609 lines
73 KiB
XML
609 lines
73 KiB
XML
<document id="D3AFCD9156CF83821057E5981EC3E8E7" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana & Grossi, Paschoal C." docDate="2022" docId="CF46879EFFA7FF84FE47775FFBB529C2" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gibboryctes szelenyii Endrodi 1974" docType="treatment" docVersion="7" lastPageNumber="210" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="203" updateTime="1699352275692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="99A4653CAE34908B82585BE0F7DCBBF6" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5C4BBCC2FF4D2F1CD3AE13B5D48EE452">
|
||
<mods:title id="065F2B2913A72C53FD3A8757C383D04C">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="BCCE3AF4B2DF59F62DA74080DEBA1486" type="personal">
|
||
<mods:role id="B078C1C15CAFBB9F422AF21850DF2AC0">
|
||
<mods:roleTerm id="3590D1C5C2A1F343763124A591C337E7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1FD980494782EEBBBB71C8422F360BB3">Costa, Leidiane O.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="4949057CBADDA34035EFDC1C2BB0CB6A" type="personal">
|
||
<mods:role id="96D02B363F18970369E3A091F871A084">
|
||
<mods:roleTerm id="C543598CC9041440C3A0769555235344">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C6B3D141EFEA248D43D7A3F1EB0EF085">Duarte, Paulo R. M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7EAFCD7AD572DE2F44E5B1714E737956" type="personal">
|
||
<mods:role id="3A2E5750C4F120C58001FD282C9F221A">
|
||
<mods:roleTerm id="56298F0BC9ED998BB3726852D1C7D648">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="262B1035E5EDCB018D33BD89DDC38D0E">Iannuzzi, Luciana</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="524EDECCBDE9DECDC79B8940F3337A5A" type="personal">
|
||
<mods:role id="926BFD3B8FE83C04B7FDA3C968AB4F21">
|
||
<mods:roleTerm id="E28963ED0199374BE94B995E87E18AE6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0AB89491E109B576BF3871DDB31C57C4">Grossi, Paschoal C.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9AF56C7340B74489885E173BDD34899A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="49BE2A7B70A20255D86DBEED1CFD200A" type="host">
|
||
<mods:titleInfo id="FA6CD6B3361DA3037D5C47ACC8F21A66">
|
||
<mods:title id="8E8C5A654C830D54E5277266746083D1">Journal of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="0BB7A132A9144D64539B3A445B7B5D1A">
|
||
<mods:date id="8DB60CC034941231DA25550E4818711D">2022</mods:date>
|
||
<mods:detail id="F2E15DCAAD76853B9DC31259BCF52FB9" type="pubDate">
|
||
<mods:number id="2F17F6BA2DB30EFD9971653F41C962A1">2022-05-17</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B7B005285F898618B5BBE1A2FE87EE87" type="volume">
|
||
<mods:number id="8942B0C4C4D15A5DC05BDB257089DA87">56</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DE3EE7A494A0882A88F1A165E89686BD" type="issue">
|
||
<mods:number id="CB8AC0F19B5B83B62A580A1EACF65265">1 - 4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9EBFB8643985D744520F173E0F00F102" unit="page">
|
||
<mods:start id="07B4362315DEE25729C58C237FDB5565">191</mods:start>
|
||
<mods:end id="CD60EFD757A10361610BB271B4636888">225</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="0C53B11745F6788EB791644C3459B9A4">
|
||
<mods:url id="C3510286F9EC6F9452A6AEDD4565F3EA">http://dx.doi.org/10.1080/00222933.2021.2017499</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="C0BDF83668810FBA1AC20569FE892DC8">journal article</mods:classification>
|
||
<mods:identifier id="D9F029336A9FC47358470A7474FE14B3" type="CLB-Dataset">82886</mods:identifier>
|
||
<mods:identifier id="B3A87347910CB248C8E869A45FB08FC1" type="DOI">10.1080/00222933.2021.2017499</mods:identifier>
|
||
<mods:identifier id="491CB75582AB0AEE713A1A5842C7B2A4" type="GBIF-Dataset">5cd01cfe-52bb-4092-97b2-804c3c7d145d</mods:identifier>
|
||
<mods:identifier id="CD732BEC5B96E8AE8072A84630EDC1C7" type="ISSN">1464-5262</mods:identifier>
|
||
<mods:identifier id="79A53CDD03F374A1B658CC79744E97A2" type="Zenodo-Dep">6758313</mods:identifier>
|
||
<mods:identifier id="57022FFBBD835E4BDDE1B686FD71F286" type="ZooBank">D766FC5E-CEC8-4184-BE90-C25350B90612</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CF46879EFFA7FF84FE47775FFBB529C2" ID-DOI="http://doi.org/10.5281/zenodo.6761924" ID-GBIF-Taxon="196400130" ID-Zenodo-Dep="6761924" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2" httpUri="http://treatment.plazi.org/id/CF46879EFFA7FF84FE47775FFBB529C2" lastPageId="20" lastPageNumber="210" pageId="13" pageNumber="203">
|
||
<subSubSection id="0FF56503FFA7FF9DFE47775FFCF42F58" box="[434,881,1326,1352]" pageId="13" pageNumber="203" type="nomenclature">
|
||
<paragraph id="47503688FFA7FF9DFE47775FFCF42F58" blockId="13.[195,1121,1326,1421]" box="[434,881,1326,1352]" pageId="13" pageNumber="203">
|
||
<heading id="1C1881E4FFA7FF9DFE47775FFCF42F58" box="[434,881,1326,1352]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="203" reason="2">
|
||
<taxonomicName id="80EF4D0BFFA7FF9DFE47775FFCF42F58" authority="Endrodi, 1974" authorityName="Endrodi" authorityYear="1974" box="[434,881,1326,1352]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA7FF9DFE47775FFCF42F58" bold="true" box="[434,881,1326,1352]" pageId="13" pageNumber="203">
|
||
Gibboryctes szelenyii
|
||
<bibRefCitation id="237E4B79FFA7FF9DFD33775FFCF42F58" author="Endrodi S" box="[710,881,1326,1352]" pageId="13" pageNumber="203" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">
|
||
<emphasis id="759BEA9AFFA7FF9DFD33775FFCF42F58" bold="true" box="[710,881,1326,1352]" italics="true" pageId="13" pageNumber="203">Endrödi, 1974</emphasis>
|
||
</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA7FF9DFF367720FBE42F7B" box="[195,1121,1361,1387]" pageId="13" pageNumber="203" type="description">
|
||
<paragraph id="47503688FFA7FF9DFF367720FBE42F7B" blockId="13.[195,1121,1326,1421]" box="[195,1121,1361,1387]" pageId="13" pageNumber="203">
|
||
<heading id="1C1881E4FFA7FF9DFF367720FBE42F7B" bold="true" box="[195,1121,1361,1387]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="203" reason="3">
|
||
<emphasis id="759BEA9AFFA7FF9DFF367720FBE42F7B" bold="true" box="[195,1121,1361,1387]" italics="true" pageId="13" pageNumber="203">
|
||
(
|
||
<figureCitation id="DFD42A0DFFA7FF9DFF3E7720FEBE2F7B" box="[203,315,1361,1387]" captionStart="Figure 1" captionStartId="4.[160,227,1366,1389]" captionTargetBox="[162,1148,150,1331]" captionTargetId="figure-77@4.[162,1154,142,1335]" captionTargetPageId="4" captionText="Figure 1. Gibboryctes szelenyii Endrödi, 1974. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale." figureDoi="http://doi.org/10.5281/zenodo.6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="13" pageNumber="203">Figures 1</figureCitation>
|
||
;
|
||
<figureCitation id="DFD42A0DFFA7FF9DFEBF7720FEDD2F7B" box="[330,344,1361,1387]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="13" pageNumber="203">6</figureCitation>
|
||
(a); 7(a); 8(a); 9(a); 10(a–b); 11(a); 12(a–b); 13(a); 14(b); 15; 16)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA7FF9DFE6E7702FC022F9D" box="[411,903,1395,1421]" pageId="13" pageNumber="203" type="reference_group">
|
||
<paragraph id="47503688FFA7FF9DFE6E7702FC022F9D" blockId="13.[195,1121,1326,1421]" box="[411,903,1395,1421]" pageId="13" pageNumber="203">
|
||
<heading id="1C1881E4FFA7FF9DFE6E7702FC022F9D" box="[411,903,1395,1421]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="203" reason="2">
|
||
<treatmentCitationGroup id="67FF11A6FFA7FF9DFE6E7702FC022F9D" box="[411,903,1395,1421]" pageId="13" pageNumber="203">
|
||
<emphasis id="759BEA9AFFA7FF9DFE6E7702FC022F9D" bold="true" box="[411,903,1395,1421]" pageId="13" pageNumber="203">
|
||
<taxonomicName id="80EF4D0BFFA7FF9DFE6E7702FD2D2F9D" authorityName="Endrodi" authorityYear="1974" box="[411,680,1395,1421]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="szelenyii">Gibboryctes szelenyii</taxonomicName>
|
||
.
|
||
<treatmentCitation id="C64E1099FFA7FF9DFD427702FC022F9D" author="Endrodi S" box="[695,903,1395,1421]" page="14" pageId="13" pageNumber="203" year="1974">
|
||
<bibRefCitation id="237E4B79FFA7FF9DFD427702FC022F9D" author="Endrodi S" box="[695,903,1395,1421]" pageId="13" pageNumber="203" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">
|
||
<emphasis id="759BEA9AFFA7FF9DFD427702FC022F9D" bold="true" box="[695,903,1395,1421]" italics="true" pageId="13" pageNumber="203">Endrödi 1974: 14</emphasis>
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
</emphasis>
|
||
</treatmentCitationGroup>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA7FF9DFF5577D9FD682C60" pageId="13" pageNumber="203" type="diagnosis">
|
||
<paragraph id="47503688FFA7FF9DFF5577D9FE912FD2" blockId="13.[160,1156,1448,1648]" box="[160,276,1448,1474]" pageId="13" pageNumber="203">
|
||
<emphasis id="759BEA9AFFA7FF9DFF5577D9FE912FD2" bold="true" box="[160,276,1448,1474]" italics="true" pageId="13" pageNumber="203">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFA7FF9DFF5577BDFD682C60" blockId="13.[160,1156,1448,1648]" pageId="13" pageNumber="203">
|
||
<taxonomicName id="80EF4D0BFFA7FF9DFF5577BDFE022FF5" authorityName="Endrodi" authorityYear="1974" box="[160,391,1484,1509]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="203" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA7FF9DFF5577BDFE022FF5" box="[160,391,1484,1509]" italics="true" pageId="13" pageNumber="203">Gibboryctes szelenyii</emphasis>
|
||
</taxonomicName>
|
||
differs from the other two
|
||
<taxonomicName id="80EF4D0BFFA7FF9DFD2277BDFC312FF5" box="[727,948,1484,1509]" pageId="13" pageNumber="203">
|
||
<emphasis id="759BEA9AFFA7FF9DFD2277BDFCD22FF5" box="[727,855,1484,1509]" italics="true" pageId="13" pageNumber="203">Gibboryctes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the following combination of characters: labrum subtriangular, weakly rounded on apical margin (
|
||
<figureCitation id="DFD42A0DFFA7FF9DFF5D7460FE8F2C3A" box="[168,266,1553,1578]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="13" pageNumber="203">Figure 6</figureCitation>
|
||
(a)); maxillary galea triangular (
|
||
<figureCitation id="DFD42A0DFFA7FF9DFD887460FD5A2C3A" box="[637,735,1553,1578]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="13" pageNumber="203">Figure 8</figureCitation>
|
||
(a)); punctures on pronotal anterior corners predominantly contiguous (
|
||
<figureCitation id="DFD42A0DFFA7FF9DFDB07445FD372C5D" box="[581,690,1588,1613]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="13" pageNumber="203">Figures 9</figureCitation>
|
||
(a) and
|
||
<figureCitation id="DFD42A0DFFA7FF9DFCE57445FC092C5D" box="[784,908,1588,1613]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="13" pageNumber="203">Figures 12</figureCitation>
|
||
(a)); elytral interstriae densely covered with large punctures (
|
||
<figureCitation id="DFD42A0DFFA7FF9DFDA27426FD462C60" box="[599,707,1623,1648]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="13" pageNumber="203">Figure 12</figureCitation>
|
||
(b)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13906600FFA4FF9EFF5577D0FB912FE3" ID-DOI="http://doi.org/10.5281/zenodo.6758341" ID-Zenodo-Dep="6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="14" pageNumber="204" startId="14.[160,225,1441,1464]" targetBox="[168,1146,144,1407]" targetPageId="14">
|
||
<paragraph id="47503688FFA4FF9EFF5577D0FB912FE3" blockId="14.[160,1156,1441,1523]" pageId="14" pageNumber="204">
|
||
<emphasis id="759BEA9AFFA4FF9EFF5577D0FE8C2FA8" bold="true" box="[160,265,1441,1464]" pageId="14" pageNumber="204">Figure 11.</emphasis>
|
||
Detail of head, pronotum and prothoracic legs of females in dorsal view: (a)
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFBE077D0FEC62FC6" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA4FF9EFBE077D0FF6A2FC6" italics="true" pageId="14" pageNumber="204">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFE9E77CEFDB62FC6" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[363,563,1471,1494]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA4FF9EFE9E77CEFDB62FC6" box="[363,563,1471,1494]" italics="true" pageId="14" pageNumber="204">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA4FF9EFDCD77CEFD032FC6" box="[568,646,1471,1494]" pageId="14" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFD5E77CEFCF42FC6" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[683,881,1471,1494]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="endroedi" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA4FF9EFD5E77CEFCF42FC6" box="[683,881,1471,1494]" italics="true" pageId="14" pageNumber="204">Gibboryctes endroedi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA4FF9EFC8277CEFC402FC6" box="[887,965,1471,1494]" pageId="14" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFC1977CEFEC22FE3" authority="Prell" authorityName="Prell" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFA4FF9EFC1977CEFE972FE3" italics="true" pageId="14" pageNumber="204">Heterogomphus gracilicornis</emphasis>
|
||
Prell
|
||
</taxonomicName>
|
||
; (e)
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFE8477ADFD1C2FE3" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" box="[369,665,1500,1523]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA4FF9EFE8477ADFDB02FE3" box="[369,565,1500,1523]" italics="true" pageId="14" pageNumber="204">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
. Arrow indicates punctate frons in (b).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FF56503FFA4FF9EFF557461FC112C82" pageId="14" pageNumber="204" type="materials_examined">
|
||
<paragraph id="47503688FFA4FF9EFF557461FE722C3A" blockId="14.[160,1155,1552,1682]" box="[160,503,1552,1578]" pageId="14" pageNumber="204">
|
||
<emphasis id="759BEA9AFFA4FF9EFF557461FE722C3A" bold="true" box="[160,503,1552,1578]" italics="true" pageId="14" pageNumber="204">
|
||
Type material (
|
||
<figureCitation id="DFD42A0DFFA4FF9EFEA17461FE352C3A" box="[340,432,1552,1578]" captionStart="Figure 1" captionStartId="4.[160,227,1366,1389]" captionTargetBox="[162,1148,150,1331]" captionTargetId="figure-77@4.[162,1154,142,1335]" captionTargetPageId="4" captionText="Figure 1. Gibboryctes szelenyii Endrödi, 1974. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale." figureDoi="http://doi.org/10.5281/zenodo.6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="14" pageNumber="204">Figure 1</figureCitation>
|
||
(a–c))
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFA4FF9EFF557442FC112C82" blockId="14.[160,1155,1552,1682]" pageId="14" pageNumber="204">
|
||
<typeStatus id="9854882AFFA4FF9EFF557442FE822C5C" box="[160,263,1587,1612]" pageId="14" pageNumber="204" type="holotype">Holotype</typeStatus>
|
||
male, labelled: (a) ‘
|
||
<typeStatus id="9854882AFFA4FF9EFE117442FDDE2C5C" box="[484,603,1587,1612]" pageId="14" pageNumber="204" type="holotype">Holotypus</typeStatus>
|
||
/
|
||
<emphasis id="759BEA9AFFA4FF9EFD977442FCC62C5C" box="[610,835,1587,1612]" italics="true" pageId="14" pageNumber="204">
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFD977442FD5B2C5C" box="[610,734,1587,1612]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="genus">Gibboryctes</taxonomicName>
|
||
/
|
||
<taxonomicName id="80EF4D0BFFA4FF9EFD1C7442FCC62C5C" authorityName="Endrodi" authorityYear="1974" box="[745,835,1587,1612]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="204" phylum="Arthropoda" rank="species" species="szelenyii">szelenyii</taxonomicName>
|
||
</emphasis>
|
||
/Endr’. [old white label, bordered in red, partially handwritten]; (b) ‘Vidit 1976/R. -P. Dechambre’ [white label, partially handwritten]; (c) ‘Paraguay’ [old white label, handwritten] (
|
||
<collectionCode id="21FEAE4DFFA4FF9EFCC37408FC0C2C82" box="[822,905,1657,1682]" country="USA" name="John May Museum of Natural History" pageId="14" pageNumber="204">MNHM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13906600FFA5FF9FFF5576D4FE342F04" ID-DOI="http://doi.org/10.5281/zenodo.6758343" ID-Zenodo-Dep="6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="15" pageNumber="205" startId="15.[160,225,1189,1212]" targetBox="[167,1149,146,1154]" targetPageId="15">
|
||
<paragraph id="47503688FFA5FF9FFF5576D4FE342F04" blockId="15.[160,1156,1189,1300]" pageId="15" pageNumber="205">
|
||
<emphasis id="759BEA9AFFA5FF9FFF5576D4FE802EAC" bold="true" box="[160,261,1189,1212]" pageId="15" pageNumber="205">Figure 12.</emphasis>
|
||
Females of
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFE8C76D4FDB12EAC" box="[377,564,1189,1212]" pageId="15" pageNumber="205">
|
||
<emphasis id="759BEA9AFFA5FF9FFE8C76D4FE6D2EAC" box="[377,488,1189,1212]" italics="true" pageId="15" pageNumber="205">Gibboryctes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b)
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFD4F76B3FC522EC9" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[698,983,1218,1241]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="205" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA5FF9FFD4F76B3FC052EC9" box="[698,896,1218,1241]" italics="true" pageId="15" pageNumber="205">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (c,d)
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFBE076B3FF732EE7" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="205" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA5FF9FFBE076B3FF732EE7" italics="true" pageId="15" pageNumber="205">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA5FF9FFEF47691FED32EE7" box="[257,342,1248,1271]" pageId="15" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (e,f)
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFE607691FDEF2EE7" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[405,618,1248,1271]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="205" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA5FF9FFE607691FDEF2EE7" box="[405,618,1248,1271]" italics="true" pageId="15" pageNumber="205">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA5FF9FFD807691FD4D2EE7" box="[629,712,1248,1271]" pageId="15" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FF56503FFA5FF9FFF55773FFEFC2F78" box="[160,377,1358,1384]" pageId="15" pageNumber="205" type="description">
|
||
<paragraph id="47503688FFA5FF9FFF55773FFEFC2F78" blockId="15.[160,1156,1358,1627]" box="[160,377,1358,1384]" pageId="15" pageNumber="205">
|
||
<emphasis id="759BEA9AFFA5FF9FFF55773FFEFC2F78" bold="true" box="[160,377,1358,1384]" italics="true" pageId="15" pageNumber="205">Non-type material</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA5FF80FF557703FEC02F1C" lastPageId="16" lastPageNumber="206" pageId="15" pageNumber="205" type="materials_examined">
|
||
<paragraph id="47503688FFA5FF80FF557703FEC02F1C" blockId="15.[160,1156,1358,1627]" lastBlockId="16.[160,1155,1198,1292]" lastPageId="16" lastPageNumber="206" pageId="15" pageNumber="205">
|
||
<materialsCitation id="F7873CD5FFA5FF9FFF557703FE642FBE" ID-GBIF-Occurrence="3819562311" collectingDate="2004-01" collectorName="E. J. Grossi & P. C. Grossi" country="Brazil" location="Brasilia" pageId="15" pageNumber="205" specimenCount="1">
|
||
<collectingCountry id="3FF87618FFA5FF9FFF557703FF772F9B" box="[160,242,1394,1419]" name="Brazil" pageId="15" pageNumber="205">BRAZIL</collectingCountry>
|
||
:
|
||
<location id="42306053FFA5FF9FFF097703FEC82F9B" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFF097703FEC82F9B" box="[252,333,1394,1419]" country="Brazil" name="Brasilia" pageId="15" pageNumber="205">Brasília</location>
|
||
,
|
||
<location id="42306053FFA5FF9FFEA27703FD802F9B" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFEA27703FD802F9B" box="[343,517,1394,1419]" country="Brazil" name="Distrito Federal" pageId="15" pageNumber="205">Distrito Federal</location>
|
||
,
|
||
<location id="42306053FFA5FF9FFDE57703FD032F9B" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFDE57703FD032F9B" box="[528,646,1394,1419]" country="Brazil" name="Brazlandia" pageId="15" pageNumber="205">Brazlândia</location>
|
||
,
|
||
<location id="42306053FFA5FF9FFD647703FC022F9B" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFD647703FC022F9B" box="[657,903,1394,1419]" country="Brazil" name="Fazenda Sao Joaquim" pageId="15" pageNumber="205">Fazenda São Joaquim</location>
|
||
,
|
||
<date id="33511048FFA5FF9FFC677703FBAC2F9B" box="[914,1065,1394,1419]" pageId="15" pageNumber="205" value="2004-01">
|
||
<collectingDate id="2315E9A0FFA5FF9FFC677703FBAC2F9B" box="[914,1065,1394,1419]" pageId="15" pageNumber="205" value="2004-01">January 2004</collectingDate>
|
||
</date>
|
||
, luz,
|
||
<collectorName id="EA1A535EFFA5FF9FFB947703FF602FBE" pageId="15" pageNumber="205">
|
||
E.J.
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFF5577E4FF602FBE" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[160,229,1429,1454]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
</collectorName>
|
||
and
|
||
<collectorName id="EA1A535EFFA5FF9FFEDC77E4FE212FBE" box="[297,420,1429,1454]" pageId="15" pageNumber="205">
|
||
P.C.
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFEAA77E4FE212FBE" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[351,420,1429,1454]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
</collectorName>
|
||
legs
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="F7873CD5FFA5FF9FFDFC77E4FC2D2FC0" ID-GBIF-Occurrence="3819562307" collectingDate="2011-10-28" collectingDateMax="2011-10-30" collectingDateMin="2011-10-28" collectionCode="CERPE" collectorName="Grossi & Parizotto & Leivas & Santos" country="Brazil" elevation="2010900" location="Parque Estadual do Guartela" municipality="Tibagi" pageId="15" pageNumber="205" specimenCount="1" specimenCount-female="1" specimenCount-male="8" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFA5FF9FFDFC77E4FDF52FBE" box="[521,624,1429,1454]" pageId="15" pageNumber="205" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFA5FF9FFD7777E4FD572FBE" box="[642,722,1429,1454]" collectionName="CERPE" pageId="15" pageNumber="205">CERPE</collectionCode>
|
||
).
|
||
<collectingRegion id="852BF86AFFA5FF9FFD1077E4FCB32FBE" box="[741,822,1429,1454]" country="Brazil" name="Parana" pageId="15" pageNumber="205">Paraná</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="A734ACF2FFA5FF9FFCB077E4FC0B2FBE" box="[837,910,1429,1454]" pageId="15" pageNumber="205">Tibagi</collectingMunicipality>
|
||
,
|
||
<location id="42306053FFA5FF9FFC6877E4FE872FC0" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFC6877E4FE872FC0" country="Brazil" municipality="Tibagi" name="Parque Estadual do Guartela" pageId="15" pageNumber="205" stateProvince="Parana">Parque Estadual do Guartelá</location>
|
||
,
|
||
<date id="33511048FFA5FF9FFEF877C6FE732FC0" box="[269,502,1463,1488]" pageId="15" pageNumber="205" value="2011-10-28" valueMax="2011-10-30" valueMin="2011-10-28">
|
||
<collectingDate id="2315E9A0FFA5FF9FFEF877C6FE732FC0" box="[269,502,1463,1488]" pageId="15" pageNumber="205" value="2011-10-28" valueMax="2011-10-30" valueMin="2011-10-28">28–30 October 2011</collectingDate>
|
||
</date>
|
||
,
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFDF777C6FDCC2FC0" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[514,585,1463,1488]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">
|
||
<collectorName id="EA1A535EFFA5FF9FFDF777C6FDCC2FC0" box="[514,585,1463,1488]" pageId="15" pageNumber="205">Grossi</collectorName>
|
||
</taxonomicName>
|
||
,
|
||
<collectorName id="EA1A535EFFA5FF9FFDA177C6FD3E2FC0" box="[596,699,1463,1488]" pageId="15" pageNumber="205">Parizotto</collectorName>
|
||
,
|
||
<collectorName id="EA1A535EFFA5FF9FFD3277C6FC8E2FC0" box="[711,779,1463,1488]" pageId="15" pageNumber="205">Leivas</collectorName>
|
||
and
|
||
<collectorName id="EA1A535EFFA5FF9FFCB277C6FC172FC0" box="[839,914,1463,1488]" pageId="15" pageNumber="205">Santos</collectorName>
|
||
–
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFA5FF9FFC4477C6FB8E2FE3" ID-GBIF-Occurrence="3819562302" collectingDate="2012-11" collectingDateMax="2012-11" collectingDateMin="2010-12" collectionCode="CERPE, CE-UFPE, EPGC" collectorName="Parizotto" country="Brazil" elevation="2010900" location="Parque Estadual do Guartela" municipality="Tibagi" pageId="15" pageNumber="205" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFA5FF9FFC4477C6FB8E2FC0" box="[945,1035,1463,1488]" pageId="15" pageNumber="205" type="male">7 males</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFA5FF9FFBE377C6FB062FC0" box="[1046,1155,1463,1488]" pageId="15" pageNumber="205" type="female">5 females</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFA5FF9FFF5077ABFF742FE3" box="[165,241,1498,1523]" collectionName="CERPE" pageId="15" pageNumber="205">CERPE</collectionCode>
|
||
,
|
||
<collectionCode id="21FEAE4DFFA5FF9FFF0B77ABFED92FE3" box="[254,348,1498,1523]" pageId="15" pageNumber="205">CE-UFPE</collectionCode>
|
||
and
|
||
<collectionCode id="21FEAE4DFFA5FF9FFE6877ABFE5B2FE3" box="[413,478,1498,1523]" pageId="15" pageNumber="205">EPGC</collectionCode>
|
||
);
|
||
<date id="33511048FFA5FF9FFE0477ABFD2F2FE3" box="[497,682,1498,1523]" pageId="15" pageNumber="205" value="2012-11">
|
||
<collectingDate id="2315E9A0FFA5FF9FFE0477ABFD2F2FE3" box="[497,682,1498,1523]" pageId="15" pageNumber="205" value="2012-11">November 2012</collectingDate>
|
||
</date>
|
||
, breed,
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFCFF77ABFCCB2FE3" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[778,846,1498,1523]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
and
|
||
<collectorName id="EA1A535EFFA5FF9FFC7A77ABFC762FE3" box="[911,1011,1498,1523]" pageId="15" pageNumber="205">Parizotto</collectorName>
|
||
–
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFA5FF9FFBE377ABFEA52C28" ID-GBIF-Occurrence="3819562312" collectingDate="2010-12" collectingDateMax="2012-11" collectingDateMin="2010-12" collectionCode="CERPE" collectorName="Nasutitermis" country="Brazil" elevation="2010900" location="Parizotto" municipality="Tibagi" pageId="15" pageNumber="205" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFA5FF9FFBE377ABFBE22FE3" box="[1046,1127,1498,1523]" pageId="15" pageNumber="205" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFA5FF9FFB8077ABFF6B2C06" pageId="15" pageNumber="205" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFA5FF9FFEF7778CFED72C06" box="[258,338,1533,1558]" collectionName="CERPE" pageId="15" pageNumber="205">CERPE</collectionCode>
|
||
);
|
||
<date id="33511048FFA5FF9FFE9D778CFDAD2C06" box="[360,552,1533,1558]" pageId="15" pageNumber="205" value="2010-12">
|
||
<collectingDate id="2315E9A0FFA5FF9FFE9D778CFDAD2C06" box="[360,552,1533,1558]" pageId="15" pageNumber="205" value="2010-12">
|
||
December
|
||
<quantity id="80179B6DFFA5FF9FFE18778CFDAD2C06" box="[493,552,1533,1558]" metricMagnitude="6" metricUnit="m" metricValue="2.0109" pageId="15" pageNumber="205" unit="m" value="2010900.0">2010</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
, 900 m, breeding in
|
||
<emphasis id="759BEA9AFFA5FF9FFCC3778CFC472C06" box="[822,962,1533,1558]" italics="true" pageId="15" pageNumber="205">
|
||
<collectorName id="EA1A535EFFA5FF9FFCC3778CFC472C06" box="[822,962,1533,1558]" pageId="15" pageNumber="205">Nasutitermis</collectorName>
|
||
</emphasis>
|
||
sp.,
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFBF1778CFBCF2C06" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[1028,1098,1533,1558]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
and
|
||
<location id="42306053FFA5FF9FFF55746EFE822C28" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFF55746EFE822C28" box="[160,263,1567,1592]" country="Brazil" municipality="Tibagi" name="Parizotto" pageId="15" pageNumber="205" stateProvince="Parana">Parizotto</location>
|
||
–
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFA5FF9FFED9746EFC3C2C4B" ID-GBIF-Occurrence="3819562324" collectingDate="2011-12-12" collectingDateMax="2012-11" collectingDateMin="2010-12" collectionCode="CERPE" collectorName="Nasutitermis" country="Brazil" elevation="2010900" location="Parque Estadual de Vila Velha" municipality="Ponta Grossa" pageId="15" pageNumber="205" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFA5FF9FFED9746EFE052C28" box="[300,384,1567,1592]" pageId="15" pageNumber="205" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFA5FF9FFE65746EFE732C28" box="[400,502,1567,1592]" pageId="15" pageNumber="205" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFA5FF9FFDFD746EFDDD2C28" box="[520,600,1567,1592]" collectionName="CERPE" pageId="15" pageNumber="205">CERPE</collectionCode>
|
||
).
|
||
<collectingMunicipality id="A734ACF2FFA5FF9FFD99746EFC8C2C28" box="[620,777,1567,1592]" pageId="15" pageNumber="205">Ponta Grossa</collectingMunicipality>
|
||
,
|
||
<location id="42306053FFA5FF9FFCED746EFB052C28" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFCED746EFB052C28" box="[792,1152,1567,1592]" country="Brazil" municipality="Ponta Grossa" name="Parque Estadual de Vila Velha" pageId="15" pageNumber="205" stateProvince="Parana">Parque Estadual de Vila Velha</location>
|
||
,
|
||
<date id="33511048FFA5FF9FFF557433FEF32C4B" box="[160,374,1602,1627]" pageId="15" pageNumber="205" value="2011-12-12">
|
||
<collectingDate id="2315E9A0FFA5FF9FFF557433FEF32C4B" box="[160,374,1602,1627]" pageId="15" pageNumber="205" value="2011-12-12">12 December 2011</collectingDate>
|
||
</date>
|
||
, breeding in
|
||
<emphasis id="759BEA9AFFA5FF9FFDFC7433FD142C4B" box="[521,657,1602,1627]" italics="true" pageId="15" pageNumber="205">
|
||
<collectorName id="EA1A535EFFA5FF9FFDFC7433FD142C4B" box="[521,657,1602,1627]" pageId="15" pageNumber="205">Nasutitermis</collectorName>
|
||
</emphasis>
|
||
sp.,
|
||
<taxonomicName id="80EF4D0BFFA5FF9FFD317433FC8D2C4B" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[708,776,1602,1627]" genus="Grossi" pageId="15" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
and
|
||
<location id="42306053FFA5FF9FFCB57433FC212C4B" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFA5FF9FFCB57433FC212C4B" box="[832,932,1602,1627]" country="Brazil" municipality="Ponta Grossa" name="Parizotto" pageId="15" pageNumber="205" stateProvince="Parana">Parizotto</location>
|
||
–
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFA5FF80FC357433FDDD2EFA" ID-GBIF-Occurrence="3819562316" collectingDate="2010-12" collectingDateMax="2012-11" collectingDateMin="2010-12" collectionCode="CERPE" collectorName="Faz & Nasutitermis" country="Brazil" elevation="1088" lastPageId="16" lastPageNumber="206" latitude="-24.629" location="Novvember" longLatPrecision="7" longitude="-50.2152" municipality="Damasio" pageId="15" pageNumber="205" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFA5FF9FFC357433FB8B2C4B" box="[960,1038,1602,1627]" pageId="15" pageNumber="205" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFA5FF9FFBED7433FB062C4B" box="[1048,1155,1602,1627]" pageId="15" pageNumber="205" type="female">4 females</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFBAFF80FF5076DFFF762ED7" box="[165,243,1198,1223]" collectionName="CERPE" pageId="16" pageNumber="206">CERPE</collectionCode>
|
||
);
|
||
<collectorName id="EA1A535EFFBAFF80FEF676DFFEAE2ED7" box="[259,299,1198,1223]" pageId="16" pageNumber="206">Faz</collectorName>
|
||
. S.
|
||
<collectingMunicipality id="A734ACF2FFBAFF80FEBA76DFFE362ED7" box="[335,435,1198,1223]" pageId="16" pageNumber="206">Damásio</collectingMunicipality>
|
||
, 1
|
||
<location id="42306053FFBAFF80FE2676DFFDD32ED7" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFBAFF80FE2676DFFDD32ED7" box="[467,598,1198,1223]" country="Brazil" latitude="-24.629" longLatPrecision="7" longitude="-50.2152" municipality="Damasio" name="Novvember" pageId="16" pageNumber="206" stateProvince="Parana">Novvember</location>
|
||
2007,
|
||
<quantity id="80179B6DFFBAFF80FD5476DFFD722ED7" box="[673,759,1198,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" pageId="16" pageNumber="206" unit="m" value="1088.0">
|
||
<elevation id="CCC2D1BBFFBAFF80FD5476DFFD722ED7" box="[673,759,1198,1223]" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" pageId="16" pageNumber="206" unit="m" value="1088.0">1088 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="22DB504FFFBAFF80FCF676DFFCF62ED7" box="[771,883,1198,1223]" degrees="24.6290" direction="south" orientation="latitude" pageId="16" pageNumber="206" precision="5" value="-24.629">24.6290°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="22DB504FFFBAFF80FC8876DFFC732ED7" box="[893,1014,1198,1223]" degrees="50.2152" direction="west" orientation="longitude" pageId="16" pageNumber="206" precision="5" value="-50.2152">50.2152°W</geoCoordinate>
|
||
, breeding in
|
||
<emphasis id="759BEA9AFFBAFF80FF5576A0FEAD2EFA" box="[160,296,1233,1258]" italics="true" pageId="16" pageNumber="206">
|
||
<collectorName id="EA1A535EFFBAFF80FF5576A0FEAD2EFA" box="[160,296,1233,1258]" pageId="16" pageNumber="206">Nasutitermis</collectorName>
|
||
</emphasis>
|
||
sp.,
|
||
<taxonomicName id="80EF4D0BFFBAFF80FEAB76A0FE272EFA" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[350,418,1233,1258]" genus="Grossi" pageId="16" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
|
||
and
|
||
<location id="42306053FFBAFF80FE2B76A0FDC72EFA" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFBAFF80FE2B76A0FDC72EFA" box="[478,578,1233,1258]" country="Brazil" latitude="-24.629" longLatPrecision="7" longitude="-50.2152" municipality="Damasio" name="Parizotto" pageId="16" pageNumber="206" stateProvince="Parana">Parizotto</location>
|
||
–
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFBAFF80FD9476A0FC102EFA" ID-GBIF-Occurrence="3819562318" box="[609,917,1233,1258]" collectingDate="2010-12" collectingDateMax="2012-11" collectingDateMin="2010-12" collectionCode="CERPE" country="Brazil" elevation="2010900" latitude="-24.629" location="Parque Estadual do Guartela" longLatPrecision="7" longitude="-50.2152" municipality="Tibagi" pageId="16" pageNumber="206" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Parana">
|
||
<specimenCount id="51E9FD01FFBAFF80FD9476A0FD392EFA" box="[609,700,1233,1258]" pageId="16" pageNumber="206" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFBAFF80FD3276A0FCB12EFA" box="[711,820,1233,1258]" pageId="16" pageNumber="206" type="female">4 females</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFBAFF80FCB776A0FC152EFA" box="[834,912,1233,1258]" collectionName="CERPE" pageId="16" pageNumber="206">CERPE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFBAFF80FC5476A0FEC72F1C" ID-GBIF-Occurrence="3819562315" collectionCode="EPGC" country="Brazil" location="No" pageId="16" pageNumber="206" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="42306053FFBAFF80FC5476A0FC442EFA" LSID="urn:lsid:plazi:treatment:CF46879EFFA7FF84FE47775FFBB529C2:42306053FFBAFF80FC5476A0FC442EFA" box="[929,961,1233,1258]" country="Brazil" name="No" pageId="16" pageNumber="206">No</location>
|
||
data –
|
||
<specimenCount id="51E9FD01FFBAFF80FBEF76A0FBEC2EFA" box="[1050,1129,1233,1258]" pageId="16" pageNumber="206" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFBAFF80FB8076A0FF692F1C" pageId="16" pageNumber="206" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFBAFF80FF0C7682FEB92F1C" box="[249,316,1267,1292]" pageId="16" pageNumber="206">EPGC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13906600FFBAFF80FF557653FE6E2E63" ID-DOI="http://doi.org/10.5281/zenodo.6758345" ID-Zenodo-Dep="6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="16" pageNumber="206" startId="16.[160,225,1058,1081]" targetBox="[166,1146,145,1023]" targetPageId="16">
|
||
<paragraph id="47503688FFBAFF80FF557653FE6E2E63" blockId="16.[160,1155,1058,1139]" pageId="16" pageNumber="206">
|
||
<emphasis id="759BEA9AFFBAFF80FF557653FE832E29" bold="true" box="[160,262,1058,1081]" pageId="16" pageNumber="206">Figure 13.</emphasis>
|
||
Detail of ventrites I–VI of females. (a)
|
||
<taxonomicName id="80EF4D0BFFBAFF80FD8C7653FC142E29" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[633,913,1058,1081]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="206" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFBAFF80FD8C7653FCB82E29" box="[633,829,1058,1081]" italics="true" pageId="16" pageNumber="206">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFBAFF80FC4F7653FB062E29" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[954,1155,1058,1081]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="206" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFBAFF80FC4F7653FB062E29" box="[954,1155,1058,1081]" italics="true" pageId="16" pageNumber="206">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFBAFF80FF55764EFF6F2E46" box="[160,234,1087,1110]" pageId="16" pageNumber="206" rank="species">sp. nov</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFBAFF80FEE1764EFE662E46" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[276,483,1087,1110]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="206" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFBAFF80FEE1764EFE662E46" box="[276,483,1087,1110]" italics="true" pageId="16" pageNumber="206">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFBAFF80FE1F764EFDBF2E46" box="[490,570,1087,1110]" pageId="16" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFBAFF80FD90764EFC0B2E46" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" box="[613,910,1087,1110]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="206" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFBAFF80FD90764EFCAF2E46" box="[613,810,1087,1110]" italics="true" pageId="16" pageNumber="206">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
. Red arrows indicate the proportions of ventrites V and VI.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FF56503FFBAFF84FF557739FBCB2854" lastPageId="20" lastPageNumber="210" pageId="16" pageNumber="206" type="description">
|
||
<paragraph id="47503688FFBAFF80FF557739FE0C2F72" blockId="16.[160,1156,1352,1724]" box="[160,393,1352,1378]" pageId="16" pageNumber="206">
|
||
<emphasis id="759BEA9AFFBAFF80FF557739FE0C2F72" bold="true" box="[160,393,1352,1378]" pageId="16" pageNumber="206">
|
||
Male
|
||
<emphasis id="759BEA9AFFBAFF80FF167739FE0C2F72" bold="true" box="[227,393,1352,1378]" italics="true" pageId="16" pageNumber="206">
|
||
(
|
||
<figureCitation id="DFD42A0DFFBAFF80FF1E7739FEC22F72" box="[235,327,1352,1378]" captionStart="Figure 1" captionStartId="4.[160,227,1366,1389]" captionTargetBox="[162,1148,150,1331]" captionTargetId="figure-77@4.[162,1154,142,1335]" captionTargetPageId="4" captionText="Figure 1. Gibboryctes szelenyii Endrödi, 1974. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale." figureDoi="http://doi.org/10.5281/zenodo.6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="16" pageNumber="206">Figure 1</figureCitation>
|
||
(d,e))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFBAFF84FF55771AFCCC2AFD" blockId="16.[160,1156,1352,1724]" lastBlockId="20.[160,1154,142,237]" lastPageId="20" lastPageNumber="210" pageId="16" pageNumber="206">
|
||
<emphasis id="759BEA9AFFBAFF80FF55771AFECE2F95" bold="true" box="[160,331,1387,1413]" italics="true" pageId="16" pageNumber="206">Redescription.</emphasis>
|
||
<emphasis id="759BEA9AFFBAFF80FEAE771AFE352F94" bold="true" box="[347,432,1387,1412]" pageId="16" pageNumber="206">Length</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFBAFF80FE34771AFDE22F94" box="[449,615,1387,1412]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.56" metricValueMin="2.44" pageId="16" pageNumber="206" unit="mm" value="30.0" valueMax="35.6" valueMin="24.4">24.4–35.6 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFBAFF80FD82771AFD3A2F94" bold="true" box="[631,703,1387,1412]" pageId="16" pageNumber="206">Width</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFBAFF80FD24771AFCF22F94" box="[721,887,1387,1412]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" metricValueMax="1.53" metricValueMin="1.09" pageId="16" pageNumber="206" unit="mm" value="13.1" valueMax="15.3" valueMin="10.9">10.9–15.3 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFBAFF80FC73771AFC532F94" bold="true" box="[902,982,1387,1412]" pageId="16" pageNumber="206">Colour</emphasis>
|
||
: From reddish brown to dark reddish brown.
|
||
<emphasis id="759BEA9AFFBAFF80FE0477FFFDAA2FB7" bold="true" box="[497,559,1422,1447]" pageId="16" pageNumber="206">Head</emphasis>
|
||
: Clypeus subtriangular, transverse, slightly constricted laterally towards a bidentate (teeth inconspicuous) anterior margin; lateral margins slightly produced posteriorly; surface rugopunctate, with 2 moderately setose areas on sides. Frontoclypeal suture with a transverse, flattened, subtrapezoidal tubercle; tubercle sometimes emarginate at middle, scarcely setose laterally. Frons from flattened to slightly concave, glabrous, transversely rugopunctate; punctures large, deep, coalescent; interocular width equals 4.63 times transverse eye diameter. Ocular canthus transverse, subrectangular; anterior margin slightly arched; outer corner usually pointed.
|
||
<emphasis id="759BEA9AFFBAFF80FC3C74F0FBBB2C8A" bold="true" box="[969,1086,1665,1690]" pageId="16" pageNumber="206">Antennae</emphasis>
|
||
: Club subequal in length to antenomeres II–VII combined.
|
||
<emphasis id="759BEA9AFFBAFF80FD0174D2FCFB2CAC" bold="true" box="[756,894,1699,1724]" pageId="16" pageNumber="206">Mouthparts</emphasis>
|
||
: Labrum subtriangular, weakly rounded at apical margin, densely covered with a fringe of bristles dorsally (
|
||
<figureCitation id="DFD42A0DFFBBFF81FBDF77ADFB062FE5" box="[1066,1155,1500,1525]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="17" pageNumber="207">Figure 6</figureCitation>
|
||
(a)). Mandibles with 2 lobed teeth separated by a shallow, wide notch (
|
||
<figureCitation id="DFD42A0DFFBBFF81FC5B778FFB822C07" box="[942,1031,1534,1559]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (a–e), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="17" pageNumber="207">Figure 7</figureCitation>
|
||
(a)). Maxilla with triangular galea (
|
||
<figureCitation id="DFD42A0DFFBBFF81FE5D7450FD8C2C2A" box="[424,521,1569,1594]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="17" pageNumber="207">Figure 8</figureCitation>
|
||
(a)); inner margin with 2–3 small teeth; stipe laterally expanded in triangular shape. Labium oval, surrounded with large, deep, coalescent bristled punctures; discal area with bristled punctures scattered about 3 diameters of punctures.
|
||
<emphasis id="759BEA9AFFBBFF81FEE974F8FEEB2CB2" bold="true" box="[284,366,1673,1698]" pageId="17" pageNumber="207">Thorax</emphasis>
|
||
: Pronotum with complete borders in dorsal view; middle area close to anterior margin with a small, transverse tubercle; dorsal surface densely punctate; anterior corners with large, deep, contiguous punctures (
|
||
<figureCitation id="DFD42A0DFFB8FF82FD4D7790FC972FEA" box="[696,786,1505,1530]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="18" pageNumber="208">Figure 9</figureCitation>
|
||
(a)); lateral punctures large, deep, from contiguous to separate about 1 diameter of punctures; punctures gradually becoming small, shallow, sparser towards posterior area. Scutellar plate usually subtriangular, anteriorly covered with deep, large, irregularly scattered punctures. Elytral surface usually with 9 well-marked striae; each stria covered with large, deep, ocellate, contiguous punctures; interstriae with mixed fine and large punctures, irregularly scattered from anterior to posterior elytral area.
|
||
<emphasis id="759BEA9AFFB9FF83FE6A76C1FE502ED9" bold="true" box="[415,469,1200,1225]" pageId="19" pageNumber="209">Legs</emphasis>
|
||
: Protibial outer margin with 4 triangular teeth (1 apical, 2 medial, 1 basal); apical tooth curved; first medial tooth tooth larger compared to other teeth; basal tooth smaller than previous ones. Mesotibial outer carinae and apex covered with stout spinules like setae (3–4 on basal carina, 4–6 on medial carina, 7–10 on apex). Mesofemora surbectangular, moderately setose ventrally on disc. Metatibial basal carina usually with 3 spinules; medial carina with 8–12 stout spinules; apex bordered with 8–10 spinules. Metafemoral ventral surface glabrous on disc, scarcely covered with shallow punctures.
|
||
<emphasis id="759BEA9AFFB9FF83FEE877D3FE142FAB" bold="true" box="[285,401,1442,1467]" pageId="19" pageNumber="209">Abdomen</emphasis>
|
||
: Tergite 8 strongly convex in lateral view; surface densely covered with large, deep punctures, from coalescent to contiguous. Ventrites 2–5 rugopunctate on sides, scarcely punctate on disc; area near to posterior margin with an incomplete row of setae confined on sides, disc glabrous; ventrite VI emarginate at middle posteriorly, transversely rugose anteriorly, rugopunctate on sides, finely punctate on disc; posterior margin setose.
|
||
<emphasis id="759BEA9AFFB9FF83FEB37421FE102C79" bold="true" box="[326,405,1616,1641]" pageId="19" pageNumber="209">Venter</emphasis>
|
||
: Prosternal process wide, rounded apically. Mesoventrite completely rugopunctate, densely setose. Metaventral surface nearly completely setose, smooth on a transversal middle area at disc.
|
||
<emphasis id="759BEA9AFFB9FF83FDD474E4FD102CBE" bold="true" box="[545,661,1685,1710]" pageId="19" pageNumber="209">Aedeagus</emphasis>
|
||
: Parameres, in dorsal view, finely punctate, divergent and wide at basal half, convergent and narrow at apical half, scarcely bristled at inner margin of apex (
|
||
<figureCitation id="DFD42A0DFFBEFF84FE5572C0FD8F2ADA" box="[416,522,177,202]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="20" pageNumber="210">Figure 10</figureCitation>
|
||
(a)). Parameres, in lateral view, arched dorsally, ventrally bearing a middle carina, apex slightly rounded (
|
||
<figureCitation id="DFD42A0DFFBEFF84FD4072A5FC9A2AFD" box="[693,799,212,237]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="20" pageNumber="210">Figure 10</figureCitation>
|
||
(b)).
|
||
</paragraph>
|
||
<caption id="13906600FFBBFF81FF55771DFE6C2FB1" ID-DOI="http://doi.org/10.5281/zenodo.6758347" ID-Zenodo-Dep="6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="17" pageNumber="207" startId="17.[160,227,1388,1411]" targetBox="[168,1147,145,1355]" targetPageId="17">
|
||
<paragraph id="47503688FFBBFF81FF55771DFE6C2FB1" blockId="17.[160,1155,1388,1441]" pageId="17" pageNumber="207">
|
||
<emphasis id="759BEA9AFFBBFF81FF55771DFE972F93" bold="true" box="[160,274,1388,1411]" pageId="17" pageNumber="207">Figure 14.</emphasis>
|
||
Geographic distribution maps. (a)
|
||
<taxonomicName id="80EF4D0BFFBBFF81FD6A771DFCCB2F93" box="[671,846,1388,1411]" pageId="17" pageNumber="207">
|
||
<emphasis id="759BEA9AFFBBFF81FD6A771DFD712F93" box="[671,756,1388,1411]" italics="true" pageId="17" pageNumber="207">Tetragus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFBBFF81FC7E771DFBDC2F93" box="[907,1113,1388,1411]" pageId="17" pageNumber="207">
|
||
<emphasis id="759BEA9AFFBBFF81FC7E771DFC782F93" box="[907,1021,1388,1411]" italics="true" pageId="17" pageNumber="207">Gibboryctes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFBBFF81FF5577FBFE6C2FB1" authority="Prell." authorityName="Prell." box="[160,489,1418,1441]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="207" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFBBFF81FF5577FBFE342FB1" box="[160,433,1418,1441]" italics="true" pageId="17" pageNumber="207">Heterogomphus gracilicornis</emphasis>
|
||
Prell.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFB8FF82FF55771CFD3D2F94" ID-DOI="http://doi.org/10.5281/zenodo.6758349" ID-Zenodo-Dep="6758349" box="[160,696,1389,1412]" httpUri="https://zenodo.org/record/6758349/files/figure.png" pageId="18" pageNumber="208" startId="18.[160,225,1389,1412]" targetBox="[168,1146,235,1354]" targetPageId="18">
|
||
<paragraph id="47503688FFB8FF82FF55771CFD3D2F94" blockId="18.[160,696,1389,1412]" box="[160,696,1389,1412]" pageId="18" pageNumber="208">
|
||
<emphasis id="759BEA9AFFB8FF82FF55771CFE822F94" bold="true" box="[160,263,1389,1412]" pageId="18" pageNumber="208">Figure 15.</emphasis>
|
||
(a) Rupestrian grassland; (b) rock outcrops.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFB9FF83FF55767AFE0B2E4C" ID-DOI="http://doi.org/10.5281/zenodo.6758351" ID-Zenodo-Dep="6758351" httpUri="https://zenodo.org/record/6758351/files/figure.png" pageId="19" pageNumber="209" startId="19.[160,225,1035,1058]" targetBox="[166,1148,142,1003]" targetPageId="19">
|
||
<paragraph id="47503688FFB9FF83FF55767AFE0B2E4C" blockId="19.[160,1156,1035,1116]" pageId="19" pageNumber="209">
|
||
<emphasis id="759BEA9AFFB9FF83FF55767AFE832E32" bold="true" box="[160,262,1035,1058]" pageId="19" pageNumber="209">Figure 16.</emphasis>
|
||
(a) Adult of
|
||
<taxonomicName id="80EF4D0BFFB9FF83FE88767AFD142E32" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[381,657,1035,1058]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="209" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFB9FF83FE88767AFDC52E32" box="[381,576,1035,1058]" italics="true" pageId="19" pageNumber="209">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
at central portion of termite nest (arrow); (b) detail of the adult
|
||
<emphasis id="759BEA9AFFB9FF83FEEB7659FEDC2E2F" box="[286,345,1064,1087]" italics="true" pageId="19" pageNumber="209">in situ</emphasis>
|
||
(arrow); (c) pupal chamber with its structures in the shape of pellets (arrow); (d) recently emerged adult.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="47503688FFBEFF84FF557359FE0E2B52" blockId="20.[160,1156,296,460]" box="[160,395,296,322]" pageId="20" pageNumber="210">
|
||
<emphasis id="759BEA9AFFBEFF84FF557359FE0E2B52" bold="true" box="[160,395,296,322]" pageId="20" pageNumber="210">
|
||
Female
|
||
<emphasis id="759BEA9AFFBEFF84FEF57359FE0E2B52" bold="true" box="[256,395,296,322]" italics="true" pageId="20" pageNumber="210">
|
||
(
|
||
<figureCitation id="DFD42A0DFFBEFF84FEFC7359FEE02B52" box="[265,357,296,322]" captionStart="Figure 1" captionStartId="4.[160,227,1366,1389]" captionTargetBox="[162,1148,150,1331]" captionTargetId="figure-77@4.[162,1154,142,1335]" captionTargetPageId="4" captionText="Figure 1. Gibboryctes szelenyii Endrödi, 1974. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale." figureDoi="http://doi.org/10.5281/zenodo.6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="20" pageNumber="210">Figure 1</figureCitation>
|
||
(f))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFBEFF84FF55733AFCC02BDC" blockId="20.[160,1156,296,460]" pageId="20" pageNumber="210">
|
||
<emphasis id="759BEA9AFFBEFF84FF55733AFECD2B75" bold="true" box="[160,328,331,357]" italics="true" pageId="20" pageNumber="210">Redescription.</emphasis>
|
||
<emphasis id="759BEA9AFFBEFF84FEAD733DFE282B75" bold="true" box="[344,429,332,357]" pageId="20" pageNumber="210">Length</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFBEFF84FE4A733DFDE02B75" box="[447,613,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" metricValueMax="3.55" metricValueMin="2.81" pageId="20" pageNumber="210" unit="mm" value="31.8" valueMax="35.5" valueMin="28.1">28.1–35.5 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFBEFF84FD81733DFD382B75" bold="true" box="[628,701,332,357]" pageId="20" pageNumber="210">Width</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFBEFF84FD3B733DFCF02B75" box="[718,885,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.5699999999999998" metricValueMin="1.09" pageId="20" pageNumber="210" unit="mm" value="13.3" valueMax="15.7" valueMin="10.9">10.9–15.7 mm</quantity>
|
||
. Quite similar to male, except in the following aspects:
|
||
<emphasis id="759BEA9AFFBEFF84FDEA731FFDD82B97" bold="true" box="[543,605,366,391]" pageId="20" pageNumber="210">Head</emphasis>
|
||
: Clypeus slightly rounded anteriorly (
|
||
<figureCitation id="DFD42A0DFFBEFF84FBE3731FFB062B97" box="[1046,1155,366,391]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="20" pageNumber="210">Figure 11</figureCitation>
|
||
(a)).
|
||
<emphasis id="759BEA9AFFBEFF84FF2473E0FEC32BBA" bold="true" box="[209,326,401,426]" pageId="20" pageNumber="210">Abdomen</emphasis>
|
||
: Tergite VIII weakly convex in lateral view; ventrite VI parabolic in shape, not emarginate at middle of posterior margin (
|
||
<figureCitation id="DFD42A0DFFBEFF84FD4473C2FC9E2BDC" box="[689,795,435,460]" captionStart="Figure 13" captionStartId="16.[160,225,1058,1081]" captionTargetBox="[166,1146,145,1023]" captionTargetId="figure-155@16.[166,1149,142,1026]" captionTargetPageId="16" captionText="Figure 13. Detail of ventrites I–VI of females. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov; (c) Gibboryctes endroedii sp. nov.; (d) Tetragus waldenfelsi (Endrödi). Red arrows indicate the proportions of ventrites V and VI." figureDoi="http://doi.org/10.5281/zenodo.6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="20" pageNumber="210">Figure 13</figureCitation>
|
||
(a)).
|
||
</paragraph>
|
||
<paragraph id="47503688FFBEFF84FF557079FDE82832" blockId="20.[160,1102,520,580]" box="[160,621,520,546]" pageId="20" pageNumber="210">
|
||
<emphasis id="759BEA9AFFBEFF84FF557079FDE82832" bold="true" box="[160,621,520,546]" pageId="20" pageNumber="210">
|
||
Geographic distribution
|
||
<emphasis id="759BEA9AFFBEFF84FE397079FDE82832" bold="true" box="[460,621,520,546]" italics="true" pageId="20" pageNumber="210">
|
||
(
|
||
<figureCitation id="DFD42A0DFFBEFF84FE207079FDC42832" box="[469,577,520,546]" captionStart="Figure 14" captionStartId="17.[160,227,1388,1411]" captionTargetBox="[168,1147,145,1355]" captionTargetId="figure-89@17.[166,1149,142,1357]" captionTargetPageId="17" captionText="Figure 14. Geographic distribution maps. (a) Tetragus species; (b) Gibboryctes species; (c) Heterogomphus gracilicornis Prell." figureDoi="http://doi.org/10.5281/zenodo.6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="20" pageNumber="210">Figure 14</figureCitation>
|
||
(b))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFBEFF84FF55705AFBCB2854" blockId="20.[160,1102,520,580]" box="[160,1102,555,580]" pageId="20" pageNumber="210">
|
||
<collectingCountry id="3FF87618FFBEFF84FF55705AFE8F2854" box="[160,266,555,580]" name="Paraguay" pageId="20" pageNumber="210">Paraguay</collectingCountry>
|
||
, with no further data;
|
||
<collectingCountry id="3FF87618FFBEFF84FDF0705AFDC42854" box="[517,577,555,580]" name="Brazil" pageId="20" pageNumber="210">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="852BF86AFFBEFF84FDBA705AFD252854" box="[591,672,555,580]" country="Brazil" name="Parana" pageId="20" pageNumber="210">Paraná</collectingRegion>
|
||
, Distrito Federal), new country record.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFBEFF84FF5570F1FBB529C2" pageId="20" pageNumber="210" type="discussion">
|
||
<paragraph id="47503688FFBEFF84FF5570F1FE83288A" blockId="20.[160,1156,640,978]" box="[160,262,640,666]" pageId="20" pageNumber="210">
|
||
<emphasis id="759BEA9AFFBEFF84FF5570F1FE83288A" bold="true" box="[160,262,640,666]" italics="true" pageId="20" pageNumber="210">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFBEFF84FF5570D2FBB529C2" blockId="20.[160,1156,640,978]" pageId="20" pageNumber="210">
|
||
For a long time, no other record of
|
||
<taxonomicName id="80EF4D0BFFBEFF84FDDB70D2FD2028AC" authorityName="Endrodi" authorityYear="1974" box="[558,677,675,700]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFBEFF84FDDB70D2FD2028AC" box="[558,677,675,700]" italics="true" pageId="20" pageNumber="210">G. szelenyii</emphasis>
|
||
</taxonomicName>
|
||
had been made since the species description by
|
||
<bibRefCitation id="237E4B79FFBEFF84FF0870B7FE2228CF" author="Endrodi S" box="[253,423,710,735]" pageId="20" pageNumber="210" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">Endrödi (1974)</bibRefCitation>
|
||
, only the
|
||
<typeStatus id="9854882AFFBEFF84FDD570B7FDD728CF" box="[544,594,710,735]" pageId="20" pageNumber="210">type</typeStatus>
|
||
specimen being known – and without accurate locality data – from
|
||
<collectingCountry id="3FF87618FFBEFF84FE757098FE6F2912" box="[384,490,745,770]" name="Paraguay" pageId="20" pageNumber="210">Paraguay</collectingCountry>
|
||
. The species is recorded for the first time from
|
||
<collectingCountry id="3FF87618FFBEFF84FC0E7098FBB22912" box="[1019,1079,745,770]" name="Brazil" pageId="20" pageNumber="210">Brazil</collectingCountry>
|
||
in this study, where
|
||
<specimenCount id="51E9FD01FFBEFF84FECB717DFE5F2935" box="[318,474,780,805]" pageId="20" pageNumber="210" type="generic">30 specimens</specimenCount>
|
||
were collected in termite nests within the Campos Gerais region, located at the second
|
||
<collectingRegion id="852BF86AFFBEFF84FE1A715FFDBE2957" box="[495,571,814,839]" country="Brazil" name="Parana" pageId="20" pageNumber="210">Parana</collectingRegion>
|
||
plateau (
|
||
<figureCitation id="DFD42A0DFFBEFF84FD51715FFC8A2957" box="[676,783,814,839]" captionStart="Figure 15" captionStartId="18.[160,225,1389,1412]" captionTargetBox="[168,1146,235,1354]" captionTargetId="figure-89@18.[166,1149,142,1357]" captionTargetPageId="18" captionText="Figure 15. (a) Rupestrian grassland; (b) rock outcrops." figureDoi="http://doi.org/10.5281/zenodo.6758349" httpUri="https://zenodo.org/record/6758349/files/figure.png" pageId="20" pageNumber="210">Figure 15</figureCitation>
|
||
(a,b)). The termites sampled were probably occupying an old termite nest of another species, and were themselves identified as belonging to multiple genera and species (R. Constantino pers. comm., 2011) as follows:
|
||
<taxonomicName id="80EF4D0BFFBEFF84FF0B71E7FD4E29BF" authority="(Silvestri, 1901)" baseAuthorityName="Silvestri" baseAuthorityYear="1901" box="[254,715,918,943]" class="Insecta" family="Termitidae" genus="Diversitermes" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="diversimiles">
|
||
<emphasis id="759BEA9AFFBEFF84FF0B71E7FD9029BF" box="[254,533,918,943]" italics="true" pageId="20" pageNumber="210">Diversitermes diversimiles</emphasis>
|
||
(Silvestri, 1901)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80EF4D0BFFBEFF84FD2271E7FB0529BF" authority="(Hagen, 1858)" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[727,1152,918,943]" class="Insecta" family="Termitidae" genus="Neocapritermes" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="759BEA9AFFBEFF84FD2271E7FC5229BF" box="[727,983,918,943]" italics="true" pageId="20" pageNumber="210">Neocapritermes opacus</emphasis>
|
||
(Hagen, 1858)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80EF4D0BFFBEFF84FF5571C8FDF829C2" authority="(Holmgren, 1910)" baseAuthorityName="Holmgren" baseAuthorityYear="1910" box="[160,637,953,978]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="coxipoensis">
|
||
<emphasis id="759BEA9AFFBEFF84FF5571C8FE3529C2" box="[160,432,953,978]" italics="true" pageId="20" pageNumber="210">Nasutitermes coxipoensis</emphasis>
|
||
(Holmgren, 1910)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80EF4D0BFFBEFF84FD7D71C8FCC329C2" box="[648,838,953,978]" class="Insecta" family="Termitidae" genus="Anoplotermes" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="759BEA9AFFBEFF84FD7D71C8FC9B29C2" box="[648,798,953,978]" italics="true" pageId="20" pageNumber="210">Anoplotermes</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80EF4D0BFFBEFF84FC8B71C8FBB529C2" box="[894,1072,953,978]" class="Insecta" family="Termitidae" genus="Grigiotermes" kingdom="Animalia" order="Blattodea" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="759BEA9AFFBEFF84FC8B71C8FB8D29C2" box="[894,1032,953,978]" italics="true" pageId="20" pageNumber="210">Grigiotermes</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |