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<document id="4793F0764E63F456ECE12E45ADE12B20" ID-CLB-Dataset="60704" ID-DOI="10.5852/ejt.2023.885.2199" ID-GBIF-Dataset="d205aec5-8b36-4f96-b447-afe09885bdf0" ID-ISSN="2118-9773" ID-Zenodo-Dep="8222417" ID-ZooBank="8432EEFF-391F-4778-81F6-14F3F6ECAA5F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691437419139" checkinUser="felipe" docAuthor="Korn, Dieter &amp; Klug, Christian" docDate="2023" docId="AE7E87800766FF9EFDEC135CFB76F92C" docLanguage="en" docName="ejt-885_156-194.pdf" docOrigin="European Journal of Taxonomy 885" docSource="http://dx.doi.org/10.5852/ejt.2023.885.2199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Temnocheilus M'Coy 1844" docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="5247FFF80761FF99FFA71124FF8FFFA1" masterDocTitle="Early Carboniferous coiled nautiloids from the Anti-Atlas (Morocco)" masterLastPageNumber="194" masterPageNumber="156" pageNumber="163" updateTime="1699551382698" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3DD4E649EB362B8BBEC188B89C05FBAB">Early Carboniferous coiled nautiloids from the Anti-Atlas (Morocco)</mods:title>
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<mods:namePart id="D457AC9A7382EFF238A11B92D4DE4E5A">Korn, Dieter</mods:namePart>
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<mods:affiliation id="7DC5E726CCFB1EE735E869C2A2F1AA5E">Museum für Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
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<mods:affiliation id="B3242BA56E33FDB41A3D2F28E5ACB0D7">Paläontologisches Institut und Museum, Universität Zürich, Karl-Schmid-Strasse 4, 8006 Zürich, Switzerland.</mods:affiliation>
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<subSubSection id="6ECD651D0766FF9EFDEC135CFC66FD33" box="[587,1001,632,659]" pageId="7" pageNumber="163" type="nomenclature">
<paragraph id="266836960766FF9EFDEC135CFC66FD33" blockId="7.[587,1001,632,659]" box="[587,1001,632,659]" pageId="7" pageNumber="163">
<heading id="7D2081FA0766FF9EFDEC135CFC66FD33" box="[587,1001,632,659]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="163" reason="2">
Genus
<taxonomicName id="E1D74D150766FF9EFD3A135CFC66FD33" authority="M'Coy, 1844" authorityName="M'Coy" authorityYear="1844" box="[669,1001,632,659]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA840766FF9EFD3A135CFCCAFD32" bold="true" box="[669,837,632,659]" italics="true" pageId="7" pageNumber="163">Temnocheilus</emphasis>
<bibRefCitation id="42464B670766FF9EFCEA135CFC66FD33" author="M'Coy F." box="[845,1001,632,658]" pageId="7" pageNumber="163" refId="ref16278" refString="M'Coy F. 1844. A Synopsis of the Characters of the Carboniferous Fossils of Ireland. University Press, Dublin. https: // doi. org / 10.5962 / bhl. title. 11559" type="book" year="1844">MCoy, 1844</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="6ECD651D0766FF9EFF1A139FFAA1FCA4" pageId="7" pageNumber="163" type="type_taxon">
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<heading id="7D2081FA0766FF9EFF1A139FFED7FD75" bold="true" box="[189,344,698,725]" fontSize="11" level="3" pageId="7" pageNumber="163" reason="3">
<emphasis id="14A3EA840766FF9EFF1A139FFED7FD75" bold="true" box="[189,344,698,725]" pageId="7" pageNumber="163">
<typeStatus id="F96C88340766FF9EFF1A139FFF75FD74" box="[189,250,699,725]" pageId="7" pageNumber="163">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="266836960766FF9EFF1A13CEFAA1FCA4" blockId="7.[189,1326,746,773]" box="[189,1326,746,773]" pageId="7" pageNumber="163">
<taxonomicName id="E1D74D150766FF9EFF1A13CEFD71FCA5" authority="M'Coy, 1844" authorityName="M'Coy" authorityYear="1844" box="[189,766,746,772]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="coronatus" subGenus="Temnocheilus">
<emphasis id="14A3EA840766FF9EFF1A13CEFEAFFCA5" box="[189,288,746,772]" italics="true" pageId="7" pageNumber="163">Nautilus</emphasis>
(
<emphasis id="14A3EA840766FF9EFE9513CEFE5DFCA5" box="[306,466,746,772]" italics="true" pageId="7" pageNumber="163">Temnocheilus</emphasis>
)
<emphasis id="14A3EA840766FF9EFE4413CFFDD8FCA5" box="[483,599,747,772]" italics="true" pageId="7" pageNumber="163">coronatus</emphasis>
<bibRefCitation id="42464B670766FF9EFDF913CEFD71FCA5" author="M'Coy F." box="[606,766,746,772]" pageId="7" pageNumber="163" refId="ref16278" refString="M'Coy F. 1844. A Synopsis of the Characters of the Carboniferous Fossils of Ireland. University Press, Dublin. https: // doi. org / 10.5962 / bhl. title. 11559" type="book" year="1844">MCoy, 1844</bibRefCitation>
</taxonomicName>
; subsequent designation by
<bibRefCitation id="42464B670766FF9EFBE013CFFB5CFCA5" author="Hyatt A." box="[1095,1235,746,773]" pageId="7" pageNumber="163" pagination="253 - 338" refId="ref15510" refString="Hyatt A. 1883 - 1884. Genera of fossil cephalopods. Proceedings of the Boston Society of Natural History 22: 253 - 338." type="journal article" year="1883">Hyatt (1883</bibRefCitation>
1884).
</paragraph>
</subSubSection>
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<heading id="7D2081FA0766FF9EFF1A1208FEB9FCE7" bold="true" box="[189,310,812,838]" fontSize="11" level="3" pageId="7" pageNumber="163" reason="3">
<emphasis id="14A3EA840766FF9EFF1A1208FEB9FCE7" bold="true" box="[189,310,812,838]" pageId="7" pageNumber="163">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="266836960766FF9EFF1A1278FC07FBF1" blockId="7.[188,1399,860,1104]" pageId="7" pageNumber="163">
Genus of the family
<taxonomicName id="E1D74D150766FF9EFE091278FDFAFCD7" authorityName="Hyatt" authorityYear="1884" box="[430,629,860,886]" class="Cephalopoda" family="Trigonoceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="family">Trigonoceratidae</taxonomicName>
with discoidal to pachyconic, usually subevolute conch. The first whorl is about
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in diameter with an umbilical foramen about
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wide; the conch is rapidly increasing in height with a high coiling rate (WER usually higher than 2.00). Whorls weakly embracing, their profile triangular with more or less angular umbilical margin. Sculpture with longitudinal tubercles along the ventral margin.
<collectingRegion id="E413F8740766FF9EFE5B12C9FDB2FBA6" box="[508,573,1005,1031]" country="Spain" name="Ceuta" pageId="7" pageNumber="163">Septa</collectingRegion>
usually without inflexions, rather deeply concave. Suture line with shallow ventral and deeper lateral, rounded internal lobe. The siphuncle has a position between the centre of the aperture and the venter (after
<bibRefCitation id="42464B670766FF9EFD141512FCF7FBF1" author="Shimansky V. N." box="[691,888,1078,1104]" pageId="7" pageNumber="163" pagination="1 - 244" refId="ref16954" refString="Shimansky V. N. 1967. Kamennougol'nye Nautilida. Trudy paleontologicheskogo Instituta Akademiya Nauk SSSR 115: 1 - 244." type="journal article" year="1967">Shimansky 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D0766FF9EFF1A155CFE27F9BB" pageId="7" pageNumber="163" type="distribution">
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<emphasis id="14A3EA840766FF9EFF1A155CFD1BFB33" bold="true" box="[189,660,1144,1170]" pageId="7" pageNumber="163">Included Early Carboniferous species</emphasis>
</heading>
</paragraph>
<paragraph id="266836960766FF9EFF1A158CFC7CFB47" blockId="7.[189,1397,1192,1255]" pageId="7" pageNumber="163">
Most of the species of
<taxonomicName id="E1D74D150766FF9EFE18158CFDD0FB63" authorityName="M'Coy" authorityYear="1844" box="[447,607,1192,1218]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="14A3EA840766FF9EFE18158CFDD0FB63" box="[447,607,1192,1218]" italics="true" pageId="7" pageNumber="163">Temnocheilus</emphasis>
</taxonomicName>
are known from Late Carboniferous and Permian strata (
<bibRefCitation id="42464B670766FF9EFB56158CFEA9FB47" author="Shimansky V. N." pageId="7" pageNumber="163" pagination="1 - 244" refId="ref16954" refString="Shimansky V. N. 1967. Kamennougol'nye Nautilida. Trudy paleontologicheskogo Instituta Akademiya Nauk SSSR 115: 1 - 244." type="journal article" year="1967">Shimansky 1967: 98</bibRefCitation>
). Early Carboniferous species are known from three regions:
</paragraph>
<paragraph id="266836960766FF9EFF1A142AFBA2FAD0" blockId="7.[189,1399,1294,1393]" pageId="7" pageNumber="163">
British Isles (
<bibRefCitation id="42464B670766FF9EFEC6142AFD9BFA88" author="Sowerby J." box="[353,532,1294,1321]" pageId="7" pageNumber="163" refId="ref16976" refString="Sowerby J. 1821. The Mineral Conchology of Great Britain; or Coloured Figures and Descriptions of those Remains of Testaceous Animals or Shells, which have been Preserved at Various Times and Depths in the Earth. Vol. 3. Arding, London." type="book" year="1821">Sowerby 1821</bibRefCitation>
;
<bibRefCitation id="42464B670766FF9EFD83142BFD4DFA88" author="M'Coy F." box="[548,706,1295,1321]" pageId="7" pageNumber="163" refId="ref16278" refString="M'Coy F. 1844. A Synopsis of the Characters of the Carboniferous Fossils of Ireland. University Press, Dublin. https: // doi. org / 10.5962 / bhl. title. 11559" type="book" year="1844">MCoy 1844</bibRefCitation>
;
<bibRefCitation id="42464B670766FF9EFD76142AFCDAFA88" author="Hind W." box="[721,853,1294,1321]" pageId="7" pageNumber="163" pagination="18 - 19" refId="ref15395" refString="Hind W. 1914. Temnocheilus derbiensis, sp. nov. Proceedings of the Yorkshire Geological Society 19: 18 - 19. https: // doi. org / 10.1144 / pygs. 19.1.18" type="journal article" year="1914">Hind 1914</bibRefCitation>
;
<bibRefCitation id="42464B670766FF9EFCC2142AFC6DFA88" author="Platt M. I." box="[869,994,1294,1321]" pageId="7" pageNumber="163" pagination="454 - 459" refId="ref16735" refString="Platt M. I. 1938. A new species of Temnocheilus McCoy from Holy Island, Northumberland. Geological Magazine 75: 454 - 459. https: // doi. org / 10.1017 / S 0016756800091895" type="journal article" year="1938">Platt 1938</bibRefCitation>
):
<taxonomicName id="E1D74D150766FF9EFC59142AFF75FAEC" authority="Sowerby, 1821" authorityName="Sowerby" authorityYear="1821" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="tuberculatus">
<emphasis id="14A3EA840766FF9EFC59142AFB71FA89" box="[1022,1278,1294,1320]" italics="true" pageId="7" pageNumber="163">Nautilus tuberculatus</emphasis>
<bibRefCitation id="42464B670766FF9EFAAE142AFF75FAEC" author="Sowerby J." pageId="7" pageNumber="163" refId="ref16976" refString="Sowerby J. 1821. The Mineral Conchology of Great Britain; or Coloured Figures and Descriptions of those Remains of Testaceous Animals or Shells, which have been Preserved at Various Times and Depths in the Earth. Vol. 3. Arding, London." type="book" year="1821">Sowerby, 1821</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC076060766FF9EFEAC1417FEFAFAEC" box="[267,373,1331,1357]" name="United Kingdom" pageId="7" pageNumber="163">Scotland</collectingCountry>
;
<taxonomicName id="E1D74D150766FF9EFE211417FC56FAEC" authority="M'Coy, 1844" authorityName="M'Coy" authorityYear="1844" box="[390,985,1331,1357]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="coronatus" subGenus="Temnocheilus">
<emphasis id="14A3EA840766FF9EFE211417FE66FAEC" box="[390,489,1331,1357]" italics="true" pageId="7" pageNumber="163">Nautilus</emphasis>
(
<emphasis id="14A3EA840766FF9EFE581417FD10FAEC" box="[511,671,1331,1357]" italics="true" pageId="7" pageNumber="163">Temnocheilus</emphasis>
)
<emphasis id="14A3EA840766FF9EFD121417FCA6FAED" box="[693,809,1331,1356]" italics="true" pageId="7" pageNumber="163">coronatus</emphasis>
<bibRefCitation id="42464B670766FF9EFC911417FC56FAEC" author="M'Coy F." box="[822,985,1331,1357]" pageId="7" pageNumber="163" refId="ref16278" refString="M'Coy F. 1844. A Synopsis of the Characters of the Carboniferous Fossils of Ireland. University Press, Dublin. https: // doi. org / 10.5962 / bhl. title. 11559" type="book" year="1844">MCoy, 1844</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC076060766FF9EFC4D1417FBCEFAEC" box="[1002,1089,1331,1357]" name="Ireland" pageId="7" pageNumber="163">Ireland</collectingCountry>
;
<taxonomicName id="E1D74D150766FF9EFBF61417FECCFAD0" authority="Hind, 1914" authorityName="Hind" authorityYear="1914" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="derbiensis">
<emphasis id="14A3EA840766FF9EFBF61417FAF8FAEC" box="[1105,1399,1331,1357]" italics="true" pageId="7" pageNumber="163">Temnocheilus derbiensis</emphasis>
<bibRefCitation id="42464B670766FF9EFF1A1473FECCFAD0" author="Hind W." box="[189,323,1367,1393]" pageId="7" pageNumber="163" pagination="18 - 19" refId="ref15395" refString="Hind W. 1914. Temnocheilus derbiensis, sp. nov. Proceedings of the Yorkshire Geological Society 19: 18 - 19. https: // doi. org / 10.1144 / pygs. 19.1.18" type="journal article" year="1914">Hind, 1914</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC076060766FF9EFEF71473FE39FAD0" box="[336,438,1367,1393]" name="United Kingdom" pageId="7" pageNumber="163">England</collectingCountry>
;
<taxonomicName id="E1D74D150766FF9EFE661473FC3DFAD0" authority="Platt, 1938" authorityName="Platt" authorityYear="1938" box="[449,946,1367,1393]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="ventro-concavus">
<emphasis id="14A3EA840766FF9EFE661473FCA6FAD0" box="[449,809,1367,1393]" italics="true" pageId="7" pageNumber="163">Temnocheilus ventro-concavus</emphasis>
<bibRefCitation id="42464B670766FF9EFC961473FC3DFAD0" author="Platt M. I." box="[817,946,1367,1393]" pageId="7" pageNumber="163" pagination="454 - 459" refId="ref16735" refString="Platt M. I. 1938. A new species of Temnocheilus McCoy from Holy Island, Northumberland. Geological Magazine 75: 454 - 459. https: // doi. org / 10.1017 / S 0016756800091895" type="journal article" year="1938">Platt, 1938</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="5EC076060766FF9EFC191473FBA6FAD0" box="[958,1065,1367,1393]" name="United Kingdom" pageId="7" pageNumber="163">Scotland</collectingCountry>
.
</paragraph>
<paragraph id="266836960766FF9EFF1A14BDFA87FA12" blockId="7.[189,1288,1433,1460]" box="[189,1288,1433,1460]" pageId="7" pageNumber="163">
South Urals (
<bibRefCitation id="42464B670766FF9EFEF014BDFD90FA12" author="Shimansky V. N." box="[343,543,1433,1459]" pageId="7" pageNumber="163" pagination="1 - 244" refId="ref16954" refString="Shimansky V. N. 1967. Kamennougol'nye Nautilida. Trudy paleontologicheskogo Instituta Akademiya Nauk SSSR 115: 1 - 244." type="journal article" year="1967">Shimansky 1967</bibRefCitation>
):
<taxonomicName id="E1D74D150766FF9EFD9014BDFBE6FA12" authority="Shimansky, 1967" authorityName="Shimansky" authorityYear="1967" box="[567,1129,1433,1459]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="coronatiformae">
<emphasis id="14A3EA840766FF9EFD9014BDFC1BFA12" box="[567,916,1433,1459]" italics="true" pageId="7" pageNumber="163">Temnocheilus coronatiformae</emphasis>
<bibRefCitation id="42464B670766FF9EFC3C14BDFBE6FA12" author="Shimansky V. N." box="[923,1129,1433,1459]" pageId="7" pageNumber="163" pagination="1 - 244" refId="ref16954" refString="Shimansky V. N. 1967. Kamennougol'nye Nautilida. Trudy paleontologicheskogo Instituta Akademiya Nauk SSSR 115: 1 - 244." type="journal article" year="1967">Shimansky, 1967</bibRefCitation>
</taxonomicName>
; South Urals.
</paragraph>
<paragraph id="266836960766FF9EFF1A14F8FE27F9BB" blockId="7.[189,1399,1500,1563]" pageId="7" pageNumber="163">
North Africa (this paper):
<taxonomicName id="E1D74D150766FF9EFE4314F8FCCDFA57" authority="Korn &amp; Klug, 2023" authorityName="Korn &amp; Klug" authorityYear="2023" box="[484,834,1500,1526]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="imazighenorum" status="sp. nov.">
<emphasis id="14A3EA840766FF9EFE4314F8FCCDFA57" box="[484,834,1500,1526]" italics="true" pageId="7" pageNumber="163">Temnocheilus imazighenorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0F9057FF0766FF9EFCE014F8FC2DFA57" box="[839,930,1500,1526]" pageId="7" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="E413F8740766FF9EFC0D14F8FBA5FA57" box="[938,1066,1500,1526]" pageId="7" pageNumber="163">Anti-Atlas</collectingRegion>
;
<taxonomicName id="E1D74D150766FF9EFB9714F8FAF8FA57" authority="Korn &amp; Klug, 2023" authorityName="Korn &amp; Klug" authorityYear="2023" box="[1072,1399,1500,1526]" class="Cephalopoda" family="Koninckioceratidae" genus="Temnocheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="7" pageNumber="163" phylum="Mollusca" rank="species" species="aubrechtovae" status="sp. nov.">
<emphasis id="14A3EA840766FF9EFB9714F8FAF8FA57" box="[1072,1399,1500,1526]" italics="true" pageId="7" pageNumber="163">Temnocheilus aubrechtovae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0F9057FF0766FF9EFF1A1725FE95F9BA" box="[189,282,1537,1563]" pageId="7" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="E413F8740766FF9EFE831724FE2BF9BB" box="[292,420,1536,1562]" pageId="7" pageNumber="163">Anti-Atlas</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="6ECD651D0766FF9EFF1A1767FB76F92C" box="[189,1273,1603,1677]" pageId="7" pageNumber="163" type="discussion">
<paragraph id="266836960766FF9EFF1A1767FEA0F9FC" blockId="7.[189,303,1603,1629]" box="[189,303,1603,1629]" pageId="7" pageNumber="163">
<heading id="7D2081FA0766FF9EFF1A1767FEA0F9FC" bold="true" box="[189,303,1603,1629]" fontSize="11" level="3" pageId="7" pageNumber="163" reason="3">
<emphasis id="14A3EA840766FF9EFF1A1767FEA0F9FC" bold="true" box="[189,303,1603,1629]" pageId="7" pageNumber="163">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="266836960766FF9EFF1A1757FB76F92C" blockId="7.[189,1273,1650,1677]" box="[189,1273,1650,1677]" pageId="7" pageNumber="163">
For a discussion of the composition and limits of the genus, see
<bibRefCitation id="42464B670766FF9EFC0A1757FB7DF92D" author="Korn D. &amp; Bockwinkel J." box="[941,1266,1650,1677]" pageId="7" pageNumber="163" pagination="67 - 108" refId="ref15872" refString="Korn D. &amp; Bockwinkel J. 2022. Early Carboniferous nautiloids from the Central Sahara, southern Algeria. European Journal of Taxonomy 831: 67 - 108. https: // doi. org / 10.5852 / ejt. 2022.831.1871" type="journal article" year="2022">Korn &amp; Bockwinkel (2022)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>