pageNumber="37">
<emphasis id="B946EA9CFFB7FFF8FED75C35FE371325" bold="true" box="[264,401,1816,1840]" pageId="18" pageNumber="37">FIGURE 19</emphasis>
.
<taxonomicName id="4C324D0DFFB7FFF8FE415C34FD361325" box="[414,656,1817,1840]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="18" pageNumber="37" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFB7FFF8FE415C34FD361325" box="[414,656,1817,1840]" italics="true" pageId="18" pageNumber="37">Uroobovella anatolica</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFB7FFF8FD495C35FD4D1325" bold="true" box="[662,747,1816,1840]" pageId="18" pageNumber="37">
<taxonomicNameLabel id="A27557E7FFB7FFF8FD495C35FD4D1325" box="[662,747,1816,1840]" pageId="18" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal idiosoma of male; scale: 200 µm.
</paragraph>
</caption>
<paragraph id="8B8D368EFFB6FFF9FEE75A31FCDB16D3" blockId="19.[264,1323,284,1230]" pageId="19" pageNumber="38">
The new species shares some morphological characters with the species of the
<emphasis id="B946EA9CFFB6FFF9FED75A69FED31548" box="[264,373,324,349]" italics="true" pageId="19" pageNumber="38">pulchella</emphasis>
group. The members of this species-group have 5 pairs of sternal setae in males and
<quantity id="4CCA9B6BFFB6FFF9FEE45A41FECF1593" box="[315,361,364,390]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="19" pageNumber="38" unit="in" value="4.0">4 in</quantity>
females; the idiosoma is egg shaped or widely oval; endopodal lines are present, legs I lack claws; with a well-developed marginal plate and with structural plates are present; and the marginal plate is smooth sided medially. The new species is closely related to
<taxonomicName id="4C324D0DFFB6FFF9FE5A5AC9FC5E15EB" authority="Hirschmann &amp; Zirngiebl-Nicol 1972" authorityName="Hirschmann &amp; Zirngiebl-Nicol" authorityYear="1972" box="[389,1016,484,510]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B946EA9CFFB6FFF9FE5A5AC9FD9715E8" box="[389,561,484,509]" italics="true" pageId="19" pageNumber="38">U. denticulata</emphasis>
<bibRefCitation id="EFA34B7FFFB6FFF9FDE35AC9FC5E15EB" author="Hirschmann" box="[572,1016,484,510]" pageId="19" pageNumber="39" refString="Hirschmann, W. (1972) Gangsystematik der Parasitiformes. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 18, 68 - 92" type="journal article" year="1972">
Hirschmann &amp; Zirngiebl-Nicol
<quantity id="4CCA9B6BFFB6FFF9FC625AC9FC5E15EB" box="[957,1016,484,510]" metricMagnitude="1" metricUnit="m" metricValue="5.00888" pageId="19" pageNumber="38" unit="in" value="1972.0">1972</quantity>
</bibRefCitation>
</taxonomicName>
in the
<emphasis id="B946EA9CFFB6FFF9FB8B5AC9FB6715E8" box="[1108,1217,484,509]" italics="true" pageId="19" pageNumber="38">pulchella</emphasis>
speciesgroup, which has some very similar morphological characters with the new species: with web-like patterns on plates, presence of well developed endopodal lines, and the existence of short simple setae on the dorsal, marginal, lateral, sternal and ventral plates. Hovewer, the epigynium is the most distinctive character for
<emphasis id="B946EA9CFFB6FFF9FCB359A9FBBA1688" box="[876,1052,644,669]" italics="true" pageId="19" pageNumber="38">
<taxonomicName id="4C324D0DFFB6FFF9FCB359A9FBBF1688" box="[876,1049,644,669]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulata">U. denticulata</taxonomicName>
,
</emphasis>
which has a wide and notched extension apically, as wide as the epigynium.
</paragraph>
<paragraph id="8B8D368EFFB6FFF9FEE759F9FEDB10DB" blockId="19.[264,1323,284,1230]" pageId="19" pageNumber="38">
In summary,
<taxonomicName id="4C324D0DFFB6FFF9FE0959F9FDCD16F8" box="[470,619,724,749]" class="Arachnida" family="Urodinychidae" genus="Uroobovella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="anatolica" status="sp. nov.">
<emphasis id="B946EA9CFFB6FFF9FE0959F9FDCD16F8" box="[470,619,724,749]" italics="true" pageId="19" pageNumber="38">U. anatolica</emphasis>
</taxonomicName>
<emphasis id="B946EA9CFFB6FFF9FDAB59F9FD7216FB" bold="true" box="[628,724,724,750]" pageId="19" pageNumber="38">
<taxonomicNameLabel id="A27557E7FFB6FFF9FDAB59F9FD7216FB" box="[628,724,724,750]" pageId="19" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all of its congeners by the following combinaton of character: males with five (and females four) pairs of sternal setae; tarsi I without claws; dorsal plate smooth and without ornamentation; tongue-like epigynium with a dentate anterior prolongation; leg pedofossae well-developed; marginal plate present and not fragmented posteriorly; all dorsal, marginal, lateral and ventral setae simple, smooth and thorn-like; sternal plate with a chitinous wrinkled surface ornamentations near anterior end of epigynium, ventri-anal and epigynial plates ornamented with micropunctations and small circular pores; soft cuticle supporting dorsal and ventral plates absent, and cuticle of this region hardened and sclerotised; medial borders of marginal plate smooth and without chitinous undulations; hyaline prolongation of fixed digit of chelicera knife-like; peritremes hook-like anteriorly and the anterior end inclined inward, and posterior prolongation smooth; and males with apophysis in leg femora II.
</paragraph>
</subSubSection>
</treatment>
</document>
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