treatments-xml/data/00/AC/B4/00ACB419898B5ED9B6932DA84F528885.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.47088" ID-GBIF-Dataset="e62a9840-3a2f-4f8b-9572-85485be2b102" ID-GBIF-Taxon="164486286" ID-Pensoft-Pub="1860-0743-1-263" ID-Pensoft-UUID="60B681D82CA852FA996FB0787B79567C" ID-ZooBank="7F7AC9881EF840148DDC0BE0ABA09B4F" LSID="urn:lsid:zoobank.org:act:80E35235-9E39-48EC-ACB0-525952C098BC" ModsDocID="1860-0743-1-263" checkinTime="1591462957541" checkinUser="pensoft" docAuthor="Rakotoarison, Andolalao, Scherz, Mark D., Koehler, Joern, Ratsoavina, Fanomezana M., Hawlitschek, Oliver, Megson, Steven, Vences, Miguel &amp; Glaw, Frank" docDate="2020" docId="00ACB419898B5ED9B6932DA84F528885" docLanguage="en" docName="ZoosystEvol 96(1): 263-274" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.47088" docTitle="Platypelis laetus Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw 2020, sp. nov." docType="treatment" docUuid="80E35235-9E39-48EC-ACB0-525952C098BC" docUuidSource="ZooBank" docVersion="3" id="60B681D82CA852FA996FB0787B79567C" lastPageNumber="263" masterDocId="60B681D82CA852FA996FB0787B79567C" masterDocTitle="Frogs of the genus Platypelis from the Sorata massif in northern Madagascar: description of a new species and reports of range extensions" masterLastPageNumber="274" masterPageNumber="263" pageId="0" pageNumber="263" updateTime="1643444340996" updateUser="ExternalLinkService">
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<mods:title>Frogs of the genus Platypelis from the Sorata massif in northern Madagascar: description of a new species and reports of range extensions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakotoarison, Andolalao</mods:namePart>
<mods:affiliation>Zoological Institute, Technical University of Braunschweig, Mendelssohnstr, 4, 38106, Braunschweig, Germany &amp; Mention Zoologie et Biodiversite Animale, Faculte des Sciences, Universite d'Antananarivo, BP 906, Madagascar</mods:affiliation>
<mods:nameIdentifier type="email">andomailaka@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scherz, Mark D.</mods:namePart>
<mods:affiliation>Zoological Institute, Technical University of Braunschweig, Mendelssohnstr, 4, 38106, Braunschweig, Germany &amp; Zoologische Staatssammlung Muenchen (ZSM-SNSB), Muenchhausenstr, 21, 81247 Muenchen, Germany</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4613-7761</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koehler, Joern</mods:namePart>
<mods:affiliation>Hessisches Landesmuseum Darmstadt, Friedensplatz 1, 64283 Darmstadt, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ratsoavina, Fanomezana M.</mods:namePart>
<mods:affiliation>Mention Zoologie et Biodiversite Animale, Faculte des Sciences, Universite d'Antananarivo, BP 906, Madagascar</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hawlitschek, Oliver</mods:namePart>
<mods:affiliation>Zoologische Staatssammlung Muenchen (ZSM-SNSB), Muenchhausenstr, 21, 81247 Muenchen, Germany &amp; Centrum fuer Naturkunde (CeNak), Universitaet Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Megson, Steven</mods:namePart>
<mods:affiliation>School of Science and the Environment, Manchester Metropolitan University, Manchester, M 1 5 GD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vences, Miguel</mods:namePart>
<mods:affiliation>Zoological Institute, Technical University of Braunschweig, Mendelssohnstr, 4, 38106, Braunschweig, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glaw, Frank</mods:namePart>
<mods:affiliation>Zoologische Staatssammlung Muenchen (ZSM-SNSB), Muenchhausenstr, 21, 81247 Muenchen, Germany</mods:affiliation>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>96</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zse.96.47088</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.47088</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-263</mods:identifier>
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<treatment ID-GBIF-Taxon="164486286" LSID="urn:lsid:zoobank.org:act:80E35235-9E39-48EC-ACB0-525952C098BC" httpUri="http://treatment.plazi.org/id/00ACB419898B5ED9B6932DA84F528885" lastPageNumber="263" pageId="0" pageNumber="263">
<subSubSection pageId="0" pageNumber="263" type="nomenclature">
<paragraph pageId="0" pageNumber="263">
<taxonomicName LSID="00ACB419-898B-5ED9-B693-2DA84F528885" authority="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw, 2020" authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" family="Microhylidae" genus="Platypelis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Platypelis laetus" order="Anura" pageId="0" pageNumber="263" rank="species" species="laetus" status="sp. nov.">Platypelis laetus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="263">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="remarks">
<paragraph pageId="0" pageNumber="263">Remark.</paragraph>
<paragraph pageId="0" pageNumber="263">
This species has been listed as 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
sp. CaNEW2 Sorata by 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2016.04.019" author="Scherz, MD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="263" pagination="372 - 381" refId="B28" refString="Scherz, MD, Vences, M, Rakotoarison, A, Andreone, F, Koehler, J, Glaw, F, Crottini, A, 2016. Reconciling molecular phylogeny, morphological divergence and classification of Madagascan narrow-mouthed frogs (Amphibia: Microhylidae). Molecular Phylogenetics and Evolution 100: 372 - 381, DOI: https://doi.org/10.1016/j.ympev.2016.04.019" title="Reconciling molecular phylogeny, morphological divergence and classification of Madagascan narrow-mouthed frogs (Amphibia: Microhylidae)." url="https://doi.org/10.1016/j.ympev.2016.04.019" volume="100" year="2016">Scherz et al. (2016</bibRefCitation>
, 2017) and 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2017.03.016" author="Peloso, PLV" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="263" pagination="56 - 64" refId="B15" refString="Peloso, PLV, Raxworthy, CJ, Wheeler, WC, Frost, DR, 2017. Nomenclatural stability does not justify recognition of paraphyletic taxa: a response to Scherz et al. (2016). Molecular Phylogenetics and Evolution 111: 56 - 64, DOI: https://doi.org/10.1016/j.ympev.2017.03.016" title="Nomenclatural stability does not justify recognition of paraphyletic taxa: a response to Scherz et al. (2016)." url="https://doi.org/10.1016/j.ympev.2017.03.016" volume="111" year="2017">Peloso et al. (2017)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="holotype">
<paragraph pageId="0" pageNumber="263">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="263">
Holotype ZSM 5652/2012 (FGZC 3761) (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs in life of Platypelis laetus sp. nov. from the Sorata massif: a-c ZSM 5652 / 2012 (FGZC 3761), holotype; d, e ZSM 5651 / 2012 (FGZC 3588), paratype; f, g ZSM 5653 / 2012 (FGZC 3762), paratype; h Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652 / 2012 and ZSM 5653 / 2012." figureDoi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263">3a-c</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved holotype of Platypelis laetus sp. nov., ZSM 5652 / 2012 (FGZC 3761), in dorsal and ventral view." figureDoi="10.3897/zse.96.47088.figure4" httpUri="https://binary.pensoft.net/fig/418560" pageId="0" pageNumber="263">4</figureCitation>
), adult male, collected on 30 November 2012, in the Sorata Massif (near 
<geoCoordinate degrees="13.6817" direction="south" orientation="latitude" precision="5" value="-13.6817">13.6817S</geoCoordinate>
, 
<geoCoordinate degrees="49.4411" direction="east" orientation="longitude" precision="5" value="49.4411">49.4411E</geoCoordinate>
, 1339 m a.s.l.), northern Madagascar, by F. Glaw, O. Hawlitschek, T. Rajoafiarison, A. Rakotoarison, F.M. Ratsoavina, and A. Razafimanantsoa.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="paratypes">
<paragraph pageId="0" pageNumber="263">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="263">
ZSM 5651/2012 (FGZC 3588) (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs in life of Platypelis laetus sp. nov. from the Sorata massif: a-c ZSM 5652 / 2012 (FGZC 3761), holotype; d, e ZSM 5651 / 2012 (FGZC 3588), paratype; f, g ZSM 5653 / 2012 (FGZC 3762), paratype; h Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652 / 2012 and ZSM 5653 / 2012." figureDoi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263">3d, e</figureCitation>
) collected on 26 November 2012 from the high elevation of the Sorata Massif (near 
<geoCoordinate degrees="13.6745" direction="south" orientation="latitude" precision="5" value="-13.6745">13.6745S</geoCoordinate>
, 
<geoCoordinate degrees="49.4402" direction="east" orientation="longitude" precision="5" value="49.4402">49.4402E</geoCoordinate>
, 1541 m a.s.l.), northern Madagascar, by the same collectors as the holotype; ZSM 5653/2012 (FGZC 3762) (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs in life of Platypelis laetus sp. nov. from the Sorata massif: a-c ZSM 5652 / 2012 (FGZC 3761), holotype; d, e ZSM 5651 / 2012 (FGZC 3588), paratype; f, g ZSM 5653 / 2012 (FGZC 3762), paratype; h Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652 / 2012 and ZSM 5653 / 2012." figureDoi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263">3f, g</figureCitation>
), adult male, collected on 30 November 2012, from the same site and by the same collectors as the holotype; UADBA-A (FGZC 3767) and UADBA-A (FGZC 3768), both ovigerous adult females, collected on 30 November 2012 from the same site and by the same collectors as the holotype.
</paragraph>
<caption doi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="263">
<emphasis bold="true" pageId="0" pageNumber="263">Figure 3.</emphasis>
Photographs in life of 
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov. from the Sorata massif: 
<emphasis bold="true" pageId="0" pageNumber="263">a-c</emphasis>
ZSM 5652/2012 (FGZC 3761), holotype; 
<emphasis bold="true" pageId="0" pageNumber="263">d, e</emphasis>
ZSM 5651/2012 (FGZC 3588), paratype; 
<emphasis bold="true" pageId="0" pageNumber="263">f, g</emphasis>
ZSM 5653/2012 (FGZC 3762), paratype; 
<emphasis bold="true" pageId="0" pageNumber="263">h</emphasis>
Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652/2012 and ZSM 5653/2012.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="diagnosis">
<paragraph pageId="0" pageNumber="263">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="263">
Assigned to the genus 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
in the microhylid subfamily 
<taxonomicName authorityName="Cope" authorityYear="1889" lsidName="" pageId="0" pageNumber="263" rank="subFamily" subFamily="Cophylinae">Cophylinae</taxonomicName>
based on enlarged terminal discs on fingers and toes, absence of nuptial pads, and molecular phylogenetic relationships. The species can be identified among other cophylines by the combination of the following character states: (1) medium-sized species (adult male SVL 24.3-25.6 mm); (2) manus with second finger slightly shorter than fourth and pes with third toe much shorter to very slightly shorter than fifth; (3) males with prepollical tubercle but lacking a finger-like prepollex as typical for 
<taxonomicName authorityName="Muller" authorityYear="1892" class="Amphibia" family="Microhylidae" genus="Anodonthyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anodonthyla" order="Anura" pageId="0" pageNumber="263" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Anodonthyla</emphasis>
</taxonomicName>
; (4) throat greenish in life; (5) chest and anterior belly translucent gray, with distinct white spotting that is absent on the posterior belly; (5) absence of red color on limbs and ventral side.
</paragraph>
<caption doi="10.3897/zse.96.47088.figure4" httpUri="https://binary.pensoft.net/fig/418560" pageId="0" pageNumber="263" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="263">
<emphasis bold="true" pageId="0" pageNumber="263">Figure 4.</emphasis>
Preserved holotype of 
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov., ZSM 5652/2012 (FGZC 3761), in dorsal and ventral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="263">
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov. can be distinguished from 
<taxonomicName genus="P." lsidName="P. grandis" pageId="0" pageNumber="263" rank="species" species="grandis">
<emphasis italics="true" pageId="0" pageNumber="263">P. grandis</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. alticola" pageId="0" pageNumber="263" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="263">P. alticola</emphasis>
</taxonomicName>
by smaller body size (verified adult male SVL 24.3-25.6 mm vs 30-105 mm), and furthermore from 
<taxonomicName genus="P." lsidName="P. grandis" pageId="0" pageNumber="263" rank="species" species="grandis">
<emphasis italics="true" pageId="0" pageNumber="263">P. grandis</emphasis>
</taxonomicName>
by a largely smooth dorsal skin (vs strongly granular and bumpy) and from 
<taxonomicName genus="P." lsidName="P. tuberifera" pageId="0" pageNumber="263" rank="species" species="tuberifera">
<emphasis italics="true" pageId="0" pageNumber="263">P. tuberifera</emphasis>
</taxonomicName>
by dark pattern on the dorsum (vs uniformly beige-yellowish) and body not conspicuously flattened (vs flattened); from 
<taxonomicName genus="P." lsidName="P. pollicaris" pageId="0" pageNumber="263" rank="species" species="pollicaris">
<emphasis italics="true" pageId="0" pageNumber="263">P. pollicaris</emphasis>
</taxonomicName>
by a slightly smaller body size (verified adult male SVL 24.3-25.6 mm vs 26-28 mm), third toe shorter than fifth (vs both toes of similar length) and less elongated body (vs conspicuously elongated); from 
<taxonomicName genus="P." lsidName="P. tuberifera" pageId="0" pageNumber="263" rank="species" species="tuberifera">
<emphasis italics="true" pageId="0" pageNumber="263">P. tuberifera</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. cowanii" pageId="0" pageNumber="263" rank="species" species="cowanii">
<emphasis italics="true" pageId="0" pageNumber="263">P. cowanii</emphasis>
</taxonomicName>
by smaller body size (verified adult male SVL 24.3-25.6 mm vs 30-40 mm) and by third toe shorter than fifth (vs third toe longer than fifth); from 
<taxonomicName genus="P." lsidName="P. tsaratananaensis" pageId="0" pageNumber="263" rank="species" species="tsaratananaensis">
<emphasis italics="true" pageId="0" pageNumber="263">P. tsaratananaensis</emphasis>
</taxonomicName>
by having a plump body (vs slender with a distinctively elongated 
<normalizedToken originalValue="“neck”">&quot;neck&quot;</normalizedToken>
region), absence of vomerine teeth (vs presence), and especially in call structure (see below); from 
<taxonomicName genus="P." lsidName="P. tetra" pageId="0" pageNumber="263" rank="species" species="tetra">
<emphasis italics="true" pageId="0" pageNumber="263">P. tetra</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. barbouri" pageId="0" pageNumber="263" rank="species" species="barbouri">
<emphasis italics="true" pageId="0" pageNumber="263">P. barbouri</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. karenae" pageId="0" pageNumber="263" rank="species" species="karenae">
<emphasis italics="true" pageId="0" pageNumber="263">P. karenae</emphasis>
</taxonomicName>
, 
<taxonomicName genus="P." lsidName="P. ravus" pageId="0" pageNumber="263" rank="species" species="ravus">
<emphasis italics="true" pageId="0" pageNumber="263">P. ravus</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="P." lsidName="P. ando" pageId="0" pageNumber="263" rank="species" species="ando">
<emphasis italics="true" pageId="0" pageNumber="263">P. ando</emphasis>
</taxonomicName>
by larger body size (SVL 24.3-25.6 mm vs 16-19 mm); from 
<taxonomicName genus="P." lsidName="P. milloti" pageId="0" pageNumber="263" rank="species" species="milloti">
<emphasis italics="true" pageId="0" pageNumber="263">P. milloti</emphasis>
</taxonomicName>
by throat greenish colored (vs brownish) and by absence of red ventral color and of a very distinct dorsal pattern (vs presence); from 
<taxonomicName genus="P." lsidName="P. barbouri" pageId="0" pageNumber="263" rank="species" species="barbouri">
<emphasis italics="true" pageId="0" pageNumber="263">P. barbouri</emphasis>
</taxonomicName>
and 
<taxonomicName genus="P." lsidName="P. ranjomena" pageId="0" pageNumber="263" rank="species" species="ranjomena">
<emphasis italics="true" pageId="0" pageNumber="263">P. ranjomena</emphasis>
</taxonomicName>
by the absence of red color on ventral side or limbs (vs presence); from 
<taxonomicName genus="P." lsidName="P. mavomavo" pageId="0" pageNumber="263" rank="species" species="mavomavo">
<emphasis italics="true" pageId="0" pageNumber="263">P. mavomavo</emphasis>
</taxonomicName>
by third toe shorter than fifth (vs third toe longer than fifth) and grayish with white spots on the chest (vs uniformly yellow on venter). Among the described species of 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
, the new species is phylogenetically closest to 
<taxonomicName genus="P." lsidName="P. olgae" pageId="0" pageNumber="263" rank="species" species="olgae">
<emphasis italics="true" pageId="0" pageNumber="263">P. olgae</emphasis>
</taxonomicName>
. It differs from this species by larger body size (adult male SVL 24.3-25.6 mm vs 20.3-21.9 mm), and by third toe often shorter than fifth (vs third toe equal to or longer than fifth).
</paragraph>
<paragraph pageId="0" pageNumber="263">
Bioacoustically, 
<taxonomicName genus="P." lsidName="P. laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">P. laetus</emphasis>
</taxonomicName>
sp. nov. can be distinguished from most congeners by its rather short note duration of only 73-88 ms. The following 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
species all emit longer notes: 
<taxonomicName genus="P." lsidName="P. alticola" pageId="0" pageNumber="263" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="263">P. alticola</emphasis>
</taxonomicName>
(411-466 ms), 
<taxonomicName genus="P." lsidName="P. ando" pageId="0" pageNumber="263" rank="species" species="ando">
<emphasis italics="true" pageId="0" pageNumber="263">P. ando</emphasis>
</taxonomicName>
(424-441 ms), 
<taxonomicName genus="P." lsidName="P. barbouri" pageId="0" pageNumber="263" rank="species" species="barbouri">
<emphasis italics="true" pageId="0" pageNumber="263">P. barbouri</emphasis>
</taxonomicName>
(142-160 ms), 
<taxonomicName genus="P." lsidName="P. karenae" pageId="0" pageNumber="263" rank="species" species="karenae">
<emphasis italics="true" pageId="0" pageNumber="263">P. karenae</emphasis>
</taxonomicName>
(131-145 ms), 
<taxonomicName genus="P." lsidName="P. pollicaris" pageId="0" pageNumber="263" rank="species" species="pollicaris">
<emphasis italics="true" pageId="0" pageNumber="263">P. pollicaris</emphasis>
</taxonomicName>
(160-180 ms), 
<taxonomicName genus="P." lsidName="P. ranjomena" pageId="0" pageNumber="263" rank="species" species="ranjomena">
<emphasis italics="true" pageId="0" pageNumber="263">P. ranjomena</emphasis>
</taxonomicName>
(303-379 ms), 
<taxonomicName genus="P." lsidName="P. ravus" pageId="0" pageNumber="263" rank="species" species="ravus">
<emphasis italics="true" pageId="0" pageNumber="263">P. ravus</emphasis>
</taxonomicName>
(384-443 ms), and 
<taxonomicName genus="P." lsidName="P. tuberifera" pageId="0" pageNumber="263" rank="species" species="tuberifera">
<emphasis italics="true" pageId="0" pageNumber="263">P. tuberifera</emphasis>
</taxonomicName>
(280 ms) (
<bibRefCitation author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlag, Koeln" pageId="0" pageNumber="263" refId="B7" refString="Glaw, F, Vences, M, 1994. A Fieldguide to the Amphibians and Reptiles of Madagascar, 2nd edition. Vences &amp; Glaw Verlag, Koeln" title="A Fieldguide to the Amphibians and Reptiles of Madagascar, 2 nd edition." year="1994">Glaw and Vences 1994</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2012.9" author="Glaw, F" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="263" pagination="1 - 9" refId="B6" refString="Glaw, F, Koehler, J, Vences, M, 2012. A tiny new species of Platypelis from the Marojejy National Park in northeastern Madagascar (Amphibia: Microhylidae). European Journal of Taxonomy 9: 1 - 9, DOI: https://doi.org/10.5852/ejt.2012.9" title="A tiny new species of Platypelis from the Marojejy National Park in northeastern Madagascar (Amphibia: Microhylidae)." url="https://doi.org/10.5852/ejt.2012.9" volume="9" year="2012">Glaw et al. 2012</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.26049/vz70-2-2020-04" author="Glaw, F" journalOrPublisher="Vertebrate Zoology" pageId="0" pageNumber="263" pagination="141 - 156" refId="B9" refString="Glaw, F, Scherz, MD, Rakotoarison, A, Crottini, A, Raselimanana, A, Andreone, F, Koehler, J, Vences, M, 2020. Genetic variability and partial integrative revision of Platypelis frogs (Microhylidae) with red flash marks from eastern Madagascar. Vertebrate Zoology 70: 141 - 156, DOI: https://doi.org/10.26049/vz70-2-2020-04" title="Genetic variability and partial integrative revision of Platypelis frogs (Microhylidae) with red flash marks from eastern Madagascar." url="https://doi.org/10.26049/vz70-2-2020-04" volume="70" year="2020">2020</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa3563.1.1" author="Rakotoarison, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="263" pagination="1 - 25" refId="B19" refString="Rakotoarison, A, Glaw, F, Vieites, DR, Raminosoa, RN, Vences, M, 2012. Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species. Zootaxa 3563: 1 - 25, DOI: https://doi.org/10.11646/zootaxa3563.1.1" title="Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species." url="https://doi.org/10.11646/zootaxa3563.1.1" volume="3563" year="2012">Rakotoarison et al. 2012</bibRefCitation>
; 
<bibRefCitation author="Rosa, GM" journalOrPublisher="Salamandra" pageId="0" pageNumber="263" pagination="201 - 214" refId="B25" refString="Rosa, GM, Crottini, A, Noel, J, Rabibisoa, N, Raxworthy, CJ, Andreone, F, 2014. A new phytotelmic species of Platypelis (Microhylidae: Cophylinae) from the Betampona Reserve, eastern Madagascar. Salamandra 50: 201 - 214" title="A new phytotelmic species of Platypelis (Microhylidae: Cophylinae) from the Betampona Reserve, eastern Madagascar." volume="50" year="2014">Rosa et al. 2014</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.3897/evolsyst.3.33417" author="Scherz, MD" journalOrPublisher="Evolutionary Systematics" pageId="0" pageNumber="263" pagination="75 - 83" refId="B33" refString="Scherz, MD, Koehler, J, Vences, M, Glaw, F, 2019. A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar. Evolutionary Systematics 3: 75 - 83, DOI: https://doi.org/10.3897/evolsyst.3.33417" title="A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar." url="https://doi.org/10.3897/evolsyst.3.33417" volume="3" year="2019">Scherz et al. 2019</bibRefCitation>
). Calls of 
<taxonomicName genus="P." lsidName="P. grandis" pageId="0" pageNumber="263" rank="species" species="grandis">
<emphasis italics="true" pageId="0" pageNumber="263">P. grandis</emphasis>
</taxonomicName>
are non-tonal, those of 
<taxonomicName genus="P." lsidName="P. milloti" pageId="0" pageNumber="263" rank="species" species="milloti">
<emphasis italics="true" pageId="0" pageNumber="263">P. milloti</emphasis>
</taxonomicName>
are shorter in duration (55-65 ms) and the call of 
<taxonomicName genus="P." lsidName="P. tsaratananaensis" pageId="0" pageNumber="263" rank="species" species="tsaratananaensis">
<emphasis italics="true" pageId="0" pageNumber="263">P. tsaratananaensis</emphasis>
</taxonomicName>
consist of two notes, rather than a single note (
<bibRefCitation author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlag, Koeln" pageId="0" pageNumber="263" refId="B7" refString="Glaw, F, Vences, M, 1994. A Fieldguide to the Amphibians and Reptiles of Madagascar, 2nd edition. Vences &amp; Glaw Verlag, Koeln" title="A Fieldguide to the Amphibians and Reptiles of Madagascar, 2 nd edition." year="1994">Glaw and Vences 1994</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa3563.1.1" author="Rakotoarison, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="263" pagination="1 - 25" refId="B19" refString="Rakotoarison, A, Glaw, F, Vieites, DR, Raminosoa, RN, Vences, M, 2012. Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species. Zootaxa 3563: 1 - 25, DOI: https://doi.org/10.11646/zootaxa3563.1.1" title="Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species." url="https://doi.org/10.11646/zootaxa3563.1.1" volume="3563" year="2012">Rakotoarison et al. 2012</bibRefCitation>
). Known calls of 
<taxonomicName authorityName="Boettger" authorityYear="1880" class="Amphibia" family="Microhylidae" genus="Cophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cophyla" order="Anura" pageId="0" pageNumber="263" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Cophyla</emphasis>
</taxonomicName>
species (
<taxonomicName genus="C." lsidName="C. berara" pageId="0" pageNumber="263" rank="species" species="berara">
<emphasis italics="true" pageId="0" pageNumber="263">C. berara</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. fortuna" pageId="0" pageNumber="263" rank="species" species="fortuna">
<emphasis italics="true" pageId="0" pageNumber="263">C. fortuna</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. maharipeo" pageId="0" pageNumber="263" rank="species" species="maharipeo">
<emphasis italics="true" pageId="0" pageNumber="263">C. maharipeo</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. noromalalae" pageId="0" pageNumber="263" rank="species" species="noromalalae">
<emphasis italics="true" pageId="0" pageNumber="263">C. noromalalae</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. occultans" pageId="0" pageNumber="263" rank="species" species="occultans">
<emphasis italics="true" pageId="0" pageNumber="263">C. occultans</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. phyllodactyla" pageId="0" pageNumber="263" rank="species" species="phyllodactyla">
<emphasis italics="true" pageId="0" pageNumber="263">C. phyllodactyla</emphasis>
</taxonomicName>
, 
<taxonomicName genus="C." lsidName="C. puellarum" pageId="0" pageNumber="263" rank="species" species="puellarum">
<emphasis italics="true" pageId="0" pageNumber="263">C. puellarum</emphasis>
</taxonomicName>
) all exhibit a much longer note duration compared to the new species, ranging from 326 ms in the shortest to 1346 ms in the longest notes (see 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3937.1.3" author="Rakotoarison, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="263" pagination="61 - 89" refId="B20" refString="Rakotoarison, A, Crottini, A, Mueller, J, Roedel, MO, Glaw, F, Vences, M, 2015. Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species. Zootaxa 3937: 61 - 89, DOI: https://doi.org/10.11646/zootaxa.3937.1.3" title="Revision and phylogeny of narrow-mouthed treefrogs (Cophyla) from northern Madagascar: integration of molecular, osteological, and bioacoustic data reveals three new species." url="https://doi.org/10.11646/zootaxa.3937.1.3" volume="3937" year="2015">Rakotoarison et al. 2015</bibRefCitation>
, 
<bibRefCitation author="Rakotoarison, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="263" pagination="271 - 288" refId="B23" refString="Rakotoarison, A, Scherz, MD, Bletz, MC, Razafindraibe, JH, Glaw, F, Vences, M, 2019b. Description of the lucky Cophyla (Microhylidae, Cophylinae), a new arboreal frog from Marojejy National Park in north-eastern Madagascar. Zootaxa 4651: 271 - 288" title="Description of the lucky Cophyla (Microhylidae, Cophylinae), a new arboreal frog from Marojejy National Park in north-eastern Madagascar." volume="4651" year="2019 b">2019b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="description">
<paragraph pageId="0" pageNumber="263">Description of the holotype.</paragraph>
<paragraph pageId="0" pageNumber="263">
Adult male in good state of preservation, some muscle tissue removed from right thigh; snout-vent length 25.6 mm (for further measurements see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Original morphometric measurements of Platypelis from the Sorata Massif, ND: Not determined. See Materials and Methods for other abbreviations. Specimens marked with UADBA-A are to be catalogued in the UADBA collection. The bolded row refers to the holotype of the new species." httpUri="http://table.plazi.org/id/A4D066085CE482AE3CEF7C5BE748372C" pageId="0" pageNumber="263" tableUuid="A4D066085CE482AE3CEF7C5BE748372C">1</tableCitation>
); body plump; head slightly wider than long, not wider than body; snout slightly rounded in dorsal view and blunt in lateral view; nostrils not protuberant, nearer to tip of snout than to eye; canthus rostralis distinct, straight; loreal region straight, slightly oblique; tympanum distinct, 60% of eye diameter; supratympanic fold distinct, starting at the posterior border of the eye and ending anterior to the forelimb; tongue long, broadening posteriorly, attached anteriorly, not notched; maxillary teeth present and vomerine teeth absent; choanae rounded. Forelimbs robust; subarticular tubercles single, indistinct; outer metacarpal tubercle small, rounded; prepollex/inner metacarpal tubercle distinct, forming a distinct protuberance at base of first finger; hand without webbing; fingers distinctly broadly rounded to slightly bilobate, with lateral fringes, smaller than discs of fingers; relative length of fingers 1&lt;2&lt;4&lt;3; nuptial pads absent. Hindlimbs slender; tibiotarsal articulation reaching between forelimb and tympanum when hindlimb adpressed along body; tibia length 47% of SVL; inner metatarsal tubercle small, oval; outer metatarsal tubercle absent; webbing between toes weakly developed, with traces of webbing between third and fourth toe; subarticular tubercles on toes single; toes flattened and their discs relatively broad, broadly rounded to slightly bilobate; relative length of toes 1&lt;2&lt;3&lt;5&lt;4; third toe slightly shorter than fifth, dorsal skin smooth, without dorsolateral folds, ventral skin smooth on throat and chest and moderately granular on belly.
</paragraph>
<caption ID-Table-UUID="A4D066085CE482AE3CEF7C5BE748372C" httpUri="http://table.plazi.org/id/A4D066085CE482AE3CEF7C5BE748372C" pageId="0" pageNumber="263" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="263">
<emphasis bold="true" pageId="0" pageNumber="263">Table 1.</emphasis>
Original morphometric measurements of 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
from the Sorata Massif, ND: Not determined. See Materials and Methods for other abbreviations. Specimens marked with UADBA-A are to be catalogued in the UADBA collection. The bolded row refers to the holotype of the new species.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="263">
<table pageId="0" pageNumber="263">
<tr pageId="0" pageNumber="263">
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">Collection/Catalogue number</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">Field number</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">Sex</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">SVL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">HW</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">HL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">TD</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">ED</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">END</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">NSD</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">NND</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">FORL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">HAL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">HIL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">FOTL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">FOL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">TIBL</th>
<th colspan="1" pageId="0" pageNumber="263" rowspan="1">RHL</th>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="18" pageId="0" pageNumber="263" rowspan="1">
<taxonomicName genus="P." lsidName="P. laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">P. laetus</emphasis>
</taxonomicName>
sp. nov.
</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 5651/2012</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3588</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">male</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">24.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">16.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">37.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">17.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">ZSM 5652/2012</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">FGZC 3761</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">male</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">25.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">8.6</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">8.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">2.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">3.3</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">1.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">1.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">2.5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">16.2</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">7.7</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">36.8</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">17.4</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">10.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">12.0</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="263">reaches tympanum</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 5653/2012</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3762</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">16.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">34.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">16.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">9.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="18" pageId="0" pageNumber="263" rowspan="1">
<taxonomicName genus="P." lsidName="P." pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">P.</emphasis>
</taxonomicName>
sp. Ca12
</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">UADBA-A</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3781</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">juvenile</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">16.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">5.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">5.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">20.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">5.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches eye</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">UADBA-A</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3782</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">juvenile</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">12.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">4.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">0.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">0.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">9.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">18.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches eye</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="18" pageId="0" pageNumber="263" rowspan="1">
<taxonomicName genus="P." lsidName="P. tsaratananaensis" pageId="0" pageNumber="263" rank="species" species="tsaratananaensis">
<emphasis italics="true" pageId="0" pageNumber="263">P. tsaratananaensis</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">UADBA-A</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3646</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">15.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">32.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">15.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 5654/2012</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGCC 3647</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">14.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">31.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">14.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 5655/2012</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3648</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">20.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">13.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">28.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">13.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 5656/2012</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3649</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">22.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">13.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">13.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">UADBA-A</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3765</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">7.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.4</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">14.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">32.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">15.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="18" pageId="0" pageNumber="263" rowspan="1">
<taxonomicName genus="P." lsidName="P. alticola" pageId="0" pageNumber="263" rank="species" species="alticola">
<emphasis italics="true" pageId="0" pageNumber="263">P. alticola</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="263">
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ZSM 1615/2018</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">FGZC 3669</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">ND</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">32.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">25.5</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">10.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">48.3</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">23.2</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">14.7</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">13.6</td>
<td colspan="1" pageId="0" pageNumber="263" rowspan="1">reaches tympanum</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="263">
After nine years of preservation in 70% ethanol, the dorsum is light beige with a brown, teddy-bear-shaped patch flecked with cream from between the eyes to the inguinal region. The nostril is surrounded with brown. The lateral surface is homogenously light beige flecked with small brown dots. The flank coloration merges with the ventral coloration. The ventral trunk and the chin are light beige. The ventral thigh is beige spotted with brown in the cloacal region. The shank, tarsus and foot are ventrally beige. Dorsally, the thigh is beige with light brown crossbands. The posterodorsal surface of the thigh is beige spotted with light brown. The shank is beige with a brown crossband. The coloration of the distal portion of the shank merges with the coloration of the tarsus, extending to the toes. The arms are beige with a brown crossband. The coloration in life is shown in Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs in life of Platypelis laetus sp. nov. from the Sorata massif: a-c ZSM 5652 / 2012 (FGZC 3761), holotype; d, e ZSM 5651 / 2012 (FGZC 3588), paratype; f, g ZSM 5653 / 2012 (FGZC 3762), paratype; h Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652 / 2012 and ZSM 5653 / 2012." figureDoi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263">3</figureCitation>
(in dorsal, lateral and ventral view).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="variation">
<paragraph pageId="0" pageNumber="263">Variation.</paragraph>
<paragraph pageId="0" pageNumber="263">
For variation in measurements, see Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Original morphometric measurements of Platypelis from the Sorata Massif, ND: Not determined. See Materials and Methods for other abbreviations. Specimens marked with UADBA-A are to be catalogued in the UADBA collection. The bolded row refers to the holotype of the new species." httpUri="http://table.plazi.org/id/A4D066085CE482AE3CEF7C5BE748372C" pageId="0" pageNumber="263" tableUuid="A4D066085CE482AE3CEF7C5BE748372C">1</tableCitation>
. In general, the two examined adult paratypes agree well with the holotype, but with the following differences: The holotype and FGZC 3588 have plump bodies and FGZC 3762 is rather slim. Relative toe length is variable: the third toe is distinctly shorter than fifth on one foot, and slightly shorter on the other foot in FGZC 3588, slightly shorter in the holotype FGZC 3761, and very slightly shorter to equal in FGZC 3762. Overall, specimens in life have a dorsal surface with some irregular granules. In coloration, FGZC 3762 is the darkest specimen of the type series in preservative and FGZC 3588 is the lightest. The teddy-bear pattern of the dorsum of the holotype and FGZC 3588 is absent in FGZC 3762.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="distribution">
<paragraph pageId="0" pageNumber="263">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="263">
The species is known only from the Sorata Massif, northern Madagascar, at elevations of 1339-1541 m above sea level (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Approximate geographical location of occurrence records of the focal Platypelis species in our study (modified from Rakotoarison et al. 2012; showing only records for which molecular data are available). The map indicates elevation (green, lowest; red-yellow, highest). Localities and species occurring at these sites are as follows: (1 - 4) campsites on the Mangindrano-Maromokotro trail, Tsaratanana Massif (Antevialambazaha, Matsabory Maiky, Bepia, Andranomadio); (5) Bemanevika forest; (6) Andrevorevo; (7) Marojejy; (8) Sorata; (9) Andravory. Occurrence records of P. alticola (at sites 1 - 3, and a deep genetic lineage at 8), P. tsaratananaensis (at sites 1 - 3, and deep genetic lineages at 7 and 8), P. olgae (between sites 3 and 4), P. laetus sp. nov. and P. Ca 12 (site 8), and P. sp. Ca 13 (site 9)." figureDoi="10.3897/zse.96.47088.figure2" httpUri="https://binary.pensoft.net/fig/418558" pageId="0" pageNumber="263">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="natural history">
<paragraph pageId="0" pageNumber="263">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="263">
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov. occurs in rainforest on the Sorata Massif, but most of the specimens were collected in the bamboo forest of the massif. The holotype was calling from a bamboo hole at about 5 m above the ground. The bamboo node was occupied by another specimen, was water-filled, and contained 35 whitish eggs and embryos, probably of this species, in at least two different developmental stages (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs in life of Platypelis laetus sp. nov. from the Sorata massif: a-c ZSM 5652 / 2012 (FGZC 3761), holotype; d, e ZSM 5651 / 2012 (FGZC 3588), paratype; f, g ZSM 5653 / 2012 (FGZC 3762), paratype; h Eggs and embryos found in the same bamboo hole with the specimens ZSM 5652 / 2012 and ZSM 5653 / 2012." figureDoi="10.3897/zse.96.47088.figure3" httpUri="https://binary.pensoft.net/fig/418559" pageId="0" pageNumber="263">3h</figureCitation>
). Another male was found on a palm tree at around 4 m above the ground. Calling occurred around dusk.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="etymology">
<paragraph pageId="0" pageNumber="263">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="263">
The specific epithet is a masculine Latin adjective meaning 
<normalizedToken originalValue="“happy”">&quot;happy&quot;</normalizedToken>
. The new species is so named in reflection of the joy and happiness of the first author to get to work on the cophyline microhylid frogs of Madagascar.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="vocalization">
<paragraph pageId="0" pageNumber="263">Vocalization.</paragraph>
<paragraph pageId="0" pageNumber="263">
The advertisement call of 
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov. consists of a single short tonal note (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5.. Spectrogram and oscillogram of one advertisement call of Platypelis laetus sp. nov. from a call series recorded on 30 November 2012 at the type locality in Sorata, northern Madagascar. Although not unequivocally verifiable, the depicted call most probably was emitted by the male holotype (ZSM 5652 / 2012)." figureDoi="10.3897/zse.96.47088.figure5" httpUri="https://binary.pensoft.net/fig/418561" pageId="0" pageNumber="263">5</figureCitation>
) repeated at regular intervals in long call series. Analysis of 43 calls belonging to three different males recorded on 30 November 2012 in Sorata revealed the following numerical parameters (ca. 15 °C air temperature): note duration (= call duration) 73-88 ms (83.4 
<normalizedToken originalValue="±">+/-</normalizedToken>
4.3 ms), inter-note interval 1465-2378 ms (1940 
<normalizedToken originalValue="±">+/-</normalizedToken>
255 ms), and dominant frequency 4707-4793 Hz (4742 
<normalizedToken originalValue="±">+/-</normalizedToken>
38 Hz). Each note exhibits some regular upward frequency modulation starting at around 4600-4700 Hz and terminating at 4770-4900 Hz. This narrow frequency band also constitutes the prevalent bandwidth of the call, although two additional rather weak frequency bands are present at around 2400 Hz and 7200 Hz. There is also a slight degree of amplitude modulation recognizable within notes, but the pattern of these modulations seems to be rather variable among notes (and possibly may constitute an artifact because of some recording device damage; see Materials and methods).
</paragraph>
<caption doi="10.3897/zse.96.47088.figure5" httpUri="https://binary.pensoft.net/fig/418561" pageId="0" pageNumber="263" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="263">
<emphasis bold="true" pageId="0" pageNumber="263">Figure 5.</emphasis>
. Spectrogram and oscillogram of one advertisement call of 
<taxonomicName authorityName="Rakotoarison &amp; Scherz &amp; Köhler &amp; Ratsoavina &amp; Hawlitschek &amp; Megson &amp; Vences &amp; Glaw" authorityYear="2020" genus="Platypelis" lsidName="Platypelis laetus" pageId="0" pageNumber="263" rank="species" species="laetus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis laetus</emphasis>
</taxonomicName>
sp. nov. from a call series recorded on 30 November 2012 at the type locality in Sorata, northern Madagascar. Although not unequivocally verifiable, the depicted call most probably was emitted by the male holotype (ZSM 5652/2012).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="263" type="range extensions of other platypelis species">
<paragraph pageId="0" pageNumber="263">
Range extensions of other 
<taxonomicName genus="Platypelis" lsidName="Platypelis" pageId="0" pageNumber="263" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="263">Platypelis</emphasis>
</taxonomicName>
species
</paragraph>
</subSubSection>
</treatment>
</document>