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<document id="D269704FC68FB45C9126F9F4891E0E3E" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. &amp; Congdon, Bradley C." docDate="2021" docId="039B87838757FFFDFF1EF4A51153F97D" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Seraphs minus" docType="treatment" docVersion="7" lastPageNumber="414" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="414" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2324AD022089C599B2CC7A88971635E8">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="5EA032923D18D62A4A864AB945512E80">Maxwell, Stephen J.</mods:namePart>
<mods:nameIdentifier id="9AC7201A4F6CA87AF347ACFEEBFFE1A7" type="ORCID">0000-0002-4328-6171</mods:nameIdentifier>
<mods:affiliation id="2B4971FED9302EB00A8D9FBFA94FA8D7">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="80B1E85A1FA5099995C91176D653FD08">Rymer, Tasmin L.</mods:namePart>
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<mods:affiliation id="6323D316486B8DBE4290B1798BFD9534">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
<mods:nameIdentifier id="024CC795B39A51F2C335F25E3D4FE64C" type="email">amsin.rymer@jcu.edu.au</mods:nameIdentifier>
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<mods:namePart id="8E7434396DFE8AFEB12B39CC2889A92E">Congdon, Bradley C.</mods:namePart>
<mods:nameIdentifier id="1D3FCEEFC66D26932D15CA656D8FB84B" type="ORCID">0000-0002-8751-0892</mods:nameIdentifier>
<mods:affiliation id="FF1C2A3CED9B9B4176015706FAE7E5A4">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
<mods:nameIdentifier id="D72DED52F599AC6BA494AB8B147B06B8" type="email">brad.congdon@jcu.edu.au</mods:nameIdentifier>
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<mods:title id="176D7E1351416C87A5A89818C7A3B71D">Zootaxa</mods:title>
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<mods:date id="618968F291AC6F8DB94018573018DF13">2021</mods:date>
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<mods:number id="50CDA3E5382D46D651547283EF56B6E0">2021-06-22</mods:number>
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<treatment id="039B87838757FFFDFF1EF4A51153F97D" ID-DOI="http://doi.org/10.5281/zenodo.5088686" ID-GBIF-Taxon="183411019" ID-Zenodo-Dep="5088686" LSID="urn:lsid:plazi:treatment:039B87838757FFFDFF1EF4A51153F97D" httpUri="http://treatment.plazi.org/id/039B87838757FFFDFF1EF4A51153F97D" lastPageNumber="414" pageId="13" pageNumber="414">
<subSubSection id="C328651E8757FFFDFF1EF4A5113EFB2B" box="[151,528,1183,1210]" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="8B8D36958757FFFDFF1EF4A5113EFB2B" blockId="13.[151,528,1183,1246]" box="[151,528,1183,1210]" pageId="13" pageNumber="414">
<heading id="D0C581F98757FFFDFF1EF4A5113EFB2B" bold="true" box="[151,528,1183,1210]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
<taxonomicName id="4C324D168757FFFDFF1EF4A5113EFB2B" authority="(Vincent, 1913)" baseAuthorityName="Vincent" baseAuthorityYear="1913" box="[151,528,1183,1210]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="minus">
<emphasis id="B946EA878757FFFDFF1EF4A5113EFB2B" bold="true" box="[151,528,1183,1210]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF1EF4A51265FB2B" bold="true" box="[151,331,1183,1210]" italics="true" pageId="13" pageNumber="414">Seraphs minus</emphasis>
(
<bibRefCitation id="EFA34B648757FFFDFED2F4A51126FB2B" author="Vincent, E." box="[347,520,1183,1209]" pageId="13" pageNumber="414" pagination="1 - 46" refId="ref42812" refString="Vincent, E. (1913) Contribution a la paleontologie dea falaises de Landana (Bas-Congo), Mollusques. Annales du Musee du Congo, 1, 1 - 46." type="journal article" year="1913">Vincent, 1913</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C328651E8757FFFDFF1EF4F91224FB4C" box="[151,266,1219,1246]" pageId="13" pageNumber="414" type="description">
<paragraph id="8B8D36958757FFFDFF1EF4F91224FB4C" blockId="13.[151,528,1183,1246]" box="[151,266,1219,1246]" pageId="13" pageNumber="414">
<figureCitation id="13092A108757FFFDFF1EF4F91224FB4C" box="[151,266,1219,1246]" captionStart="FIGURE 5" captionStartId="9.[151,250,1632,1656]" captionTargetBox="[169,1422,191,1598]" captionTargetId="figure-19@9.[151,1435,181,1609]" captionTargetPageId="9" captionText="FIGURE 5. Seraphs Montfort, 1810. A. Seraphs minus (Vincent, 1913), length 7.1 mm (Jung 1974, pl. 6, figs. 9, 10). B. Seraphs naricus (Vredenberg, 1925), holotype, GSI 12614, length 38.6 mm (Jung 1974, pl. 6, figs. 20–22). C. Seraphs olivaceus (Cossmann, 1889), MNHN A28577, Collection de Paleontologie, Paris, length 32 mm (Caze et al. 2010, figs. 6H, I). D. Seraphs peterjungi Caze et al., 2010, holotype, MNHN A28578, Collection de Paleontologie, length 16 mm (Caze et al. 2010, figs. 15J, K, N). E. Seraphs placiferus (Bayan, 1870), lectotype, MNHN G2100, length 33.7 mm (Jung 1974, pl. 7, figs. 5–7). F. Seraphs sopita (Brander, 1766), lectotype, NHMUK GG21010, length 26.4 mm (Jung 1974, pl. 1, figs. 1–3). G. Seraphs squamosus (Martin, 1914), Lectotype, RGM, Leiden St10344, length 37.1 mm (Jung 1974, pl. 6, figs. 17–19). H. Seraphs striatus (von Koenen, 1889), holotype, length 38.5 mm (von Koenen 1889, pl. I, figs. 7A–C). I. Seraphs subconvolutus (d’Orbigny, 1852), lectotype, MNHN UBT65-2-120, length 28 mm (Caze et al. 2010, figs. 14A, B). J. Seraphs volutatus (Brander, 1766), MNHN A28776, Collection de Paleontologie, Paris, length 17 mm (Caze et al. 2010, figs. 6A–C)." figureDoi="http://doi.org/10.5281/zenodo.5026789" httpUri="https://zenodo.org/record/5026789/files/figure.png" pageId="13" pageNumber="414">Figure 5A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF1EF5321118FAAD" pageId="13" pageNumber="414" type="reference_group">
<paragraph id="8B8D36958757FFFDFF1EF5321118FAAD" blockId="13.[151,1436,1288,1343]" pageId="13" pageNumber="414">
1913
<taxonomicName id="4C324D168757FFFDFF58F532116AFAB2" authority="Vincent" authorityName="Vincent" box="[209,580,1288,1312]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="minus" subGenus="Seraphs">
<emphasis id="B946EA878757FFFDFF58F532126FFAB2" box="[209,321,1288,1312]" italics="true" pageId="13" pageNumber="414">Terebellum</emphasis>
(
<emphasis id="B946EA878757FFFDFED8F532128DFAB2" box="[337,419,1288,1312]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
)
<emphasis id="B946EA878757FFFDFE3BF53312C1FAB2" box="[434,495,1289,1312]" italics="true" pageId="13" pageNumber="414">minus</emphasis>
Vincent
</taxonomicName>
,
<emphasis id="B946EA878757FFFDFDD8F5331036FAB2" box="[593,792,1288,1312]" italics="true" pageId="13" pageNumber="414">p. 19, pl. 1, figs. 23</emphasis>
&amp;
<emphasis id="B946EA878757FFFDFCB2F5321075FAB2" box="[827,859,1288,1312]" italics="true" pageId="13" pageNumber="414">24.</emphasis>
<bibRefCitation id="EFA34B648757FFFDFCE8F53210E3FAB2" author="Jung, P." box="[865,973,1288,1312]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
, p. 25. =
<taxonomicName id="4C324D168757FFFDFBA6F53216ADFAB2" authority="Vincent" authorityName="Vincent" box="[1071,1411,1288,1312]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="minus" subGenus="Seraphs">
<emphasis id="B946EA878757FFFDFBA6F53217AFFAB2" box="[1071,1153,1288,1312]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
(
<emphasis id="B946EA878757FFFDFB19F53217CCFAB2" box="[1168,1250,1288,1312]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
)
<emphasis id="B946EA878757FFFDFB78F5331600FAB2" box="[1265,1326,1289,1312]" italics="true" pageId="13" pageNumber="414">minus</emphasis>
Vincent
</taxonomicName>
—
<bibRefCitation id="EFA34B648757FFFDFF4EF51D121CFAAD" author="Jung, P." box="[199,306,1319,1343]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
, p. 25, pl. 6, figs. 9 &amp; 10.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF1EF55711D6FA39" pageId="13" pageNumber="414" type="materials_examined">
<paragraph id="8B8D36958757FFFDFF1EF5571656FA15" blockId="13.[151,1436,1389,1775]" box="[151,1400,1389,1415]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF1EF557126BFA15" bold="true" box="[151,325,1389,1415]" pageId="13" pageNumber="414">Type material.</emphasis>
<typeStatus id="548988378757FFFDFEC5F557129CFA15" box="[332,434,1389,1415]" pageId="13" pageNumber="414" type="holotype">Holotype</typeStatus>
—
<bibRefCitation id="EFA34B648757FFFDFE46F5571147FA15" author="Vincent, E." box="[463,617,1389,1415]" pageId="13" pageNumber="414" pagination="1 - 46" refId="ref42812" refString="Vincent, E. (1913) Contribution a la paleontologie dea falaises de Landana (Bas-Congo), Mollusques. Annales du Musee du Congo, 1, 1 - 46." type="journal article" year="1913">
Vincent
<emphasis id="B946EA878757FFFDFDA7F5571147FA15" box="[558,617,1389,1415]" italics="true" pageId="13" pageNumber="414">1913</emphasis>
</bibRefCitation>
, pl. 1, figs. 23, 24. The type measures
<quantity id="4CCA9B708757FFFDFB9EF5571776FA15" box="[1047,1112,1389,1415]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="414" unit="mm" value="8.0">8 mm</quantity>
in length (Vincent 2013).
</paragraph>
<paragraph id="8B8D36958757FFFDFF4EF5AB11D6FA39" blockId="13.[151,1436,1389,1775]" box="[199,760,1424,1451]" pageId="13" pageNumber="414">
<materialsCitation id="3B5A3CC88757FFFDFF4EF5AB11DAFA39" ID-GBIF-Occurrence="3320081322" box="[199,756,1424,1451]" country="Angola" location="Landra" pageId="13" pageNumber="414" specimenCount="1" typeStatus="holotype">
<emphasis id="B946EA878757FFFDFF4EF5AB1248FA39" bold="true" box="[199,358,1425,1451]" pageId="13" pageNumber="414">
<typeStatus id="548988378757FFFDFF4EF5AB122CFA39" box="[199,258,1425,1451]" pageId="13" pageNumber="414">Type</typeStatus>
locality.
</emphasis>
<location id="8EED604E8757FFFDFEE5F5AB1290FA39" LSID="urn:lsid:plazi:treatment:039B87838757FFFDFF1EF4A51153F97D:8EED604E8757FFFDFEE5F5AB1290FA39" box="[364,446,1425,1451]" country="Angola" name="Landra" pageId="13" pageNumber="414">Landra</location>
,
<collectingCountry id="F32576058757FFFDFE43F5AB1104FA39" box="[458,554,1425,1451]" name="Angola" pageId="13" pageNumber="414">Cabinda</collectingCountry>
,
<collectingCountry id="F32576058757FFFDFDBAF5AB11ABFA39" box="[563,645,1425,1451]" name="Angola" pageId="13" pageNumber="414">Angola</collectingCountry>
[Danian]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF4EF58F12D0F985" pageId="13" pageNumber="414" type="diagnosis">
<paragraph id="8B8D36958757FFFDFF4EF58F12D0F985" blockId="13.[151,1436,1389,1775]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF58F1214FA5D" bold="true" box="[199,314,1461,1487]" pageId="13" pageNumber="414">Diagnosis</emphasis>
. Known only from immature specimens with the features of juvenile
<taxonomicName id="4C324D168757FFFDFBEBF58F1792FA5D" authorityName="Montfort" authorityYear="1810" box="[1122,1212,1461,1487]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878757FFFDFBEBF58F1792FA5D" box="[1122,1212,1461,1487]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
</taxonomicName>
, with smooth body whorl and very little ornamentation. This species can be recognised as the only member of the
<taxonomicName id="4C324D168757FFFDFB33F5E3166FFA61" authorityName="Jung" authorityYear="1974" box="[1210,1345,1497,1523]" class="Gastropoda" family="Seraphsidae" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="family">Seraphsidae</taxonomicName>
coming from
<collectingCountry id="F32576058757FFFDFF5AF5C7120BF985" box="[211,293,1533,1559]" name="Angola" pageId="13" pageNumber="414">Angola</collectingCountry>
during the Danian.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF4EF61B10C1F9A9" box="[199,1007,1569,1595]" pageId="13" pageNumber="414" type="distribution">
<paragraph id="8B8D36958757FFFDFF4EF61B10C1F9A9" blockId="13.[151,1436,1389,1775]" box="[199,1007,1569,1595]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF61B124FF9A9" bold="true" box="[199,353,1569,1595]" pageId="13" pageNumber="414">Distribution.</emphasis>
? PALEOCENE—Danian—
<collectingCountry id="F32576058757FFFDFD2DF61B1077F9A9" box="[676,857,1569,1595]" name="Angola Cabinda" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFD2DF61B11DBF9A9" box="[676,757,1569,1595]" italics="true" pageId="13" pageNumber="414">Angola</emphasis>
Cabinda
</collectingCountry>
(
<bibRefCitation id="EFA34B648757FFFDFCE1F61B10CCF9A9" author="Jung, P." box="[872,994,1569,1595]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C328651E8757FFFDFF4EF67F1153F97D" pageId="13" pageNumber="414" type="discussion">
<paragraph id="8B8D36958757FFFDFF4EF67F1153F97D" blockId="13.[151,1436,1389,1775]" pageId="13" pageNumber="414">
<emphasis id="B946EA878757FFFDFF4EF67F1215F9CD" bold="true" box="[199,315,1605,1631]" pageId="13" pageNumber="414">Remarks.</emphasis>
<bibRefCitation id="EFA34B648757FFFDFECAF67F12E4F9CD" author="Jung, P." box="[323,458,1605,1631]" pageId="13" pageNumber="414" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung (1974)</bibRefCitation>
noted that the
<taxonomicName id="4C324D168757FFFDFDFAF67F1092F9CD" authority="Vincent, 1913" authorityName="Vincent" authorityYear="1913" box="[627,956,1605,1631]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="species" species="minus">
<emphasis id="B946EA878757FFFDFDFAF67F1036F9CD" box="[627,792,1605,1631]" italics="true" pageId="13" pageNumber="414">Seraphs minus</emphasis>
<bibRefCitation id="EFA34B648757FFFDFC96F67F1092F9CD" author="Vincent, E." box="[799,956,1605,1631]" pageId="13" pageNumber="414" pagination="1 - 46" refId="ref42812" refString="Vincent, E. (1913) Contribution a la paleontologie dea falaises de Landana (Bas-Congo), Mollusques. Annales du Musee du Congo, 1, 1 - 46." type="journal article" year="1913">Vincent, 1913</bibRefCitation>
</taxonomicName>
<typeStatus id="548988378757FFFDFC4DF67F10DAF9CD" box="[964,1012,1605,1631]" pageId="13" pageNumber="414">type</typeStatus>
material could not be located, and he recommended that the taxon should be considered a
<emphasis id="B946EA878757FFFDFD68F65010A8F911" box="[737,902,1641,1667]" italics="true" pageId="13" pageNumber="414">nomen dubium</emphasis>
as the morphology is, at present, best described as unclear, the
<typeStatus id="548988378757FFFDFEB6F6B71241F935" box="[319,367,1677,1703]" pageId="13" pageNumber="414">type</typeStatus>
material being a small juvenile specimen. However, given the restricted geographic range, and as the only representative of
<taxonomicName id="4C324D168757FFFDFE56F68B11C0F959" authority="Montfort, 1810" authorityName="Montfort" authorityYear="1810" box="[479,750,1713,1739]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="414" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878757FFFDFE56F68B1117F959" box="[479,569,1713,1739]" italics="true" pageId="13" pageNumber="414">Seraphs</emphasis>
<bibRefCitation id="EFA34B648757FFFDFDCBF68B11C0F959" author="Montfort, D." box="[578,750,1713,1739]" pageId="13" pageNumber="414" refId="ref39226" refString="Montfort, D. (1810) Conchyliologie Systematique, et Classification Methodique Des Coquille. Vol. II. F. Schoell, Paris, 675 pp. https: // doi. org / 10.5962 / bhl. title. 10571" type="book" year="1810">Montfort, 1810</bibRefCitation>
</taxonomicName>
from this region in Africa, it is herein considered taxonomically significant and worthy of recognition.
</paragraph>
</subSubSection>
</treatment>
</document>
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