636 lines
88 KiB
XML
636 lines
88 KiB
XML
<document id="D0468CE5AFE564231B2FAE1C4B194CE6" ID-DOI="10.5281/zenodo.183467" ID-GBIF-Dataset="a63daeed-ef54-4cf9-adec-42e9dd1ab497" ID-ISSN="1175-5326" ID-Zenodo-Dep="183467" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459907824163" checkinUser="plazi" docAuthor="D, Eduardo Flórez, Botero-Trujillo, Ricardo & Acosta, Luis E." docDate="2008" docId="1A6087DBFFF7DA60FF30FC79FBBEF954" docLanguage="en" docName="zt01853p044.pdf" docOrigin="Zootaxa 1853" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Vachoniochactas humboldti D, Botero-Trujillo & Acosta, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="E659FFA3FFF5DA66FFA7FFF6FFF6FFA6" masterDocTitle="Description of Vachoniochactas humboldti sp. nov. from Colombia, with complementary notes on the genus (Scorpiones, Chactidae)" masterLastPageNumber="44" masterPageNumber="31" pageNumber="33" updateTime="1698229169680" updateUser="plazi">
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<mods:title id="F947CC26ADF268CA4E34F1B86128C04F">Description of Vachoniochactas humboldti sp. nov. from Colombia, with complementary notes on the genus (Scorpiones, Chactidae)</mods:title>
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<mods:name id="9F3A86C43F44E7CE338F0A6E3BE5B595" type="personal">
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<mods:namePart id="E1C53A48C1A28B50A0C76275A7C57542">D, Eduardo Flórez</mods:namePart>
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<mods:roleTerm id="77BA069EE4F5475F623706239DFACB0C">Author</mods:roleTerm>
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<mods:namePart id="F3ACED2B10BD6F60A347E6448FFEDABE">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:roleTerm id="E354D57D2CDFCD8A21A5FDE2DD5C72DB">Author</mods:roleTerm>
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<mods:namePart id="0B46A59B10688E746299E29A049E4D58">Acosta, Luis E.</mods:namePart>
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<mods:typeOfResource id="B681B1CCBCAEB674689B26AA77944A80">text</mods:typeOfResource>
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<mods:title id="4A3BFD0DE5C463E56EE9E7B5F7F3F39E">Zootaxa</mods:title>
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<mods:part id="A102290A5369F503C2B985F98C4029D9">
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<mods:date id="AC3B2794558C48E13ED0613C1BF587C3">2008</mods:date>
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<mods:number id="42F1B6F4B1D7FB9722A0850A445754DC">1853</mods:number>
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<mods:classification id="D8DC23111A777991EE603EB999BB78DB">journal article</mods:classification>
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<mods:identifier id="11BBC03D169149D3C9D2BED315143707" type="DOI">10.5281/zenodo.183467</mods:identifier>
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<treatment id="1A6087DBFFF7DA60FF30FC79FBBEF954" ID-DOI="http://doi.org/10.5281/zenodo.5674674" ID-GBIF-Taxon="119373597" ID-Zenodo-Dep="5674674" LSID="urn:lsid:plazi:treatment:1A6087DBFFF7DA60FF30FC79FBBEF954" httpUri="http://treatment.plazi.org/id/1A6087DBFFF7DA60FF30FC79FBBEF954" lastPageId="6" lastPageNumber="37" pageId="2" pageNumber="33">
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<subSubSection id="DAD36546FFF7DA64FF30FC79FE60FC6A" pageId="2" pageNumber="33" type="nomenclature">
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<paragraph id="927636CDFFF7DA64FF30FC79FDBDFC0F" blockId="2.[151,587,911,972]" box="[151,587,911,937]" pageId="2" pageNumber="33">
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<heading id="C93E81A1FFF7DA64FF30FC79FDBDFC0F" bold="true" box="[151,587,911,937]" fontSize="11" level="1" pageId="2" pageNumber="33" reason="1">
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<emphasis id="A0BDEADFFFF7DA64FF30FC79FDBDFC0F" bold="true" box="[151,587,911,937]" pageId="2" pageNumber="33">
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<taxonomicName id="55C94D4EFFF7DA64FF30FC79FE11FC0F" ID-CoL="7F99M" box="[151,487,911,937]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
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<emphasis id="A0BDEADFFFF7DA64FF30FC79FE11FC0F" bold="true" box="[151,487,911,937]" italics="true" pageId="2" pageNumber="33">Vachoniochactas humboldti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BB8E57A4FFF7DA64FE4AFC79FDBDFC0F" box="[493,587,911,937]" pageId="2" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="927636CDFFF7DA64FF30FC44FE60FC6A" blockId="2.[151,587,911,972]" box="[151,406,946,972]" pageId="2" pageNumber="33">
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<figureCitation id="0AF22A48FFF7DA64FF30FC44FEC5FC6A" box="[151,307,946,972]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 5" captionStart-2="FIGURES 6 – 10" captionStart-3="FIGURES 11 – 18" captionStart-4="FIGURES 19 – 21" captionStartId-0="3.[151,255,1235,1259]" captionStartId-1="7.[151,269,1808,1832]" captionStartId-2="8.[151,269,1759,1783]" captionStartId-3="9.[151,269,1877,1901]" captionStartId-4="10.[151,269,1306,1330]" captionTargetBox-0="[151,1435,185,1208]" captionTargetBox-1="[207,1379,195,1783]" captionTargetBox-2="[152,1399,222,1709]" captionTargetBox-3="[177,1431,214,1817]" captionTargetBox-4="[177,1421,234,1257]" captionTargetId-0="figure@3.[151,1436,185,1209]" captionTargetId-1="figure@7.[207,1379,194,1784]" captionTargetId-2="figure@8.[151,1436,194,1735]" captionTargetId-3="figure@9.[151,1436,194,1853]" captionTargetId-4="figure@10.[151,1436,194,1282]" captionTargetPageId-0="3" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURE 1. Distribution of the genus Vachoniochactas: V. amazonicus (); V. ashleeae (); V. humboldti sp. nov. (); V. lasallei ()." captionText-1="FIGURES 2 – 5. Vachoniochactas humboldti sp. nov. 2 – 3. Male holotype (IAvH-E 100799), dorsal and ventral views. 4 – 5. Female allotype (ICN-MHN-As- 651), dorsal and ventral views. Scale bars = 5 mm." captionText-2="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." captionText-3="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." captionText-4="FIGURES 19 – 21. Vachoniochactas humboldti sp. nov., left hemispermatophore of a male paratype (IAvH-E 100808). 19. External view. 20. Rear view. 21. Internal view. Scale bars = 0.5 mm." httpUri-0="https://zenodo.org/record/183468/files/figure.png" httpUri-1="https://zenodo.org/record/183469/files/figure.png" httpUri-2="https://zenodo.org/record/183470/files/figure.png" httpUri-3="https://zenodo.org/record/183471/files/figure.png" httpUri-4="https://zenodo.org/record/183472/files/figure.png" pageId="2" pageNumber="33">Figures 1–21</figureCitation>
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,
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<tableCitation id="DF4B0376FFF7DA64FE98FC44FE60FC6A" box="[319,406,946,972]" captionStart="TABLE 1" captionStartId="4.[151,239,271,295]" captionTargetBox="[159,1411,405,1700]" captionTargetPageId="4" captionText="TABLE 1. Vachoniochactas humboldti sp. nov., measurements (in mm). L = length, W = width, D = depth." httpUri="http://table.plazi.org/id/C6B66645FFF1DA62FF30FEF9FB1CFE81" pageId="2" pageNumber="33" tableUuid="C6B66645FFF1DA62FF30FEF9FB1CFE81">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DAD36546FFF7DA64FF30FBF4FE1AFB42" pageId="2" pageNumber="33" type="materials_examined">
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<paragraph id="927636CDFFF7DA64FF30FBF4FE6AFBCA" blockId="2.[151,1436,1026,2012]" pageId="2" pageNumber="33">
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<emphasis id="A0BDEADFFFF7DA64FF30FBF4FEDEFBBA" bold="true" box="[151,296,1026,1052]" pageId="2" pageNumber="33">
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<typeStatus id="4D72886FFFF7DA64FF30FBF4FF22FBBA" box="[151,212,1026,1052]" pageId="2" pageNumber="33">Type</typeStatus>
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series.
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</emphasis>
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<emphasis id="A0BDEADFFFF7DA64FE88FBF4FE52FBBD" box="[303,420,1026,1051]" italics="true" pageId="2" pageNumber="33">
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<typeStatus id="4D72886FFFF7DA64FE88FBF4FE6BFBBD" box="[303,413,1026,1051]" pageId="2" pageNumber="33" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="EADE765DFFF7DA64FE0CFBF4FDBAFBBA" box="[427,588,1026,1052]" name="Colombia" pageId="2" pageNumber="33">COLOMBIA</collectingCountry>
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:
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<emphasis id="A0BDEADFFFF7DA64FDF1FBF4FCA4FBBD" box="[598,850,1026,1051]" italics="true" pageId="2" pageNumber="33">Vichada Department:</emphasis>
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||
adult ɗ, Cumaribo, Matavén Forestry, "bosque de planicies arenosas",
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<quantity id="55319B28FFF7DA64FE2FFBDCFE38FBE2" box="[392,462,1066,1092]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="33" unit="m" value="260.0">260m</quantity>
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,
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<geoCoordinate id="F7FD500AFFF7DA64FE7CFBDCFD99FBE2" box="[475,623,1066,1092]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.486944">04°29’13’’N</geoCoordinate>
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<geoCoordinate id="F7FD500AFFF7DA64FDDFFBDCFCE4FBE2" box="[632,786,1066,1092]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.00611">68°00’22’’W</geoCoordinate>
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,
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<date id="E677100DFFF7DA64FCB8FBDCFC23FBE2" box="[799,981,1066,1092]" pageId="2" pageNumber="33" value="2007-03-31">31 March 2007</date>
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–
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<date id="E677100DFFF7DA64FC43FBDCFB78FBE2" box="[996,1166,1066,1092]" pageId="2" pageNumber="33" value="2007-04-02">02 April 2007</date>
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, L.E. Franco coll., pitfall (IAvH-E
|
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<date id="E677100DFFF7DA64FE94FBA4FE66FBCA" box="[307,400,1106,1132]" pageId="2" pageNumber="33" value="99-10-07">100799</date>
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).
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</paragraph>
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<paragraph id="927636CDFFF7DA64FF62FB8DFE1AFB42" blockId="2.[151,1436,1026,2012]" pageId="2" pageNumber="33">
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<emphasis id="A0BDEADFFFF7DA64FF62FB8DFEC6FB32" box="[197,304,1147,1172]" italics="true" pageId="2" pageNumber="33">Allotype:</emphasis>
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<collectingCountry id="EADE765DFFF7DA64FE9BFB8CFE2BFB32" box="[316,477,1146,1172]" name="Colombia" pageId="2" pageNumber="33">COLOMBIA</collectingCountry>
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:
|
||
<emphasis id="A0BDEADFFFF7DA64FE4BFB8DFD1AFB32" box="[492,748,1147,1172]" italics="true" pageId="2" pageNumber="33">Vichada Department:</emphasis>
|
||
adult Ψ, Cumaribo, Matavén Forestry, "bosque de cerro rocoso",
|
||
<quantity id="55319B28FFF7DA64FEA6FB54FEB1FB1A" box="[257,327,1186,1212]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="33" unit="m" value="300.0">300m</quantity>
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,
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||
<geoCoordinate id="F7FD500AFFF7DA64FEF3FB54FE1EFB1A" box="[340,488,1186,1212]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.6091666">04°36’33’’N</geoCoordinate>
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<geoCoordinate id="F7FD500AFFF7DA64FE56FB54FD7DFB1A" box="[497,651,1186,1212]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-67.86444">67°51’52’’W</geoCoordinate>
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, 0
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<date id="E677100DFFF7DA64FD0FFB54FCA6FB1A" box="[680,848,1186,1212]" pageId="2" pageNumber="33" value="2007-03-06">6 March 2007</date>
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–
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<date id="E677100DFFF7DA64FCF8FB54FBEEFB1A" box="[863,1048,1186,1212]" pageId="2" pageNumber="33" value="2007-03-08">08 March 2007</date>
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||
, L.E. Franco coll., trap for dung beetles (ICN-MHN-As-651).
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</paragraph>
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</subSubSection>
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<subSubSection id="DAD36546FFF7DA64FF62FB05FE10F87A" pageId="2" pageNumber="33" type="description">
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<paragraph id="927636CDFFF7DA64FF62FB05FE10F87A" blockId="2.[151,1436,1026,2012]" pageId="2" pageNumber="33">
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<emphasis id="A0BDEADFFFF7DA64FF62FB05FEB1FAAA" box="[197,327,1267,1292]" italics="true" pageId="2" pageNumber="33">
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<typeStatus id="4D72886FFFF7DA64FF62FB05FEC9FAAA" box="[197,319,1267,1292]" pageId="2" pageNumber="33" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectingCountry id="EADE765DFFF7DA64FEF7FB04FE07FAAA" box="[336,497,1266,1292]" name="Colombia" pageId="2" pageNumber="33">COLOMBIA</collectingCountry>
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:
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<emphasis id="A0BDEADFFFF7DA64FE58FB05FD08FAAA" box="[511,766,1267,1292]" italics="true" pageId="2" pageNumber="33">Vichada Department:</emphasis>
|
||
one adult Ψ and two adult ɗ, Cumaribo, Matavén Forestry, "bosque de planicies arenosas",
|
||
<quantity id="55319B28FFF7DA64FDFEFAECFD69FA92" box="[601,671,1306,1332]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="33" unit="m" value="260.0">260m</quantity>
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,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FD0BFAECFCB6FA92" box="[684,832,1306,1332]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.486944">04°29’13’’N</geoCoordinate>
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<geoCoordinate id="F7FD500AFFF7DA64FCEFFAECFC14FA92" box="[840,994,1306,1332]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.00611">68°00’22’’W</geoCoordinate>
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,
|
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<date id="E677100DFFF7DA64FC48FAECFB52FA92" box="[1007,1188,1306,1332]" pageId="2" pageNumber="33" value="2007-03-31">31 March 2007</date>
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–
|
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<date id="E677100DFFF7DA64FB13FAECFAAAFA92" box="[1204,1372,1306,1332]" pageId="2" pageNumber="33" value="2007-04-02">02 April 2007</date>
|
||
, L.E. Franco coll., trap for dung beetles (IAvH-E
|
||
<date id="E677100DFFF7DA64FD3AFAB4FD0EFAFA" box="[669,760,1346,1372]" pageId="2" pageNumber="33" value="96-10-07">100796</date>
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,
|
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<date id="E677100DFFF7DA64FCA2FAB4FC96FAFA" box="[773,864,1346,1372]" pageId="2" pageNumber="33" value="98-10-07">100798</date>
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,
|
||
<date id="E677100DFFF7DA64FCCBFAB4FC3EFAFA" box="[876,968,1346,1372]" pageId="2" pageNumber="33" value="01-10-08">100801</date>
|
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); one adult Ψ, Cumaribo, Matavén Forestry, "bosque de planicies arenosas",
|
||
<quantity id="55319B28FFF7DA64FDFEFA9CFD69FA22" box="[601,671,1386,1412]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="33" unit="m" value="260.0">260m</quantity>
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,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FD0BFA9CFCB6FA22" box="[684,832,1386,1412]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.486944">04°29’13’’N</geoCoordinate>
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<geoCoordinate id="F7FD500AFFF7DA64FCEFFA9CFC14FA22" box="[840,994,1386,1412]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.00611">68°00’22’’W</geoCoordinate>
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,
|
||
<date id="E677100DFFF7DA64FC48FA9CFB52FA22" box="[1007,1188,1386,1412]" pageId="2" pageNumber="33" value="2007-03-31">31 March 2007</date>
|
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–
|
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<date id="E677100DFFF7DA64FB13FA9CFAAAFA22" box="[1204,1372,1386,1412]" pageId="2" pageNumber="33" value="2007-04-02">02 April 2007</date>
|
||
, L.E. Franco coll., trap for dung beetles (MPUJ-SCO-378); six adult ɗ, Cumaribo, Matavén Forestry, "bosque de planicies arenosas",
|
||
<quantity id="55319B28FFF7DA64FE2FFA4CFE38FA72" box="[392,462,1466,1492]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="33" unit="m" value="260.0">260m</quantity>
|
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,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FE7CFA4CFD99FA72" box="[475,623,1466,1492]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.486944">04°29’13’’N</geoCoordinate>
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<geoCoordinate id="F7FD500AFFF7DA64FDDFFA4CFCE4FA72" box="[632,786,1466,1492]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.00611">68°00’22’’W</geoCoordinate>
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,
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<date id="E677100DFFF7DA64FCB8FA4CFC23FA72" box="[799,981,1466,1492]" pageId="2" pageNumber="33" value="2007-03-31">31 March 2007</date>
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–
|
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<date id="E677100DFFF7DA64FC43FA4CFB78FA72" box="[996,1166,1466,1492]" pageId="2" pageNumber="33" value="2007-04-02">02 April 2007</date>
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, L.E. Franco coll., pitfall (IAvH-E
|
||
<date id="E677100DFFF7DA64FE93FA14FE79FA5A" box="[308,399,1506,1532]" pageId="2" pageNumber="33" value="00-10-08">100800</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FE3CFA14FE00FA5A" box="[411,502,1506,1532]" pageId="2" pageNumber="33" value="04-10-08">100804</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FDA5FA14FDABFA5A" box="[514,605,1506,1532]" pageId="2" pageNumber="33" value="05-10-08">100805</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FDCDFA14FD33FA5A" box="[618,709,1506,1532]" pageId="2" pageNumber="33" value="06-10-08">100806</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FD76FA14FCDAFA5A" box="[721,812,1506,1532]" pageId="2" pageNumber="33" value="07-10-08">100807</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FC9FFA14FC63FA5A" box="[824,917,1506,1532]" pageId="2" pageNumber="33" value="08-10-08">100808</date>
|
||
); one adult ɗ, Cumaribo, Matavén Forestry, "bosque de planicies arenosas",
|
||
<quantity id="55319B28FFF7DA64FDAAF9FCFDA5F982" box="[525,595,1546,1572]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="33" unit="m" value="260.0">260m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FDF9F9FCFD04F982" box="[606,754,1546,1572]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.486944">04°29’13’’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FD5EF9FCFC65F982" box="[761,915,1546,1572]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.00611">68°00’22’’W</geoCoordinate>
|
||
,
|
||
<date id="E677100DFFF7DA64FC38F9FCFBA6F982" box="[927,1104,1546,1572]" pageId="2" pageNumber="33" value="2007-03-31">31 March 2007</date>
|
||
–
|
||
<date id="E677100DFFF7DA64FBC7F9FCFAF0F982" box="[1120,1286,1546,1572]" pageId="2" pageNumber="33" value="2007-04-02">02 April 2007</date>
|
||
, L.E. Franco coll., pitfall (MPUJ-SCO-379); one adult ɗ, Cumaribo, Matavén Forestry, "bosque de cerro rocoso",
|
||
<quantity id="55319B28FFF7DA64FAF5F9C4FA61F9EA" box="[1362,1431,1586,1612]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="33" unit="m" value="300.0">300m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FF30F9ACFEDCF9D2" box="[151,298,1626,1652]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.6091666">04°36’33’’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FE94F9ACFE3BF9D2" box="[307,461,1626,1652]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-67.86444">67°51’52’’W</geoCoordinate>
|
||
, 0
|
||
<date id="E677100DFFF7DA64FE4DF9ACFD64F9D2" box="[490,658,1626,1652]" pageId="2" pageNumber="33" value="2007-03-06">6 March 2007</date>
|
||
–
|
||
<date id="E677100DFFF7DA64FD06F9ACFCACF9D2" box="[673,858,1626,1652]" pageId="2" pageNumber="33" value="2007-03-08">08 March 2007</date>
|
||
, L.E. Franco coll., pitfall (IAvH-E
|
||
<date id="E677100DFFF7DA64FB5AF9ACFAACF9D2" box="[1277,1370,1626,1652]" pageId="2" pageNumber="33" value="03-10-08">100803</date>
|
||
); two juvenile ɗ, Cumaribo, Matavén Forestry, "bosque de cerro rocoso",
|
||
<quantity id="55319B28FFF7DA64FC7AF974FBD5F93A" box="[989,1059,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="33" unit="m" value="300.0">300m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FB93F974FB31F93A" box="[1076,1223,1666,1692]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="4.6091666">04°36’33’’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FB74F974FA9BF93A" box="[1235,1389,1666,1692]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-67.86444">67°51’52’’W</geoCoordinate>
|
||
, 0
|
||
<date id="E677100DFFF7DA64FA2AF974FED1F962" pageId="2" pageNumber="33" value="2007-03-06">6 March 2007</date>
|
||
–
|
||
<date id="E677100DFFF7DA64FE91F95CFE19F962" box="[310,495,1706,1732]" pageId="2" pageNumber="33" value="2007-03-08">08 March 2007</date>
|
||
, L.E. Franco coll., Winkler (IAvH-E
|
||
<date id="E677100DFFF7DA64FC0CF95CFBF0F962" box="[939,1030,1706,1732]" pageId="2" pageNumber="33" value="93-10-07">100793</date>
|
||
,
|
||
<date id="E677100DFFF7DA64FBB4F95CFB86F962" box="[1043,1136,1706,1732]" pageId="2" pageNumber="33" value="94-10-07">100794</date>
|
||
); one adult ɗ, Cumaribo, Santa Rita Municipality, El Tuparro Natural National Park, "mata de monte",
|
||
<quantity id="55319B28FFF7DA64FB0DF924FB06F94A" box="[1194,1264,1746,1772]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="33" unit="m" value="135.0">135m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FAAFF924FA6AF94A" box="[1288,1436,1746,1772]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="5.3316665">05°19’54’’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FF30F90CFEC7F8B2" box="[151,305,1786,1812]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-67.89083">67°53’27’’W</geoCoordinate>
|
||
, soil,
|
||
<date id="E677100DFFF7DA64FED0F90CFDBBF8B2" box="[375,589,1786,1812]" pageId="2" pageNumber="33" value="2004-02-10">10 February 2004</date>
|
||
, I. Quintero & E. González coll., pitfall (ICN-MHN-As-652); one juvenile ɗ, Cumaribo, Santa Rita Municipality, El Tuparro Natural National Park, "bosque de cerro",
|
||
<quantity id="55319B28FFF7DA64FAF5F8D4FA61F89A" box="[1362,1431,1826,1852]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="33" unit="m" value="135.0">135m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FF30F8BCFEDEF8C2" box="[151,296,1866,1892]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="15" value="5.3555555">05º21’20’’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FE97F8BCFE3EF8C2" box="[304,456,1866,1892]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="15" value="-68.024445">68º01’28’’W</geoCoordinate>
|
||
, soil,
|
||
<date id="E677100DFFF7DA64FDA8F8BCFD13F8C2" box="[527,741,1866,1892]" pageId="2" pageNumber="33" value="2004-02-20">20 February 2004</date>
|
||
, I. Quintero & E. González coll., pitfall (IAvH-E
|
||
<date id="E677100DFFF7DA64FA94F8BCFA66F8C2" box="[1331,1424,1866,1892]" pageId="2" pageNumber="33" value="95-10-07">100795</date>
|
||
).
|
||
<emphasis id="A0BDEADFFFF7DA64FF30F885FE65F82A" box="[151,403,1907,1932]" italics="true" pageId="2" pageNumber="33">Guainía Department:</emphasis>
|
||
one adult ɗ, Santa Rosa, Santa Rosa Indigenous Community, Bocón Pipe, Inírida River,
|
||
<quantity id="55319B28FFF7DA64FF30F86CFF2AF812" box="[151,220,1946,1972]" metricMagnitude="2" metricUnit="m" metricValue="1.01" pageId="2" pageNumber="33" unit="m" value="101.0">101m</quantity>
|
||
,
|
||
<geoCoordinate id="F7FD500AFFF7DA64FF4DF86CFEB9F812" box="[234,335,1946,1972]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="3.7166667">03°43’N</geoCoordinate>
|
||
<geoCoordinate id="F7FD500AFFF7DA64FEFFF86CFE35F812" box="[344,451,1946,1972]" direction="west" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="-68.066666">68°04’W</geoCoordinate>
|
||
, "bosque maduro de terraza no inundable", sandy soil,
|
||
<date id="E677100DFFF7DA64FBF3F86CFAE0F812" box="[1108,1302,1946,1972]" pageId="2" pageNumber="33" value="1996-11">November 1996</date>
|
||
, A. Lopera coll., pitfall (CDA 000.805).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C6B66645FFF6DA65FF30FB25FEC1FAAD" httpUri="https://zenodo.org/record/183468/files/figure.png" pageId="3" pageNumber="34" targetBox="[151,1435,185,1208]" targetPageId="3">
|
||
<paragraph id="927636CDFFF6DA65FF30FB25FEC1FAAD" blockId="3.[151,1436,1235,1291]" pageId="3" pageNumber="34">
|
||
<emphasis id="A0BDEADFFFF6DA65FF30FB25FEEFFB4D" bold="true" box="[151,281,1235,1259]" pageId="3" pageNumber="34">FIGURE 1.</emphasis>
|
||
Distribution of the genus
|
||
<taxonomicName id="55C94D4EFFF6DA65FD94FB22FD1EFB4D" box="[563,744,1236,1259]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0BDEADFFFF6DA65FD94FB22FD1EFB4D" box="[563,744,1236,1259]" italics="true" pageId="3" pageNumber="34">Vachoniochactas</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="55C94D4EFFF6DA65FD51FB22FC32FB4D" box="[758,964,1236,1259]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="A0BDEADFFFF6DA65FD51FB22FC79FB4D" box="[758,911,1236,1259]" italics="true" pageId="3" pageNumber="34">V. amazonicus</emphasis>
|
||
()
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="55C94D4EFFF6DA65FC76FB22FB81FB4D" box="[977,1143,1236,1259]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ashleeae">
|
||
<emphasis id="A0BDEADFFFF6DA65FC76FB22FBBFFB4D" box="[977,1097,1236,1259]" italics="true" pageId="3" pageNumber="34">V. ashleeae</emphasis>
|
||
()
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="55C94D4EFFF6DA65FB23FB22FAFDFB4D" box="[1156,1291,1236,1259]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF6DA65FB23FB22FAFDFB4D" box="[1156,1291,1236,1259]" italics="true" pageId="3" pageNumber="34">V. humboldti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0BDEADFFFF6DA65FAB5FB25FA9EFB4D" bold="true" box="[1298,1384,1235,1259]" pageId="3" pageNumber="34">
|
||
<taxonomicNameLabel id="BB8E57A4FFF6DA65FAB5FB25FA9EFB4D" box="[1298,1384,1235,1259]" pageId="3" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
();
|
||
<taxonomicName id="55C94D4EFFF6DA65FF30FB02FEC4FAAD" box="[151,306,1268,1291]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="lasallei">
|
||
<emphasis id="A0BDEADFFFF6DA65FF30FB02FF09FAAD" box="[151,255,1268,1291]" italics="true" pageId="3" pageNumber="34">V. lasallei</emphasis>
|
||
()
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DAD36546FFF6DA65FF62FAC5FF16FA60" pageId="3" pageNumber="34" type="etymology">
|
||
<paragraph id="927636CDFFF6DA65FF62FAC5FF16FA60" blockId="3.[151,1437,1331,2038]" pageId="3" pageNumber="34">
|
||
<emphasis id="A0BDEADFFFF6DA65FF62FAC5FEA4FAEB" bold="true" box="[197,338,1331,1357]" pageId="3" pageNumber="34">Etymology.</emphasis>
|
||
The species name is a tribute to the labor of the Instituto de Investigación de Recursos Biológicos Alexander von Humboldt (Villa de Leyva,
|
||
<collectingCountry id="EADE765DFFF6DA65FCB7FAAAFC7EFAD0" box="[784,904,1372,1398]" name="Colombia" pageId="3" pageNumber="34">Colombia</collectingCountry>
|
||
), whose collecting programs in many Colombian areas of difficult access have led to the discovery of important novelties for the Colombian scorpion fauna.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAD36546FFF6DA63FF62FA25FC95FDBC" lastPageId="5" lastPageNumber="36" pageId="3" pageNumber="34" type="diagnosis">
|
||
<paragraph id="927636CDFFF6DA65FF62FA25FE51F850" blockId="3.[151,1437,1331,2038]" pageId="3" pageNumber="34">
|
||
<emphasis id="A0BDEADFFFF6DA65FF62FA25FEB3FA4B" bold="true" box="[197,325,1491,1517]" pageId="3" pageNumber="34">Diagnosis.</emphasis>
|
||
<taxonomicName id="55C94D4EFFF6DA65FEEBFA22FD66FA4B" box="[332,656,1492,1517]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF6DA65FEEBFA22FD66FA4B" box="[332,656,1492,1517]" italics="true" pageId="3" pageNumber="34">Vachoniochactas humboldti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BB8E57A4FFF6DA65FD30FA22FD07FA48" box="[663,753,1492,1518]" pageId="3" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from the other species by (1) the presence of a ventromedian row of setae flanked by three pairs of submedian setae plus a single prolateral distal seta in telotarsi III–IV (
|
||
<figureCitation id="0AF22A48FFF6DA65FF55F9D2FE84F998" box="[242,370,1572,1598]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="3" pageNumber="34">Figs. 9–10</figureCitation>
|
||
), and (2) by having the ventral surface of metasomal segments I–IV completely smooth. In contrast, in all remaining three species the tarsal setae are not arranged in rows (
|
||
<bibRefCitation id="F6584B3CFFF6DA65FB90F9BAFABBF9C0" author="Gonzalez-Sponga" box="[1079,1357,1612,1638]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1978) Escorpiofauna de la Region Oriental del Estado Bolivar, en Venezuela. Roto-Impresos, C. A. Caracas, 217 pp." type="book" year="1978">González-Sponga 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F6584B3CFFF6DA65FAFEF9BAFA61F9C0" author="Gonzalez-Sponga" box="[1369,1431,1612,1638]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1991) Aracnidos de Venezuela. Escorpiones del Tepui " La Neblina ", Territorio Federal Amazonas, Venezuela (Scorpionida: Chactidae: Buthidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales (Caracas), 50 (163 - 164), 11 - 62." type="journal article" year="1991">1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F6584B3CFFF6DA65FF30F982FEBDF928" author="Lourenco" box="[151,331,1652,1678]" pageId="3" pageNumber="43" refString="Lourenco, W. R. (1994) Scorpion biogeographic patterns as evidence for a Neblina-Sao Gabriel endemic center in Brazilian Amazonia. Revista de la Academia Colombiana de Ciencias, 19 (72), 181 - 185." type="journal article" year="1994">Lourenço 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F6584B3CFFF6DA65FEF0F982FE55F928" author="Lourenco" box="[343,419,1652,1678]" pageId="3" pageNumber="43" refString="Lourenco, W. R. (2002 a) Scorpions of Brazil. Les Editions de l'If, Paris, 307 pp." type="book" year="2002" yearSuffix="a">2002a</bibRefCitation>
|
||
), and metasomal segments III–IV (and I–II in
|
||
<taxonomicName id="55C94D4EFFF6DA65FC19F982FBC7F92B" box="[958,1073,1652,1677]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="lasallei">
|
||
<emphasis id="A0BDEADFFFF6DA65FC19F982FBC7F92B" box="[958,1073,1652,1677]" italics="true" pageId="3" pageNumber="34">V. lasallei</emphasis>
|
||
</taxonomicName>
|
||
) have vestigial granules on the ventral surface (
|
||
<bibRefCitation id="F6584B3CFFF6DA65FEFEF96AFD85F910" author="Gonzalez-Sponga" box="[345,627,1692,1718]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1978) Escorpiofauna de la Region Oriental del Estado Bolivar, en Venezuela. Roto-Impresos, C. A. Caracas, 217 pp." type="book" year="1978">González-Sponga 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F6584B3CFFF6DA65FD25F96AFD49F910" author="Gonzalez-Sponga" box="[642,703,1692,1718]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1991) Aracnidos de Venezuela. Escorpiones del Tepui " La Neblina ", Territorio Federal Amazonas, Venezuela (Scorpionida: Chactidae: Buthidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales (Caracas), 50 (163 - 164), 11 - 62." type="journal article" year="1991">1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F6584B3CFFF6DA65FD69F96AFCFAF910" author="Gonzalez-Sponga" box="[718,780,1692,1718]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1996) Guia para identificar escorpiones de Venezuela. Cuadernos Lagoven, Caracas, 204 pp." type="book" year="1996">1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F6584B3CFFF6DA65FCBCF96AFC23F910" author="Lourenco" box="[795,981,1692,1718]" pageId="3" pageNumber="43" refString="Lourenco, W. R. (1994) Scorpion biogeographic patterns as evidence for a Neblina-Sao Gabriel endemic center in Brazilian Amazonia. Revista de la Academia Colombiana de Ciencias, 19 (72), 181 - 185." type="journal article" year="1994">Lourenço 1994</bibRefCitation>
|
||
). The new species can also be distinguished from
|
||
<emphasis id="A0BDEADFFFF6DA65FE9DF932FE5EF97B" box="[314,424,1732,1757]" italics="true" pageId="3" pageNumber="34">V. l a s a l l e i</emphasis>
|
||
because the vesicle of telson has two to three aligned subaculear tubercles in the later (see
|
||
<bibRefCitation id="F6584B3CFFF6DA65FF69F91AFE10F8A0" author="Gonzalez-Sponga" box="[206,486,1772,1798]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1978) Escorpiofauna de la Region Oriental del Estado Bolivar, en Venezuela. Roto-Impresos, C. A. Caracas, 217 pp." type="book" year="1978">González-Sponga 1978</bibRefCitation>
|
||
: fig. 65), which are absent in
|
||
<taxonomicName id="55C94D4EFFF6DA65FCEFF91AFC2BF8A3" box="[840,989,1772,1797]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF6DA65FCEFF91AFC2BF8A3" box="[840,989,1772,1797]" italics="true" pageId="3" pageNumber="34">V. humboldti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BB8E57A4FFF6DA65FC42F91AFBB7F8A0" box="[997,1089,1772,1798]" pageId="3" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="0AF22A48FFF6DA65FBF6F91AFB6FF8A0" box="[1105,1177,1772,1798]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="3" pageNumber="34">Fig. 8</figureCitation>
|
||
). In addition,
|
||
<taxonomicName id="55C94D4EFFF6DA65FAE5F91AFF2DF88B" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF6DA65FAE5F91AFF2DF88B" italics="true" pageId="3" pageNumber="34">V. humboldti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BB8E57A4FFF6DA65FF43F8E2FEB6F888" box="[228,320,1812,1838]" pageId="3" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="55C94D4EFFF6DA65FE78F8E2FD71F88B" box="[479,647,1812,1837]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="A0BDEADFFFF6DA65FE78F8E2FD71F88B" box="[479,647,1812,1837]" italics="true" pageId="3" pageNumber="34">V. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="55C94D4EFFF6DA65FD63F8E2FCCEF88B" box="[708,824,1812,1837]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="lasallei">
|
||
<emphasis id="A0BDEADFFFF6DA65FD63F8E2FCCEF88B" box="[708,824,1812,1837]" italics="true" pageId="3" pageNumber="34">V. lasallei</emphasis>
|
||
</taxonomicName>
|
||
by the position of trichobothrium
|
||
<emphasis id="A0BDEADFFFF6DA65FB72F8E2FB1AF88B" box="[1237,1260,1812,1837]" italics="true" pageId="3" pageNumber="34">dt</emphasis>
|
||
, basal to
|
||
<emphasis id="A0BDEADFFFF6DA65FAC6F8E2FA80F88B" box="[1377,1398,1812,1837]" italics="true" pageId="3" pageNumber="34">et</emphasis>
|
||
on pedipalp fixed finger (
|
||
<figureCitation id="0AF22A48FFF6DA65FE3AF8CAFE05F8F0" box="[413,499,1852,1878]" captionStart="FIGURES 11 – 18" captionStartId="9.[151,269,1877,1901]" captionTargetBox="[177,1431,214,1817]" captionTargetId="figure@9.[151,1436,194,1853]" captionTargetPageId="9" captionText="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183471/files/figure.png" pageId="3" pageNumber="34">Fig. 11</figureCitation>
|
||
); in the two later species
|
||
<emphasis id="A0BDEADFFFF6DA65FC87F8CAFCC0F8F3" box="[800,822,1852,1877]" italics="true" pageId="3" pageNumber="34">dt</emphasis>
|
||
is distal to
|
||
<emphasis id="A0BDEADFFFF6DA65FC18F8CAFC22F8F3" box="[959,980,1852,1877]" italics="true" pageId="3" pageNumber="34">et</emphasis>
|
||
(see
|
||
<bibRefCitation id="F6584B3CFFF6DA65FBB5F8CAFADFF8F0" author="Gonzalez-Sponga" box="[1042,1321,1852,1878]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1978) Escorpiofauna de la Region Oriental del Estado Bolivar, en Venezuela. Roto-Impresos, C. A. Caracas, 217 pp." type="book" year="1978">González-Sponga 1978</bibRefCitation>
|
||
: figs. 46– 47; 1991: figs. 48–49). Finally, it can be also distinguished from
|
||
<taxonomicName id="55C94D4EFFF6DA65FC39F892FBB1F8DB" box="[926,1095,1892,1917]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="amazonicus">
|
||
<emphasis id="A0BDEADFFFF6DA65FC39F892FBB1F8DB" box="[926,1095,1892,1917]" italics="true" pageId="3" pageNumber="34">V. amazonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="55C94D4EFFF6DA65FB25F892FAF0F8DB" box="[1154,1286,1892,1917]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="ashleeae">
|
||
<emphasis id="A0BDEADFFFF6DA65FB25F892FAF0F8DB" box="[1154,1286,1892,1917]" italics="true" pageId="3" pageNumber="34">V. ashleeae</emphasis>
|
||
</taxonomicName>
|
||
because
|
||
<emphasis id="A0BDEADFFFF6DA65FAD2F892FA6AF8DB" box="[1397,1436,1892,1917]" italics="true" pageId="3" pageNumber="34">esb</emphasis>
|
||
trichobothrium is considerably more basal to
|
||
<emphasis id="A0BDEADFFFF6DA65FD17F87AFD3DF803" box="[688,715,1932,1957]" italics="true" pageId="3" pageNumber="34">eb</emphasis>
|
||
on the fixed finger (
|
||
<figureCitation id="0AF22A48FFF6DA65FC67F87AFBE1F800" box="[960,1047,1932,1958]" captionStart="FIGURES 11 – 18" captionStartId="9.[151,269,1877,1901]" captionTargetBox="[177,1431,214,1817]" captionTargetId="figure@9.[151,1436,194,1853]" captionTargetPageId="9" captionText="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183471/files/figure.png" pageId="3" pageNumber="34">Fig. 12</figureCitation>
|
||
), contrasting with the mentioned species in which
|
||
<emphasis id="A0BDEADFFFF6DA65FED4F842FE6CF86B" box="[371,410,1972,1997]" italics="true" pageId="3" pageNumber="34">esb</emphasis>
|
||
is located close or slightly distal to
|
||
<emphasis id="A0BDEADFFFF6DA65FCD3F842FC66F86B" box="[884,912,1972,1997]" italics="true" pageId="3" pageNumber="34">eb</emphasis>
|
||
(see
|
||
<bibRefCitation id="F6584B3CFFF6DA65FC7CF842FB0EF868" author="Gonzalez-Sponga" box="[987,1272,1972,1998]" pageId="3" pageNumber="43" refString="Gonzalez-Sponga, M. A. (1991) Aracnidos de Venezuela. Escorpiones del Tepui " La Neblina ", Territorio Federal Amazonas, Venezuela (Scorpionida: Chactidae: Buthidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales (Caracas), 50 (163 - 164), 11 - 62." type="journal article" year="1991">González-Sponga 1991</bibRefCitation>
|
||
: figs. 48–49;
|
||
<bibRefCitation id="F6584B3CFFF6DA65FF30F82AFEBBF850" author="Lourenco" box="[151,333,2012,2038]" pageId="3" pageNumber="43" refString="Lourenco, W. R. (1994) Scorpion biogeographic patterns as evidence for a Neblina-Sao Gabriel endemic center in Brazilian Amazonia. Revista de la Academia Colombiana de Ciencias, 19 (72), 181 - 185." type="journal article" year="1994">Lourenço 1994</bibRefCitation>
|
||
: fig. 6).
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF1DA62FF62FF6EFCD3FF14" blockId="4.[197,944,152,218]" box="[197,805,152,178]" pageId="4" pageNumber="35">
|
||
Description of the male
|
||
<typeStatus id="4D72886FFFF1DA62FE47FF6EFDB1FF14" box="[480,583,152,178]" pageId="4" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
(IAvH-E
|
||
<date id="E677100DFFF1DA62FD1CFF6EFCEEFF14" box="[699,792,152,178]" pageId="4" pageNumber="35" value="99-10-07">100799</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF1DA62FF62FF36FC46FF7C" blockId="4.[197,944,152,218]" box="[197,944,192,218]" pageId="4" pageNumber="35">
|
||
Total body length 23.15 mm; detailed measurements in
|
||
<tableCitation id="DF4B0376FFF1DA62FCF5FF36FC5CFF7C" box="[850,938,192,218]" captionStart="TABLE 1" captionStartId="4.[151,239,271,295]" captionTargetBox="[159,1411,405,1700]" captionTargetPageId="4" captionText="TABLE 1. Vachoniochactas humboldti sp. nov., measurements (in mm). L = length, W = width, D = depth." httpUri="http://table.plazi.org/id/C6B66645FFF1DA62FF30FEF9FB1CFE81" pageId="4" pageNumber="35" tableUuid="C6B66645FFF1DA62FF30FEF9FB1CFE81">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="C6B66645FFF1DA62FF30FEF9FB1CFE81" ID-Table-UUID="C6B66645FFF1DA62FF30FEF9FB1CFE81" box="[151,1258,271,295]" httpUri="http://table.plazi.org/id/C6B66645FFF1DA62FF30FEF9FB1CFE81" pageId="4" pageNumber="35" targetBox="[159,1411,405,1700]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="927636CDFFF1DA62FF30FEF9FB1CFE81" blockId="4.[151,1258,271,295]" box="[151,1258,271,295]" pageId="4" pageNumber="35">
|
||
<emphasis id="A0BDEADFFFF1DA62FF30FEF9FEFEFE81" bold="true" box="[151,264,271,295]" pageId="4" pageNumber="35">TABLE 1.</emphasis>
|
||
<taxonomicName id="55C94D4EFFF1DA62FEB7FEE6FDCEFE81" box="[272,568,272,295]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF1DA62FEB7FEE6FDCEFE81" box="[272,568,272,295]" italics="true" pageId="4" pageNumber="35">Vachoniochactas humboldti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A0BDEADFFFF1DA62FD99FEF9FD62FE81" bold="true" box="[574,660,271,295]" pageId="4" pageNumber="35">
|
||
<taxonomicNameLabel id="BB8E57A4FFF1DA62FD99FEF9FD62FE81" box="[574,660,271,295]" pageId="4" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, measurements (in mm). L=length, W=width, D=depth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="927636CDFFF1DA62FCBBFEBEFB54FE27" blockId="4.[796,1411,328,385]" pageId="4" pageNumber="35">
|
||
Male
|
||
<typeStatus id="4D72886FFFF1DA62FCFDFEBEFC41FEF9" box="[858,951,328,351]" pageId="4" pageNumber="35" type="holotype">holotype</typeStatus>
|
||
(IAvH-E Female allotype (ICN-MHN-
|
||
<date id="E677100DFFF1DA62FCBBFE9CFC98FE27" box="[796,878,362,385]" pageId="4" pageNumber="35" value="99-10-07">100799</date>
|
||
) As-651)
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF1DA62FF38FE63FB8FF902" pageId="4" pageNumber="35">
|
||
<table id="E0C9C46DFFF12599FF38FE63FA75F902" box="[159,1411,405,1700]" gridcols="3" gridrows="20" pageId="4" pageNumber="35">
|
||
<tr id="2CF9348FFFF12599FF38FE63FA75FE5F" box="[159,1411,405,505]" gridrow="0" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FE63FD43FE5F" box="[159,693,405,505]" gridcol="0" gridrow="0" pageId="4" pageNumber="35">Total body L (includ- ing telson) Carapace Length</th>
|
||
<th id="6F285DF3FFF12599FCBBFE63FCAEFE5F" box="[796,856,405,505]" gridcol="1" gridrow="0" pageId="4" pageNumber="35">23.15 3.25</th>
|
||
<th id="6F285DF3FFF12599FBEDFE63FA75FE5F" box="[1098,1411,405,505]" gridcol="2" gridrow="0" pageId="4" pageNumber="35">24.31 3.55</th>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FDFBFA75FD82" box="[159,1411,525,548]" gridrow="1" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FDFBFD43FD82" box="[159,693,525,548]" gridcol="0" gridrow="1" pageId="4" pageNumber="35">Anterior W</th>
|
||
<td id="6F285DF3FFF12599FCBBFDFBFCAEFD82" box="[796,856,525,548]" gridcol="1" gridrow="1" pageId="4" pageNumber="35">1.90</td>
|
||
<td id="6F285DF3FFF12599FBEDFDFBFA75FD82" box="[1098,1411,525,548]" gridcol="2" gridrow="1" pageId="4" pageNumber="35">2.27</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FDCEFA75FDDF" box="[159,1411,568,633]" gridrow="2" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FDCEFD43FDDF" box="[159,693,568,633]" gridcol="0" gridrow="2" pageId="4" pageNumber="35">Posterior W Mesosoma Total L</th>
|
||
<td id="6F285DF3FFF12599FCBBFDCEFCAEFDDF" box="[796,856,568,633]" gridcol="1" gridrow="2" pageId="4" pageNumber="35">3.40 6.65</td>
|
||
<td id="6F285DF3FFF12599FBEDFDCEFA75FDDF" box="[1098,1411,568,633]" gridcol="2" gridrow="2" pageId="4" pageNumber="35">3.73 7.83</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FD7BFA75FD02" box="[159,1411,653,676]" gridrow="3" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FD7BFD43FD02" box="[159,693,653,676]" gridcol="0" gridrow="3" pageId="4" pageNumber="35">Metasoma Total L (including telson)</th>
|
||
<td id="6F285DF3FFF12599FCBBFD7BFCAEFD02" box="[796,856,653,676]" gridcol="1" gridrow="3" pageId="4" pageNumber="35">13.25</td>
|
||
<td id="6F285DF3FFF12599FBEDFD7BFA75FD02" box="[1098,1411,653,676]" gridcol="2" gridrow="3" pageId="4" pageNumber="35">12.93</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FD4EFA75FD5F" box="[159,1411,696,761]" gridrow="4" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FD4EFD43FD5F" box="[159,693,696,761]" gridcol="0" gridrow="4" pageId="4" pageNumber="35">Segment I L Segment I W</th>
|
||
<td id="6F285DF3FFF12599FCBBFD4EFCAEFD5F" box="[796,856,696,761]" gridcol="1" gridrow="4" pageId="4" pageNumber="35">0.95 2.25</td>
|
||
<td id="6F285DF3FFF12599FBEDFD4EFA75FD5F" box="[1098,1411,696,761]" gridcol="2" gridrow="4" pageId="4" pageNumber="35">1.06 2.47</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FCFBFA75FC82" box="[159,1411,781,804]" gridrow="5" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FCFBFD43FC82" box="[159,693,781,804]" gridcol="0" gridrow="5" pageId="4" pageNumber="35">Segment II L</th>
|
||
<td id="6F285DF3FFF12599FCBBFCFBFCAEFC82" box="[796,856,781,804]" gridcol="1" gridrow="5" pageId="4" pageNumber="35">1.40</td>
|
||
<td id="6F285DF3FFF12599FBEDFCFBFA75FC82" box="[1098,1411,781,804]" gridcol="2" gridrow="5" pageId="4" pageNumber="35">1.40</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FCCEFA75FCDF" box="[159,1411,824,889]" gridrow="6" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FCCEFD43FCDF" box="[159,693,824,889]" gridcol="0" gridrow="6" pageId="4" pageNumber="35">Segment II W Segment III L</th>
|
||
<td id="6F285DF3FFF12599FCBBFCCEFCAEFCDF" box="[796,856,824,889]" gridcol="1" gridrow="6" pageId="4" pageNumber="35">2.20 1.50</td>
|
||
<td id="6F285DF3FFF12599FBEDFCCEFA75FCDF" box="[1098,1411,824,889]" gridcol="2" gridrow="6" pageId="4" pageNumber="35">2.28 1.55</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FC7BFA75FC02" box="[159,1411,909,932]" gridrow="7" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FC7BFD43FC02" box="[159,693,909,932]" gridcol="0" gridrow="7" pageId="4" pageNumber="35">Segment III W</th>
|
||
<td id="6F285DF3FFF12599FCBBFC7BFCAEFC02" box="[796,856,909,932]" gridcol="1" gridrow="7" pageId="4" pageNumber="35">2.26</td>
|
||
<td id="6F285DF3FFF12599FBEDFC7BFA75FC02" box="[1098,1411,909,932]" gridcol="2" gridrow="7" pageId="4" pageNumber="35">2.30</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FC4EFA75FC5F" box="[159,1411,952,1017]" gridrow="8" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FC4EFD43FC5F" box="[159,693,952,1017]" gridcol="0" gridrow="8" pageId="4" pageNumber="35">Segment IV L Segment IV W</th>
|
||
<td id="6F285DF3FFF12599FCBBFC4EFCAEFC5F" box="[796,856,952,1017]" gridcol="1" gridrow="8" pageId="4" pageNumber="35">2.15 2.30</td>
|
||
<td id="6F285DF3FFF12599FBEDFC4EFA75FC5F" box="[1098,1411,952,1017]" gridcol="2" gridrow="8" pageId="4" pageNumber="35">2.12 2.36</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FBFBFA75FB82" box="[159,1411,1037,1060]" gridrow="9" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FBFBFD43FB82" box="[159,693,1037,1060]" gridcol="0" gridrow="9" pageId="4" pageNumber="35">Segment V L</th>
|
||
<td id="6F285DF3FFF12599FCBBFBFBFCAEFB82" box="[796,856,1037,1060]" gridcol="1" gridrow="9" pageId="4" pageNumber="35">3.50</td>
|
||
<td id="6F285DF3FFF12599FBEDFBFBFA75FB82" box="[1098,1411,1037,1060]" gridcol="2" gridrow="9" pageId="4" pageNumber="35">3.25</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FBCEFA75FBDF" box="[159,1411,1080,1145]" gridrow="10" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FBCEFD43FBDF" box="[159,693,1080,1145]" gridcol="0" gridrow="10" pageId="4" pageNumber="35">Segment V W Telson L</th>
|
||
<td id="6F285DF3FFF12599FCBBFBCEFCAEFBDF" box="[796,856,1080,1145]" gridcol="1" gridrow="10" pageId="4" pageNumber="35">2.33 3.75</td>
|
||
<td id="6F285DF3FFF12599FBEDFBCEFA75FBDF" box="[1098,1411,1080,1145]" gridcol="2" gridrow="10" pageId="4" pageNumber="35">2.44 3.55</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FB7BFA75FB02" box="[159,1411,1165,1188]" gridrow="11" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FB7BFD43FB02" box="[159,693,1165,1188]" gridcol="0" gridrow="11" pageId="4" pageNumber="35">Aculeus L</th>
|
||
<td id="6F285DF3FFF12599FCBBFB7BFCAEFB02" box="[796,856,1165,1188]" gridcol="1" gridrow="11" pageId="4" pageNumber="35">0.9</td>
|
||
<td id="6F285DF3FFF12599FBEDFB7BFA75FB02" box="[1098,1411,1165,1188]" gridcol="2" gridrow="11" pageId="4" pageNumber="35">0.8</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FB4EFA75FB5F" box="[159,1411,1208,1273]" gridrow="12" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FB4EFD43FB5F" box="[159,693,1208,1273]" gridcol="0" gridrow="12" pageId="4" pageNumber="35">Vesicle L Vesicle W</th>
|
||
<td id="6F285DF3FFF12599FCBBFB4EFCAEFB5F" box="[796,856,1208,1273]" gridcol="1" gridrow="12" pageId="4" pageNumber="35">2.85 1.94</td>
|
||
<td id="6F285DF3FFF12599FBEDFB4EFA75FB5F" box="[1098,1411,1208,1273]" gridcol="2" gridrow="12" pageId="4" pageNumber="35">2.75 1.75</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FAFBFA75FA82" box="[159,1411,1293,1316]" gridrow="13" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FAFBFD43FA82" box="[159,693,1293,1316]" gridcol="0" gridrow="13" pageId="4" pageNumber="35">Vesicle D</th>
|
||
<td id="6F285DF3FFF12599FCBBFAFBFCAEFA82" box="[796,856,1293,1316]" gridcol="1" gridrow="13" pageId="4" pageNumber="35">1.30</td>
|
||
<td id="6F285DF3FFF12599FBEDFAFBFA75FA82" box="[1098,1411,1293,1316]" gridcol="2" gridrow="13" pageId="4" pageNumber="35">1.20</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FACEFA75FADF" box="[159,1411,1336,1401]" gridrow="14" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FACEFD43FADF" box="[159,693,1336,1401]" gridcol="0" gridrow="14" pageId="4" pageNumber="35">Pedipalps Total L Femur L</th>
|
||
<td id="6F285DF3FFF12599FCBBFACEFCAEFADF" box="[796,856,1336,1401]" gridcol="1" gridrow="14" pageId="4" pageNumber="35">8.60 2.00</td>
|
||
<td id="6F285DF3FFF12599FBEDFACEFA75FADF" box="[1098,1411,1336,1401]" gridcol="2" gridrow="14" pageId="4" pageNumber="35">9.23 2.28</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FA7BFA75FA02" box="[159,1411,1421,1444]" gridrow="15" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FA7BFD43FA02" box="[159,693,1421,1444]" gridcol="0" gridrow="15" pageId="4" pageNumber="35">Femur W</th>
|
||
<td id="6F285DF3FFF12599FCBBFA7BFCAEFA02" box="[796,856,1421,1444]" gridcol="1" gridrow="15" pageId="4" pageNumber="35">0.95</td>
|
||
<td id="6F285DF3FFF12599FBEDFA7BFA75FA02" box="[1098,1411,1421,1444]" gridcol="2" gridrow="15" pageId="4" pageNumber="35">1.07</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38FA4EFA75FA5F" box="[159,1411,1464,1529]" gridrow="16" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38FA4EFD43FA5F" box="[159,693,1464,1529]" gridcol="0" gridrow="16" pageId="4" pageNumber="35">Patella L Patella W</th>
|
||
<td id="6F285DF3FFF12599FCBBFA4EFCAEFA5F" box="[796,856,1464,1529]" gridcol="1" gridrow="16" pageId="4" pageNumber="35">2.50 1.02</td>
|
||
<td id="6F285DF3FFF12599FBEDFA4EFA75FA5F" box="[1098,1411,1464,1529]" gridcol="2" gridrow="16" pageId="4" pageNumber="35">2.52 1.05</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38F9FBFA75F982" box="[159,1411,1549,1572]" gridrow="17" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38F9FBFD43F982" box="[159,693,1549,1572]" gridcol="0" gridrow="17" pageId="4" pageNumber="35">Chela L</th>
|
||
<td id="6F285DF3FFF12599FCBBF9FBFCAEF982" box="[796,856,1549,1572]" gridcol="1" gridrow="17" pageId="4" pageNumber="35">4.10</td>
|
||
<td id="6F285DF3FFF12599FBEDF9FBFA75F982" box="[1098,1411,1549,1572]" gridcol="2" gridrow="17" pageId="4" pageNumber="35">4.43</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38F9CEFA75F9DF" box="[159,1411,1592,1657]" gridrow="18" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38F9CEFD43F9DF" box="[159,693,1592,1657]" gridcol="0" gridrow="18" pageId="4" pageNumber="35">Chela W Chela D</th>
|
||
<td id="6F285DF3FFF12599FCBBF9CEFCAEF9DF" box="[796,856,1592,1657]" gridcol="1" gridrow="18" pageId="4" pageNumber="35">1.77 1.41</td>
|
||
<td id="6F285DF3FFF12599FBEDF9CEFA75F9DF" box="[1098,1411,1592,1657]" gridcol="2" gridrow="18" pageId="4" pageNumber="35">2.05 1.77</td>
|
||
</tr>
|
||
<tr id="2CF9348FFFF12599FF38F97BFA75F902" box="[159,1411,1677,1700]" gridrow="19" pageId="4" pageNumber="35">
|
||
<th id="6F285DF3FFF12599FF38F97BFD43F902" box="[159,693,1677,1700]" gridcol="0" gridrow="19" pageId="4" pageNumber="35">Movable finger L</th>
|
||
<td id="6F285DF3FFF12599FCBBF97BFCAEF902" box="[796,856,1677,1700]" gridcol="1" gridrow="19" pageId="4" pageNumber="35">2.07</td>
|
||
<td id="6F285DF3FFF12599FBEDF97BFA75F902" box="[1098,1411,1677,1700]" gridcol="2" gridrow="19" pageId="4" pageNumber="35">2.16</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF1DA63FF62F903FC95FDBC" blockId="4.[151,1437,1781,2007]" lastBlockId="5.[151,1437,152,2018]" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35">
|
||
<emphasis id="A0BDEADFFFF1DA62FF62F903FEB9F8A9" bold="true" box="[197,335,1781,1807]" italics="true" pageId="4" pageNumber="35">Coloration.</emphasis>
|
||
Carapace dark reddish-brown with some lighter regions, mainly in the median and posterolateral furrows; median and lateral ocular tubercles black. Tergites dark reddish-brown. Coxosternal region predominantly dark-yellow, with conspicuous brown areas on all the components. Genital operculum and pectinal basal piece yellow; pectines light-yellow, except for the basal piece of the basal lamella which is darkened near the junction with pectinal basal piece. Sternites dark-yellow, gradually becoming reddish towards sternite VII, all with inconspicuous brownish areas on the posterolateral margins. Metasomal segments reddish-brown and iridescent, with inconspicuous dark-brown mottling throughout. Telson vesicle reddish, lighter than segment V; aculeus darkened. Chelicerae predominantly yellow; manus dorsally with broad brown longitudinal reticulations and others thinner located between these, predominantly brown below junction of the movable finger; fixed finger with minute brown mottling basally and reddish teeth; movable finger with a dorsoexternal brown area located basally and reddish teeth. Pedipalps with base color red and insertion of all the trichobothria yellow; femur predominantly dark-brown dorsally and internally, yellowish-red externally and ventrally but with some brownish areas; patella variegated on dorsal and external surfaces, almost completely yellowish-red ventrally and internally; hand with longitudinal brown bands in all surfaces, suggesting the corresponding position of the carinae; fixed and movable fingers red. Legs with femur and patella predominantly brown; other segments predominantly yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAD36546FFF0DA60FF62FDD1FE4AFAC4" lastPageId="6" lastPageNumber="37" pageId="5" pageNumber="36" type="description">
|
||
<paragraph id="927636CDFFF0DA60FF62FDD1FD98FEDC" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,1778]" lastPageId="6" lastPageNumber="37" pageId="5" pageNumber="36">
|
||
<emphasis id="A0BDEADFFFF0DA63FF62FDD1FE90FDE7" bold="true" box="[197,358,551,577]" pageId="5" pageNumber="36">Morphology.</emphasis>
|
||
<emphasis id="A0BDEADFFFF0DA63FECAFDDEFE1DFDE7" box="[365,491,552,577]" italics="true" pageId="5" pageNumber="36">Carapace:</emphasis>
|
||
Uniformly covered with dense thin granulation, except for the carapace furrows which exhibit weaker granulation; granules are stronger between the lateral ocular tubercles; median and lateral ocular tubercles smooth; anterior and posterior margins of carapace almost straight, lateral margins not parallel (
|
||
<figureCitation id="0AF22A48FFF0DA63FF59FD56FEB0FD1C" box="[254,326,672,698]" captionStart="FIGURES 2 – 5" captionStartId="7.[151,269,1808,1832]" captionTargetBox="[207,1379,195,1783]" captionTargetId="figure@7.[207,1379,194,1784]" captionTargetPageId="7" captionText="FIGURES 2 – 5. Vachoniochactas humboldti sp. nov. 2 – 3. Male holotype (IAvH-E 100799), dorsal and ventral views. 4 – 5. Female allotype (ICN-MHN-As- 651), dorsal and ventral views. Scale bars = 5 mm." httpUri="https://zenodo.org/record/183469/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
|
||
); anteromedian furrow vestigial, reduced to a slight depression; posterotransverse and posteromedian furrows pronounced; lateral ocular and central lateral carinae represented by slight elevations, from which the carapace is declined until the lateral margins; other carinae absent; median ocular tubercle rhomboidal, located in the anterior half of the carapace (
|
||
<figureCitation id="0AF22A48FFF0DA63FD71FCEEFCD6FC94" box="[726,800,792,818]" captionStart="FIGURES 2 – 5" captionStartId="7.[151,269,1808,1832]" captionTargetBox="[207,1379,195,1783]" captionTargetId="figure@7.[207,1379,194,1784]" captionTargetPageId="7" captionText="FIGURES 2 – 5. Vachoniochactas humboldti sp. nov. 2 – 3. Male holotype (IAvH-E 100799), dorsal and ventral views. 4 – 5. Female allotype (ICN-MHN-As- 651), dorsal and ventral views. Scale bars = 5 mm." httpUri="https://zenodo.org/record/183469/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
|
||
); lateral ocular tubercles each with two equally sized eyes and two small ocelli behind them.
|
||
<emphasis id="A0BDEADFFFF0DA63FDD9FCB6FD14FCFF" box="[638,738,832,857]" italics="true" pageId="5" pageNumber="36">Tergites:</emphasis>
|
||
With similar granulation to that of the carapace; without median or dorsolateral carinae; each tergite with a clearly defined pretergite; tergite VII with vestigial and incomplete lateral carinae which are absent on the preceding tergites.
|
||
<emphasis id="A0BDEADFFFF0DA63FC63FC66FB46FC0F" box="[964,1200,912,937]" italics="true" pageId="5" pageNumber="36">Coxosternal region:</emphasis>
|
||
Sternum wider than long, slit-like (
|
||
<figureCitation id="0AF22A48FFF0DA63FEE4FC4EFE7DFC74" box="[323,395,952,978]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 6</figureCitation>
|
||
); all the components of the coxosternal region smooth, almost without granules but with numerous setae.
|
||
<emphasis id="A0BDEADFFFF0DA63FEFCFC16FD2CFC5F" box="[347,730,992,1017]" italics="true" pageId="5" pageNumber="36">Genital operculum and pectines:</emphasis>
|
||
Genital operculum divided longitudinally, composed of two subtriangular plates; pectinal basal piece wider than long, with anteromedian notch (
|
||
<figureCitation id="0AF22A48FFF0DA63FB35FBFEFB2AFB84" box="[1170,1244,1032,1058]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 6</figureCitation>
|
||
); pectines long, surpassing the lateral margins of sternite III, without fulcra and with 8 thick teeth each, of which the distal one is rounded (
|
||
<figureCitation id="0AF22A48FFF0DA63FE82FBAEFE99FBD4" box="[293,367,1112,1138]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 6</figureCitation>
|
||
); basal and middle lamellae with abundant setae.
|
||
<emphasis id="A0BDEADFFFF0DA63FC62FBAEFBC3FBD7" box="[965,1077,1112,1137]" italics="true" pageId="5" pageNumber="36">Sternites:</emphasis>
|
||
Densely covered with minute granulation uniformly distributed, and with few setae especially on the lateral margins; without lateral or submedian carinae; spiracles small and round.
|
||
<emphasis id="A0BDEADFFFF0DA63FD3FFB5EFCEAFB67" box="[664,796,1192,1217]" italics="true" pageId="5" pageNumber="36">Metasoma:</emphasis>
|
||
Metasomal segments with abundant setae of different sizes (
|
||
<figureCitation id="0AF22A48FFF0DA63FF78FB26FED0FB4C" box="[223,294,1232,1258]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 8</figureCitation>
|
||
); lateral margins parallel; dorsal surface of all segments deeply depressed; segments I–IV with an anterotransverse suture; segments I–IV with granulose dorsal carinae, evident and complete, and dorsolateral carinae represented by a large anterior tubercle and another posterior; segment V with only dorsolateral carinae (
|
||
<figureCitation id="0AF22A48FFF0DA63FF75FABEFEEDFAC4" box="[210,283,1352,1378]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 8</figureCitation>
|
||
); segment I with few granules located between both dorsal carinae, other segments smooth dorsally; segments I–II with few granules laterally, III–IV almost completely smooth except for some granules occurring in the dorsolateral surfaces (
|
||
<figureCitation id="0AF22A48FFF0DA63FDFFFA6EFD68FA14" box="[600,670,1432,1458]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 8</figureCitation>
|
||
); ventral surface of segments I–IV completely smooth and shiny; segment V with abundant and strong granulation in ventral and lateral surfaces, except basally where the granules are scarce (
|
||
<figureCitation id="0AF22A48FFF0DA63FEF4FA1EFE6CF9A4" box="[339,410,1512,1538]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 8</figureCitation>
|
||
). Telson with an elongated vesicle and a short and curved aculeus; vesicle smooth dorsally, with dense granulation ventrally and laterally similar to that on segment V but rather stronger; with abundant setae ventrally and laterally, especially under the aculeus (
|
||
<figureCitation id="0AF22A48FFF0DA63FC6CF9CEFBE2F9F4" box="[971,1044,1592,1618]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="5" pageNumber="36">Fig. 8</figureCitation>
|
||
); dorsal aspect of vesicle with a slight basal expansion on each side, under which a smooth longitudinal furrow is present.
|
||
<emphasis id="A0BDEADFFFF0DA63FB68F996FAACF9DF" box="[1231,1370,1632,1657]" italics="true" pageId="5" pageNumber="36">Chelicerae:</emphasis>
|
||
With abundant setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family
|
||
<taxonomicName id="55C94D4EFFF0DA63FA80F97EFEBFF96C" authority="Vachon 1963" authorityName="Vachon" authorityYear="1963" class="Arachnida" family="Chactidae" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="36" phylum="Arthropoda" rank="family">
|
||
Chactidae (
|
||
<bibRefCitation id="F6584B3CFFF0DA63FF39F946FEB6F96C" author="Vachon" box="[158,320,1712,1738]" pageId="5" pageNumber="44" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle (Paris) (2 e ser.), 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Movable finger external surface with one small basal tooth, one sub-basal pronounced tooth, one (right chelicera) or two (left chelicera) small subdistal teeth, and one distal tooth; internal surface of movable finger without teeth and with well developed serrula. Fixed finger external surface with one basal and one median tooth mounted onto a bicuspid, one subdistal pronounced tooth, and one distal tooth; internal surface of fixed finger without teeth.
|
||
<emphasis id="A0BDEADFFFF0DA63FDA3F8A6FD75F8CF" box="[516,643,1872,1897]" italics="true" pageId="5" pageNumber="36">Pedipalps:</emphasis>
|
||
Femur with four granulose carinae (dorsoexternal, dorsointernal, internal median and ventrointernal; ventroexternal carinae absent); patella with four smooth, slightly elevated carinae (ventrolateral, ventrointernal, dorsal and median dorsal carinae; others absent); chela acarinate. Femur with abundant large granules dorsally (
|
||
<figureCitation id="0AF22A48FFF0DA63FDC1F83EFD4BF844" box="[614,701,1992,2018]" captionStart="FIGURES 11 – 18" captionStartId="9.[151,269,1877,1901]" captionTargetBox="[177,1431,214,1817]" captionTargetId="figure@9.[151,1436,194,1853]" captionTargetPageId="9" captionText="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183471/files/figure.png" pageId="5" pageNumber="36">Fig. 15</figureCitation>
|
||
), with thin granulation internally, smooth ventrally and externally; patella smooth in all surfaces, with an internal well developed tubercle; chela smooth; fixed and movable fingers short and thick (
|
||
<figureCitation id="0AF22A48FFF3DA60FE54FF36FD75FF7C" box="[499,643,192,218]" captionStart="FIGURES 11 – 18" captionStartId="9.[151,269,1877,1901]" captionTargetBox="[177,1431,214,1817]" captionTargetId="figure@9.[151,1436,194,1853]" captionTargetPageId="9" captionText="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183471/files/figure.png" pageId="6" pageNumber="37">Figs. 11–14</figureCitation>
|
||
), each with six rows of granules flanked by one external and one internal larger granule. Trichobothriotaxy (
|
||
<figureCitation id="0AF22A48FFF3DA60FD36FF1EFCD7FEA4" box="[657,801,232,258]" captionStart="FIGURES 11 – 18" captionStartId="9.[151,269,1877,1901]" captionTargetBox="[177,1431,214,1817]" captionTargetId="figure@9.[151,1436,194,1853]" captionTargetPageId="9" captionText="FIGURES 11 – 18. Vachoniochactas humboldti sp. nov., distribution of the trichobothria in male holotype (IAvH-E 100799) 11. Chela, dorsal view. 12. Chela, external view. 13. Chela, ventral view. 14. Chela, internal view. 15. Femur, dorsointernal view. 16. Patella, dorsointernal view. 17. Patella, ventral view. 18. Patella, external view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183471/files/figure.png" pageId="6" pageNumber="37">Figs. 11–18</figureCitation>
|
||
):
|
||
<typeStatus id="4D72886FFFF3DA60FC90FF1EFC87FEA4" box="[823,881,232,258]" pageId="6" pageNumber="37">Type</typeStatus>
|
||
C (
|
||
<bibRefCitation id="F6584B3CFFF3DA60FC3AFF1EFBCDFEA4" author="Vachon" box="[925,1083,232,258]" pageId="6" pageNumber="44" refString="Vachon, M. (1973) Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national de Histoire naturelle (Paris) (3 e ser.), 104, 857 - 958." type="journal article" year="1973">Vachon 1973</bibRefCitation>
|
||
) with 54 trichobothria (major neobothriotaxy); femur with three trichobothria; patella with 25 of which 17 are external; chela with 26.
|
||
<emphasis id="A0BDEADFFFF3DA60FAFCFEE6FA6AFE8F" box="[1371,1436,272,297]" italics="true" pageId="6" pageNumber="37">Legs:</emphasis>
|
||
Telotarsi III–IV with a ventromedian row of setae flanked by three alternated pairs of submedian setae plus a single prolateral distal seta (
|
||
<figureCitation id="0AF22A48FFF3DA60FE47FE96FD97FEDC" box="[480,609,352,378]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="6" pageNumber="37">Figs. 9–10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62FE71FAC0FE6C" blockId="6.[151,1437,152,1778]" pageId="6" pageNumber="37">
|
||
<emphasis id="A0BDEADFFFF3DA60FF62FE71FE39FE07" bold="true" box="[197,463,391,417]" pageId="6" pageNumber="37">Hemispermatophore.</emphasis>
|
||
Lamelliform (
|
||
<figureCitation id="0AF22A48FFF3DA60FDDCFE7EFCFDFE04" box="[635,779,392,418]" captionStart="FIGURES 19 – 21" captionStartId="10.[151,269,1306,1330]" captionTargetBox="[177,1421,234,1257]" captionTargetId="figure@10.[151,1436,194,1282]" captionTargetPageId="10" captionText="FIGURES 19 – 21. Vachoniochactas humboldti sp. nov., left hemispermatophore of a male paratype (IAvH-E 100808). 19. External view. 20. Rear view. 21. Internal view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183472/files/figure.png" pageId="6" pageNumber="37">Figs. 19–21</figureCitation>
|
||
). Lamina straight and slender, with no distal ridge; capsular lobes small and simple; internal lobe with two to four sclerotized spur-like external projections.
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62FE2EFC6BFE54" blockId="6.[151,1437,152,1778]" box="[197,925,472,498]" pageId="6" pageNumber="37">Female allotype (ICN-MHN-As-651) and sexual dimorphism.</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62FDF6FC46FDBC" blockId="6.[151,1437,152,1778]" box="[197,944,512,538]" pageId="6" pageNumber="37">
|
||
Total body length 24.31 mm; detailed measurements in
|
||
<tableCitation id="DF4B0376FFF3DA60FCF5FDF6FC5CFDBC" box="[850,938,512,538]" captionStart="TABLE 1" captionStartId="4.[151,239,271,295]" captionTargetBox="[159,1411,405,1700]" captionTargetPageId="4" captionText="TABLE 1. Vachoniochactas humboldti sp. nov., measurements (in mm). L = length, W = width, D = depth." httpUri="http://table.plazi.org/id/C6B66645FFF1DA62FF30FEF9FB1CFE81" pageId="6" pageNumber="37" tableUuid="C6B66645FFF1DA62FF30FEF9FB1CFE81">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62FDDEFBC7FC24" blockId="6.[151,1437,152,1778]" pageId="6" pageNumber="37">
|
||
Coloration and morphology similar to the male
|
||
<typeStatus id="4D72886FFFF3DA60FCA7FDDEFC9CFDE4" box="[768,874,552,578]" pageId="6" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
, except for that the sternites are lighter and the pectines have 7:8 teeth. The following differences were also identified and, through the examination of the available sample, are attributed to sexual dimorphism: (1) the tergites, sternites and medially the carapace are almost completely smooth and shiny, with very weak granulation (
|
||
<figureCitation id="0AF22A48FFF3DA60FC13FD56FBDEFD1C" box="[948,1064,672,698]" captionStart="FIGURES 2 – 5" captionStartId="7.[151,269,1808,1832]" captionTargetBox="[207,1379,195,1783]" captionTargetId="figure@7.[207,1379,194,1784]" captionTargetPageId="7" captionText="FIGURES 2 – 5. Vachoniochactas humboldti sp. nov. 2 – 3. Male holotype (IAvH-E 100799), dorsal and ventral views. 4 – 5. Female allotype (ICN-MHN-As- 651), dorsal and ventral views. Scale bars = 5 mm." httpUri="https://zenodo.org/record/183469/files/figure.png" pageId="6" pageNumber="37">Figs. 4–5</figureCitation>
|
||
); (2) the pectines are short and with smaller and more slender teeth (
|
||
<figureCitation id="0AF22A48FFF3DA60FDE9FD3EFD60FD44" box="[590,662,712,738]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="6" pageNumber="37">Fig. 7</figureCitation>
|
||
); (3) the sternum is almost as long as wide in the female (
|
||
<figureCitation id="0AF22A48FFF3DA60FAE3FD3EFA7AFD44" box="[1348,1420,712,738]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="6" pageNumber="37">Fig. 7</figureCitation>
|
||
); and (4) the pectinal basal piece is straight anteriorly lacking median notch (
|
||
<figureCitation id="0AF22A48FFF3DA60FBA9FD06FBA0FCAC" box="[1038,1110,752,778]" captionStart="FIGURES 6 – 10" captionStartId="8.[151,269,1759,1783]" captionTargetBox="[152,1399,222,1709]" captionTargetId="figure@8.[151,1436,194,1735]" captionTargetPageId="8" captionText="FIGURES 6 – 10. Vachoniochactas humboldti sp. nov. 6 – 7. Coxosternal region, genital operculum, and pectines. 6. Male holotype (IAvH-E 100799). 7. Female allotype (ICN-MHN-As- 651). 8. Metasomal segments IV – V and telson of male holotype, lateral view. 9 – 10. Telotarsus IV of male holotype. 9. Prolateral view. 10. Ventral view. Scale bars = 1 mm (Figs. 6 – 7), 2 mm (Fig. 8), 0.5 mm (Figs. 9 – 10)." httpUri="https://zenodo.org/record/183470/files/figure.png" pageId="6" pageNumber="37">Fig. 7</figureCitation>
|
||
). Sexual dimorphism in the total body size, with males being smaller than females, is also suggested by the available materials (no male reaches the size of the female allotype, 24.31 mm). These differences may occur too in the other species of the genus but this still remains to be proven (in most of them males are unknown).
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62FC79FE4AFAC4" blockId="6.[151,1437,152,1778]" pageId="6" pageNumber="37">
|
||
<emphasis id="A0BDEADFFFF3DA60FF62FC79FEA7FC0F" bold="true" box="[197,337,911,937]" pageId="6" pageNumber="37">Variability.</emphasis>
|
||
<emphasis id="A0BDEADFFFF3DA60FEFBFC66FE2CFC0F" box="[348,474,912,937]" italics="true" pageId="6" pageNumber="37">Body size:</emphasis>
|
||
Total body length 19.30–24.02 mm in adult males (n = 13; mean 22.56 mm), 23.15–24.31 mm in females (n = 3; mean 23.67 mm).
|
||
<emphasis id="A0BDEADFFFF3DA60FCAAFC4EFBF7FC77" box="[781,1025,952,977]" italics="true" pageId="6" pageNumber="37">Pectinal teeth count:</emphasis>
|
||
<quantity id="55319B28FFF3DA60FBA0FC4EFB91FC74" box="[1031,1127,952,978]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="6" pageNumber="37" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7 to 9 in</quantity>
|
||
males (mode 8:8; n = 16);
|
||
<quantity id="55319B28FFF3DA60FF30FC16FF0CFC5C" box="[151,250,992,1018]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="6" pageNumber="37" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7 to 8 in</quantity>
|
||
females (mode 8:8; n = 3).
|
||
<emphasis id="A0BDEADFFFF3DA60FDE3FC16FC0DFC5F" box="[580,1019,992,1017]" italics="true" pageId="6" pageNumber="37">External trichobothria of the patella:</emphasis>
|
||
In most specimens the external trichobothria of the patella are arranged as in figure 18; however, two males (CDA 0 0 0.805, IAvH-E
|
||
<date id="E677100DFFF3DA60FA9DFBFEFA62FB84" box="[1338,1428,1032,1058]" pageId="6" pageNumber="37" value="95-10-07">100795</date>
|
||
) exhibit a different position of the
|
||
<emphasis id="A0BDEADFFFF3DA60FDBAFBC6FDB2FBEF" box="[541,580,1072,1097]" italics="true" pageId="6" pageNumber="37">esb</emphasis>
|
||
trichobothria, being the resulting line connecting these trichobothria parallel to the
|
||
<emphasis id="A0BDEADFFFF3DA60FE8CFBAEFEBBFBD7" box="[299,333,1112,1137]" italics="true" pageId="6" pageNumber="37">em</emphasis>
|
||
group.
|
||
<emphasis id="A0BDEADFFFF3DA60FE14FBAEFD5AFBD7" box="[435,684,1112,1137]" italics="true" pageId="6" pageNumber="37">Hemispermatophore:</emphasis>
|
||
Total length 3.05–4.25 mm (n = 14; mean 3.78 mm). Both hemispermatophores were extracted from seven adult males. In two specimens the number of spur-like projections in the capsule (expressed for left/right hemispermatophores respectively) is 4/3 (IAvH-E
|
||
<date id="E677100DFFF3DA60FA9AFB5EFA6EFB64" box="[1341,1432,1192,1218]" pageId="6" pageNumber="37" value="06-10-08">100806</date>
|
||
,
|
||
<date id="E677100DFFF3DA60FF30FB26FF05FB4C" box="[151,243,1232,1258]" pageId="6" pageNumber="37" value="08-10-08">100808</date>
|
||
), in one it is 3/4 (IAvH-E
|
||
<date id="E677100DFFF3DA60FD8FFB26FD73FB4C" box="[552,645,1232,1258]" pageId="6" pageNumber="37" value="04-10-08">100804</date>
|
||
), in one it is 2/3 (CDA 000.805), in one it is 3/2 (IAvH-E
|
||
<date id="E677100DFFF3DA60FA94FB26FA79FB4C" box="[1331,1423,1232,1258]" pageId="6" pageNumber="37" value="07-10-08">100807</date>
|
||
), in one it is 2/2 (IAvH-E
|
||
<date id="E677100DFFF3DA60FE17FB0EFDFAFAB4" box="[432,524,1272,1298]" pageId="6" pageNumber="37" value="05-10-08">100805</date>
|
||
), and in one it is 3/3 (IAvH-E
|
||
<date id="E677100DFFF3DA60FCCDFB0EFC31FAB4" box="[874,967,1272,1298]" pageId="6" pageNumber="37" value="03-10-08">100803</date>
|
||
). This indicates that the number of spurlike projections in the capsule is variable not only between individuals but also between hemispermatophores from the same specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAD36546FFF3DA60FF62FA99FBBEF954" pageId="6" pageNumber="37" type="distribution">
|
||
<paragraph id="927636CDFFF3DA60FF62FA99FC07F9DC" blockId="6.[151,1437,152,1778]" pageId="6" pageNumber="37">
|
||
<emphasis id="A0BDEADFFFF3DA60FF62FA99FE09FA2F" bold="true" box="[197,511,1391,1417]" pageId="6" pageNumber="37">Distribution and habitat.</emphasis>
|
||
According to the biogeographic units proposed by
|
||
<bibRefCitation id="F6584B3CFFF3DA60FBC6FA86FF12FA14" pageId="6" pageNumber="43" refString="Hernandez-Camacho, J., Hurtado, A., Ortiz, R. & Walschburger, T. (1992) Unidades biogeograficas de Colombia. In: Halffter, G. (Ed.). La diversidad biologica de Iberoamerica I. Volumen especial, Acta Zoologica Mexicana. Programa Iberoamericano de Ciencia y Tecnologia para el Desarrollo-Instituto de Ecologia, A. C. Xalapa, Mexico, pp. 103 - 151." type="book chapter">
|
||
Hernández-Camacho
|
||
<emphasis id="A0BDEADFFFF3DA60FAC7FA86FA63FA2F" box="[1376,1429,1392,1417]" italics="true" pageId="6" pageNumber="37">et al</emphasis>
|
||
. (1992)
|
||
</bibRefCitation>
|
||
the area where
|
||
<taxonomicName id="55C94D4EFFF3DA60FE06FA6EFDC3FA17" box="[417,565,1432,1457]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="humboldti" status="sp. nov.">
|
||
<emphasis id="A0BDEADFFFF3DA60FE06FA6EFDC3FA17" box="[417,565,1432,1457]" italics="true" pageId="6" pageNumber="37">V. humboldti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BB8E57A4FFF3DA60FD9AFA6EFD6FFA14" box="[573,665,1432,1458]" pageId="6" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected is enclaved in the Vaupes complex of the Guayana Biogeographic Province, adjacent to the Amazonian region. This region comprises swampy forests and noninundable terraces surrounded by rivers where the gallery forests are formed. The northernmost known locality of the new species is inhabited by the Santa Rosa Indigenous Community, and includes mature forests with non-inundable terrace growing in rocky hills and sandy floors. It is somewhat disturbed due to small cultivations and animal hunting by the indigenous community for their survival.
|
||
</paragraph>
|
||
<paragraph id="927636CDFFF3DA60FF62F97EFBBEF954" blockId="6.[151,1437,152,1778]" pageId="6" pageNumber="37">
|
||
The fact that most specimens were collected by pitfall and Winkler traps suggests that this species is a leaf litter dweller, a microhabitat where the specimen of
|
||
<taxonomicName id="55C94D4EFFF3DA60FD5DF946FC37F96F" box="[762,961,1712,1737]" class="Arachnida" family="Chactidae" genus="Vachoniochactas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0BDEADFFFF3DA60FD5DF946FC37F96F" box="[762,961,1712,1737]" italics="true" pageId="6" pageNumber="37">Vachoniochactas</emphasis>
|
||
</taxonomicName>
|
||
sp. from
|
||
<collectingCountry id="EADE765DFFF3DA60FB96F946FB5FF96C" box="[1073,1193,1712,1738]" name="Venezuela" pageId="6" pageNumber="37">Venezuela</collectingCountry>
|
||
and deposited in the California Academy of Science was also captured (M.E. Soleglad, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |