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<document id="919CD6E90A7FAF3081FF186FF7943E7A" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFDE842C8CFBFC33DA54FD51" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes quevedoensis Flowers 2009" docType="treatment" docVersion="8" lastPageNumber="73" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="68" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="71FEABF783BAA194470DD79EDFB657C1">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:namePart id="C07D9EF9A0A6FAC4474A0E1362EB97A9">Kluge, Nikita J.</mods:namePart>
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<mods:date id="DC8D2B8B751079E6A26FE00B27F6A7A5">2020</mods:date>
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<mods:number id="75560CA7D3D9F1E7AA40F6623C948B8E">2020-03-26</mods:number>
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<treatment id="039787A6FFDE842C8CFBFC33DA54FD51" ID-DOI="http://doi.org/10.5281/zenodo.3811793" ID-GBIF-Taxon="163420776" ID-Zenodo-Dep="3811793" LSID="urn:lsid:plazi:treatment:039787A6FFDE842C8CFBFC33DA54FD51" httpUri="http://treatment.plazi.org/id/039787A6FFDE842C8CFBFC33DA54FD51" lastPageId="72" lastPageNumber="73" pageId="67" pageNumber="68">
<subSubSection id="C324653BFFDE84278CFBFC33D9ADFC4D" box="[151,659,981,1008]" pageId="67" pageNumber="68" type="nomenclature">
<paragraph id="8B8136B0FFDE84278CFBFC33D9ADFC4D" blockId="67.[151,659,981,1043]" box="[151,659,981,1008]" pageId="67" pageNumber="68">
<heading id="D0C981DCFFDE84278CFBFC33D9ADFC4D" bold="true" box="[151,659,981,1008]" fontSize="10" level="2" pageId="67" pageNumber="68" reason="2">
<emphasis id="B94AEAA2FFDE84278CFBFC33D9ADFC4D" bold="true" box="[151,659,981,1008]" pageId="67" pageNumber="68">
8.
<taxonomicName id="4C3E4D33FFDE84278CD9FC33D9ADFC4D" ID-CoL="56LKZ" authority="Flowers 2009" authorityName="Flowers" authorityYear="2009" box="[181,659,981,1008]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFDE84278CD9FC33DAD8FC52" bold="true" box="[181,486,981,1008]" italics="true" pageId="67" pageNumber="68">Thraulodes quevedoensis</emphasis>
<bibRefCitation id="EFAF4B41FFDE84278D81FC33D9ADFC4D" author="Flowers, R. W." box="[493,659,981,1007]" pageId="67" pageNumber="68" pagination="55 - 61" refId="ref69757" refString="Flowers, R. W. (2009) A new species of Thraulodes (Ephemeroptera: Leptophlebiidae, Atalophlebiinae) from a highly altered river in western Ecuador. Zootaxa, 2052 (1), 55 - 61. https: // doi. org / 10.11646 / zootaxa. 2052.1.4" type="journal article" year="2009">Flowers 2009</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFDE84278CFBFC1FDA7DFBB1" box="[151,323,1016,1043]" pageId="67" pageNumber="68" type="description">
<paragraph id="8B8136B0FFDE84278CFBFC1FDA7DFBB1" blockId="67.[151,659,981,1043]" box="[151,323,1016,1043]" pageId="67" pageNumber="68">
(
<figureCitation id="13052A35FFDE84278CF3FC1FDA02FBB1" box="[159,316,1016,1043]" captionStart-0="FIGURES 250–259" captionStart-1="FIGURES 260–266" captionStart-2="FIGURES 267–273" captionStart-3="FIGURES 274–275" captionStartId-0="68.[151,264,1770,1795]" captionStartId-1="69.[151,264,1402,1427]" captionStartId-2="70.[151,264,1687,1712]" captionStartId-3="71.[151,264,1924,1949]" captionTargetBox-0="[151,1436,798,1751]" captionTargetBox-1="[151,1436,166,1382]" captionTargetBox-2="[151,1424,174,1654]" captionTargetBox-3="[151,1398,166,1905]" captionTargetId-0="figure@68.[151,1436,798,1751]" captionTargetId-1="figure@69.[151,1436,166,1382]" captionTargetId-2="figure@70.[151,1436,166,1668]" captionTargetId-3="figure@71.[151,1398,166,1905]" captionTargetPageId-0="68" captionTargetPageId-1="69" captionTargetPageId-2="70" captionTargetPageId-3="71" captionText-0="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." captionText-1="FIGURES 260–266. Thraulodes quevedoensis. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing." captionText-2="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." captionText-3="FIGURES 274–275. Thraulodes quevedoensis, eggs." figureDoi-0="http://doi.org/10.5281/zenodo.3736969" figureDoi-1="http://doi.org/10.5281/zenodo.3736971" figureDoi-2="http://doi.org/10.5281/zenodo.3736973" figureDoi-3="http://doi.org/10.5281/zenodo.3736975" httpUri-0="https://zenodo.org/record/3736969/files/figure.png" httpUri-1="https://zenodo.org/record/3736971/files/figure.png" httpUri-2="https://zenodo.org/record/3736973/files/figure.png" httpUri-3="https://zenodo.org/record/3736975/files/figure.png" pageId="67" pageNumber="68">Figs 250–275</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFDE84278CFBFBDBD888FB31" pageId="67" pageNumber="68" type="reference_group">
<paragraph id="8B8136B0FFDE84278CFBFBDBD837FBF7" blockId="67.[151,1189,1084,1171]" box="[151,777,1084,1109]" pageId="67" pageNumber="68">
<treatmentCitationGroup id="AB2E119EFFDE84278CFBFBDBD837FBF7" box="[151,777,1084,1109]" pageId="67" pageNumber="68">
<taxonomicName id="4C3E4D33FFDE84278CFBFBDBDA34FBF7" ID-CoL="648C7" authorityName="Ulmer" authorityYear="1920" box="[151,266,1085,1109]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFDE84278CFBFBDBDA34FBF7" box="[151,266,1085,1109]" italics="true" pageId="67" pageNumber="68">Thraulodes</emphasis>
</taxonomicName>
? sp. 5:
<treatmentCitation id="0A9F10A1FFDE84278D0CFBDAD922FBF7" author="Roback, S. S." box="[352,540,1084,1109]" page="146" pageId="67" pageNumber="68" year="1955">Roback 1955: 146</treatmentCitation>
, figs 199–2008 (larva).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136B0FFDE84278CFBFBBADF9BFBD6" blockId="67.[151,1189,1084,1171]" box="[151,1189,1115,1140]" pageId="67" pageNumber="68">
<treatmentCitationGroup id="AB2E119EFFDE84278CFBFBBADF9BFBD6" box="[151,1189,1115,1140]" pageId="67" pageNumber="68">
<taxonomicName id="4C3E4D33FFDE84278CFBFBBADA34FBD6" ID-CoL="648C7" authorityName="Ulmer" authorityYear="1920" box="[151,266,1116,1140]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEAA2FFDE84278CFBFBBADA34FBD6" box="[151,266,1116,1140]" italics="true" pageId="67" pageNumber="68">Thraulodes</emphasis>
</taxonomicName>
sp. «allied to
<taxonomicName id="4C3E4D33FFDE84278DF5FBBBD85CFBD6" ID-CoL="56LL3" authority="&quot;: Traver &amp; Edmunds 1967: 385" authorityName="&quot;: Traver &amp; Edmunds" authorityPageNumber="385" authorityYear="1967" box="[409,866,1115,1140]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="B94AEAA2FFDE84278DF5FBBBD929FBD6" box="[409,535,1116,1140]" italics="true" pageId="67" pageNumber="68">T. schlingeri</emphasis>
»:
<treatmentCitation id="0A9F10A1FFDE84278E5DFBBAD85CFBD6" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[561,866,1115,1140]" page="385" pageId="67" pageNumber="68" year="1967">
<bibRefCitation id="EFAF4B41FFDE84278E5DFBBAD85CFBD6" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[561,866,1115,1140]" pageId="67" pageNumber="68" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds 1967: 385</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 59 (
<specimenCount id="9D38FD39FFDE84278FD1FBBAD8EEFBD6" box="[957,976,1116,1140]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
and
<specimenCount id="9D38FD39FFDE8427886EFBBADF2BFBD1" box="[1026,1045,1116,1139]" pageId="67" pageNumber="68" type="female">♀</specimenCount>
imago, eggs).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136B0FFDE84278CFBFB9CD888FB31" blockId="67.[151,1189,1084,1171]" box="[151,950,1146,1171]" pageId="67" pageNumber="68">
<treatmentCitationGroup id="AB2E119EFFDE84278CFBFB9CD888FB31" box="[151,950,1146,1171]" pageId="67" pageNumber="68">
<treatmentCitation id="0A9F10A1FFDE84278CFBFB9CD96CFB31" ID-DOI="http://doi.org/10.5281/zenodo.6223336" ID-Zenodo-Dep="6223336" author="Flowers, R. W." box="[151,594,1146,1171]" httpUri="http://treatment.plazi.org/id/03B2EA3AB02CFFE4FF4F190104D3C84F" page="56" pageId="67" pageNumber="68" year="2009">
<taxonomicName id="4C3E4D33FFDE84278CFBFB9CD96CFB31" ID-CoL="56LKZ" authority="Flowers 2009: 56" authorityName="Flowers" authorityPageNumber="56" authorityYear="2009" box="[151,594,1146,1171]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFDE84278CFBFB9CDAA6FB30" box="[151,408,1146,1170]" italics="true" pageId="67" pageNumber="68">Thraulodes quevedoensis</emphasis>
<bibRefCitation id="EFAF4B41FFDE84278DF2FB9CD96CFB31" author="Flowers, R. W." box="[414,594,1146,1171]" pageId="67" pageNumber="68" pagination="55 - 61" refId="ref69757" refString="Flowers, R. W. (2009) A new species of Thraulodes (Ephemeroptera: Leptophlebiidae, Atalophlebiinae) from a highly altered river in western Ecuador. Zootaxa, 2052 (1), 55 - 61. https: // doi. org / 10.11646 / zootaxa. 2052.1.4" type="journal article" year="2009">Flowers 2009: 56</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–18 (
<specimenCount id="9D38FD39FFDE84278EA6FB9DD9E3FB31" box="[714,733,1147,1171]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
and
<specimenCount id="9D38FD39FFDE84278F62FB9DD81FFB30" box="[782,801,1147,1170]" pageId="67" pageNumber="68" type="female">♀</specimenCount>
imago, larva).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFDE84278CFBFB26DEB9FB5D" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="8B8136B0FFDE84278CFBFB26DEB9FB5D" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
<emphasis id="B94AEAA2FFDE84278CFBFB26DAB9FB79" bold="true" box="[151,391,1216,1243]" pageId="67" pageNumber="68">Material examined.</emphasis>
<materialsCitation id="3B563CEDFFDE84278DF8FB27DEB9FB5D" ID-GBIF-Occurrence="2594509908" collectingDate="2006-01-20" collectingDateMax="2006-02-25" collectingDateMin="2006-01-20" collectorName="N. Kluge" country="Peru" latitude="-9.5" location="Rio Cantaria" longLatPrecision="1299" longitude="-76.0" municipality="Tingo Maria" pageId="67" pageNumber="68" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Region Huanuco">
<collectingCountry id="F3297620FFDE84278DF8FB27DAE0FB79" box="[404,478,1217,1243]" name="Peru" pageId="67" pageNumber="68">PERU</collectingCountry>
,
<collectingRegion id="49FAF852FFDE84278D83FB27D98BFB79" box="[495,693,1217,1243]" country="Peru" name="Huanuco" pageId="67" pageNumber="68">Region Huanuco</collectingRegion>
,
<collectingMunicipality id="6BE5ACCAFFDE84278EAAFB27D862FB79" box="[710,860,1217,1243]" pageId="67" pageNumber="68">Tingo Maria</collectingMunicipality>
:
<location id="8EE1606BFFDE84278F00FB27DF3AFB79" LSID="urn:lsid:plazi:treatment:039787A6FFDE842C8CFBFC33DA54FD51:8EE1606BFFDE84278F00FB27DF3AFB79" box="[876,1028,1217,1243]" country="Peru" latitude="-9.5" longLatPrecision="1299" longitude="-76.0" municipality="Tingo Maria" name="Rio Cantaria" pageId="67" pageNumber="68" stateProvince="Region Huanuco">Rio Cantaria</location>
,
<location id="8EE1606BFFDE84278879FB27DF92FB79" LSID="urn:lsid:plazi:treatment:039787A6FFDE842C8CFBFC33DA54FD51:8EE1606BFFDE84278879FB27DF92FB79" box="[1045,1196,1217,1243]" country="Peru" latitude="-9.5" longLatPrecision="1299" longitude="-76.0" municipality="Tingo Maria" name="Puente Perez" pageId="67" pageNumber="68" stateProvince="Region Huanuco">Puente Perez</location>
(
<geoCoordinate id="EE0A5077FFDE842788AFFB27DE2DFB79" box="[1219,1299,1217,1243]" degrees="9" direction="south" minutes="21" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="-9.35">9°21’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFDE84278949FB26DEB1FB79" box="[1317,1423,1216,1243]" degrees="75" direction="west" minutes="58" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="-75.96667">75°58’W</geoCoordinate>
),
<date id="FF801070FFDE84278CFBFB03DA3BFB5D" box="[151,261,1253,1279]" pageId="67" pageNumber="68" value="2006-01-20">
<collectingDate id="EFC4E998FFDE84278CFBFB03DA3BFB5D" box="[151,261,1253,1279]" pageId="67" pageNumber="68" value="2006-01-20">20.I.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFDE84278D25FB03DA8AFB5D" box="[329,436,1253,1279]" pageId="67" pageNumber="68">N. Kluge</collectorName>
: 1 L-S/I
<specimenCount id="9D38FD39FFDE84278E7EFB03D918FB5D" box="[530,550,1253,1279]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
, 1 L-S-I
<specimenCount id="9D38FD39FFDE84278EE0FB03D9A4FB5D" box="[652,666,1253,1279]" pageId="67" pageNumber="68" type="female">♀</specimenCount>
;
<location id="8EE1606BFFDE84278ECAFB03D80CFB5D" LSID="urn:lsid:plazi:treatment:039787A6FFDE842C8CFBFC33DA54FD51:8EE1606BFFDE84278ECAFB03D80CFB5D" box="[678,818,1253,1279]" country="Peru" latitude="-9.5" longLatPrecision="1299" longitude="-76.0" municipality="Tingo Maria" name="Tingo Maria" pageId="67" pageNumber="68" stateProvince="Region Huanuco">Tingo Maria</location>
(
<geoCoordinate id="EE0A5077FFDE84278F2EFB03D8ACFB5D" box="[834,914,1253,1279]" degrees="9" direction="south" minutes="30" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="-9.5">9°30’S</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFDE84278FF2FB03DF36FAA2" box="[926,1032,1253,1280]" degrees="76" direction="west" minutes="00" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="-76.0">76°00’W</geoCoordinate>
),
<date id="FF801070FFDE84278870FB02DFAAFB5D" box="[1052,1172,1252,1279]" pageId="67" pageNumber="68" value="2006-02-25">
<collectingDate id="EFC4E998FFDE84278870FB02DFAAFB5D" box="[1052,1172,1252,1279]" pageId="67" pageNumber="68" value="2006-02-25">25.II.2006</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFDE842788BBFB03DE7CFB5D" box="[1239,1346,1253,1279]" pageId="67" pageNumber="68">N. Kluge</collectorName>
: 6 I
<specimenCount id="9D38FD39FFDE84278900FB03DEBDFB5D" box="[1388,1411,1253,1279]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFDE842C8CABFAEED8DEFE55" lastPageId="72" lastPageNumber="73" pageId="67" pageNumber="68" type="description">
<paragraph id="8B8136B0FFDE84278CABFAEEDAA6FA11" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
<emphasis id="B94AEAA2FFDE84278CABFAEEDA8BFA81" bold="true" box="[199,437,1288,1315]" pageId="67" pageNumber="68">
Descriptions.
<emphasis id="B94AEAA2FFDE84278D06FAEFDA8BFA81" bold="true" box="[362,437,1289,1315]" italics="true" pageId="67" pageNumber="68">Larva.</emphasis>
</emphasis>
CUTICULAR COLORATION (
<figureCitation id="13052A35FFDE84278F4FFAEFD8FCFA81" box="[803,962,1288,1315]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Figs 250–255</figureCitation>
). Dorsal side of head, thorax and abdomen mostly brown with few blanks: pronotum bordered by light laterally, with or without other blanks; fore protoptera with longitudinal veins either invisible, or slightly darker than background; abdominal terga with blanks on lateral parts, area mediad of tergalii attachments nearly unicolor brown. Femora mostly brown with blanks; tibiae and tarsi uniformly light brown.
</paragraph>
<paragraph id="8B8136B0FFDE84278CABFA5BDE67FA59" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
HYPODERMAL COLORATION. Unknown; probably, abdominal terga with paired blackish spots corresponding to midway gray spots of imago (as in
<figureCitation id="13052A35FFDE84278E05FA07D9F4FA59" box="[617,714,1505,1531]" captionStart="FIGURES 208–231" captionStartId="62.[151,264,188,213]" captionTargetId="figure@61.[151,1436,181,1894]" captionTargetPageId="61" captionText="FIGURES 208–231. Thraulodes marreroi. 208, margin of labrum with focus on posterior transverse setal row; 209, the same with focus on anterior transverse setal row (represented by field of irregularly situated setae) (arrows show four sensilla separating three denticles). 210–215 and 216–219, larval exuviae of two different specimens at the same magnification; 220–226, tergalii I–VII; 227, immature larva (with ocher and whitish cuticular coloration and blackish paired hypodermal markings on abdomen); 228, anterior side of hind femur; 229, anterior side of fore tibia; 230, hind tibia with focus on outer-posterior row of stout setae; 231, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736959" httpUri="https://zenodo.org/record/3736959/files/figure.png" pageId="67" pageNumber="68">Fig. 227</figureCitation>
). Tergalii brownish or colorless, tracheae black (
<figureCitation id="13052A35FFDE84278886FA07DE72FA59" box="[1258,1356,1504,1531]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Fig. 255</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFDE84278CABF9E3DAFBF929" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
SHAPE AND SETATION. Clypeus widened distally; labrum 1.15 times wider than clypeus (
<figureCitation id="13052A35FFDE842788A3F9E3DE0FF9BD" box="[1231,1329,1540,1567]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Fig. 251</figureCitation>
). Labrum widest at 0.4 length from base; initial fore margin (turned ventrally) without median emargination, with all 5 denticles wide; anterior transverse setal row regular (as in
<figureCitation id="13052A35FFDE84278E99F9ABD879F9C5" box="[757,839,1613,1639]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="67" pageNumber="68">Fig. 88</figureCitation>
), narrower than 3 denticles. Maxilla with 11 pectinate setae in apical-ventral row.
</paragraph>
<paragraph id="8B8136B0FFDE84278CABF973DF0AF971" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
Femora: Stout setae on anterior surface widened distally and blunt (
<figureCitation id="13052A35FFDE84278FD2F973DF1EF90D" box="[958,1056,1684,1711]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Fig. 256</figureCitation>
). Irregular row of hairs near inner margin absent on fore and middle femora, represented by few setae on hind femur.
</paragraph>
<paragraph id="8B8136B0FFDE84278CABF93BD920F8C1" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
Fore tibia (
<figureCitation id="13052A35FFDE84278D2AF93BDA95F955" box="[326,427,1756,1783]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Fig. 257</figureCitation>
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inneranterior row of stout setae represented by few (1–2) blunt stout setae near tibia base; inner field of stout pointed setae moderately dense (setae lengths subequal to distances between them), its setae subequal in lengths, forming 2 rows, setae of both rows smooth.
</paragraph>
<paragraph id="8B8136B0FFDE84278CABF88BDFA1F86D" blockId="67.[151,1437,1216,2035]" pageId="67" pageNumber="68">
Hind tibia (
<figureCitation id="13052A35FFDE84278D2BF88BDAD9F825" box="[327,487,1900,1927]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="67" pageNumber="68">Figs 258–259</figureCitation>
), each of three rows of stout setae—outer-anterior, outer-posterior and inner-anterior ones—consists of blunt stout setae of various lengths; hairs located between outer-anterior and outer-posterior rows, form one irregular row (besides row of hairs posteriad of outer-posterior row of stout setae).
</paragraph>
<paragraph id="8B8136B0FFDE84208CABF83FD901FF75" blockId="67.[151,1437,1216,2035]" lastBlockId="68.[151,1437,153,791]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
Claws of fore and middle legs with rigid portion bearing 7–6 larger denticles and 3 smaller denticles distad of them (
<figureCitation id="13052A35FFD984208CB2FF7FDA7EFF11" box="[222,320,153,179]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="68" pageNumber="69">Fig. 273</figureCitation>
); claw of hind leg with rigid portion bearing 8–9 denticles progressively larger distally; on all claws articulatory portion without denticles.
</paragraph>
<paragraph id="8B8136B0FFD984208CABFF07D8CBFF59" blockId="68.[151,1437,153,791]" box="[199,1013,224,251]" pageId="68" pageNumber="69">
Tergalii (
<figureCitation id="13052A35FFD984208D41FF07DAAEFF59" box="[301,400,224,251]" captionStart="FIGURES 250–259" captionStartId="68.[151,264,1770,1795]" captionTargetBox="[151,1436,798,1751]" captionTargetId="figure@68.[151,1436,798,1751]" captionTargetPageId="68" captionText="FIGURES 250–259. Thraulodes quevedoensis. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="68" pageNumber="69">Fig. 255</figureCitation>
): each lamella slender, tracheae without side branches.
</paragraph>
<paragraph id="8B8136B0FFD984208CABFEE3DE08FEE1" blockId="68.[151,1437,153,791]" pageId="68" pageNumber="69">
Male genitalia in last larval instar (
<figureCitation id="13052A35FFD984208E34FEE3D983FEBD" box="[600,701,261,287]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="68" pageNumber="69">Fig. 271</figureCitation>
): protogonostyli separated one from another by moderately deep emargination. Each protopenes lobe conic, without gonopore-bearing process; gonopore opened caudally.
</paragraph>
<paragraph id="8B8136B0FFD984208CABFEAAD92BFE71" blockId="68.[151,1437,153,791]" pageId="68" pageNumber="69">
<emphasis id="B94AEAA2FFD984208CABFEAADA7DFEC5" bold="true" box="[199,323,332,359]" italics="true" pageId="68" pageNumber="69">Subimago.</emphasis>
CUTICULAR COLORATION. Cuticle mostly colorless. Mesonotum with antelateroparapsidal suture contrastingly colored with dark brown; chromozones and other areas nearly completely colorless (
<figureCitation id="13052A35FFD984208940FE97DEAFFE2E" box="[1324,1425,369,396]" captionStart="FIGURES 260–266" captionStartId="69.[151,264,1402,1427]" captionTargetBox="[151,1436,166,1382]" captionTargetId="figure@69.[151,1436,166,1382]" captionTargetPageId="69" captionText="FIGURES 260–266. Thraulodes quevedoensis. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736971" httpUri="https://zenodo.org/record/3736971/files/figure.png" pageId="68" pageNumber="69">Fig. 262</figureCitation>
). On all legs tibia and tarsus tinges with light brownish. Cuticle of wings colorless, microtrichia light brownish. Other cuticle of body and legs colorless.
</paragraph>
<paragraph id="8B8136B0FFD984208CABFE3BDA1CFD9D" blockId="68.[151,1437,153,791]" pageId="68" pageNumber="69">
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of middle and hind legs (as in
<figureCitation id="13052A35FFD984208CD8FDC3DA2BFD9D" box="[180,277,549,575]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="68" pageNumber="69">Fig. 473</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFD984208CABFDAEDAB2FD25" blockId="68.[151,1437,153,791]" pageId="68" pageNumber="69">
<emphasis id="B94AEAA2FFD984208CABFDAEDA6AFDC1" bold="true" box="[199,340,584,611]" italics="true" pageId="68" pageNumber="69">Male imago</emphasis>
(
<figureCitation id="13052A35FFD984208D09FDAFD936FDC6" box="[357,520,585,612]" captionStart-0="FIGURES 260–266" captionStart-1="FIGURES 267–273" captionStartId-0="69.[151,264,1402,1427]" captionStartId-1="70.[151,264,1687,1712]" captionTargetBox-0="[151,1436,166,1382]" captionTargetBox-1="[151,1424,174,1654]" captionTargetId-0="figure@69.[151,1436,166,1382]" captionTargetId-1="figure@70.[151,1436,166,1668]" captionTargetPageId-0="69" captionTargetPageId-1="70" captionText-0="FIGURES 260–266. Thraulodes quevedoensis. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing." captionText-1="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi-0="http://doi.org/10.5281/zenodo.3736971" figureDoi-1="http://doi.org/10.5281/zenodo.3736973" httpUri-0="https://zenodo.org/record/3736971/files/figure.png" httpUri-1="https://zenodo.org/record/3736973/files/figure.png" pageId="68" pageNumber="69">Figs 260–272</figureCitation>
). Head brown. Antenna reddish, scapus apically darker. Dorsal eyes contiguous medially, dull orange.
</paragraph>
<paragraph id="8B8136B0FFD984208CABFD77DA08FCB5" blockId="68.[151,1437,153,791]" pageId="68" pageNumber="69">Pronotum, mesonotum and metanotum brownish orange. Lateral and ventral sides of prothorax uniformly ocher. Lateral sides of pterothorax brownish orange with ocher. Ventral side of pterothorax partly ocher; mesotho- racic episterna, anterior part of basisternum and furcasternal protuberances brown; median furcasternal impression lighter brown.</paragraph>
<caption id="DF416638FFD984208CFBF90CD8A7F8CC" ID-DOI="http://doi.org/10.5281/zenodo.3736969" ID-Zenodo-Dep="3736969" httpUri="https://zenodo.org/record/3736969/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,264,1770,1795]" targetBox="[151,1436,798,1751]" targetPageId="68">
<paragraph id="8B8136B0FFD984208CFBF90CD8A7F8CC" blockId="68.[151,1436,1770,1903]" pageId="68" pageNumber="69">
<emphasis id="B94AEAA2FFD984208CFBF90CDA52F8A1" bold="true" box="[151,364,1770,1795]" pageId="68" pageNumber="69">FIGURES 250–259.</emphasis>
<taxonomicName id="4C3E4D33FFD984208D1EF90DD94CF8A1" authorityName="Flowers" authorityYear="2009" box="[370,626,1771,1795]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFD984208D1EF90DD94CF8A1" box="[370,626,1771,1795]" italics="true" pageId="68" pageNumber="69">Thraulodes quevedoensis</emphasis>
</taxonomicName>
. 250–255, larval exuviae at the same magnification: 250–251, parts of head with labrum and mandibles; 252, right half of pronotum and mesonotum; 253–254, fore and hind legs; 255, abdominal terga; 256, anterior side of hind femur; 257, anterior side of fore tibia; 258, hind tibia with focus on outer-posterior row of stout setae; 259, the same with focus on outer-anterior and inner-anterior rows of stout setae.
</paragraph>
</caption>
<caption id="DF416638FFD884218CFBFA9CD91AFA79" ID-DOI="http://doi.org/10.5281/zenodo.3736971" ID-Zenodo-Dep="3736971" httpUri="https://zenodo.org/record/3736971/files/figure.png" pageId="69" pageNumber="70" startId="69.[151,264,1402,1427]" targetBox="[151,1436,166,1382]" targetPageId="69">
<paragraph id="8B8136B0FFD884218CFBFA9CD91AFA79" blockId="69.[151,1436,1402,1499]" pageId="69" pageNumber="70">
<emphasis id="B94AEAA2FFD884218CFBFA9CDA52FA31" bold="true" box="[151,364,1402,1427]" pageId="69" pageNumber="70">FIGURES 260–266.</emphasis>
<taxonomicName id="4C3E4D33FFD884218D1DFA9DD94FFA31" authorityName="Flowers" authorityYear="2009" box="[369,625,1403,1427]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFD884218D1DFA9DD94FFA31" box="[369,625,1403,1427]" italics="true" pageId="69" pageNumber="70">Thraulodes quevedoensis</emphasis>
</taxonomicName>
. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing.
</paragraph>
</caption>
<paragraph id="8B8136B0FFD884218CABF9EFDEB6F92D" blockId="69.[151,1437,1545,2039]" pageId="69" pageNumber="70">Cuticle of legs mostly colorless; on fore leg apex of femur, base of tibia, apex of tibia and 1st tarsal segment with cuticle orange. Legs with following hypodermal pigmentation on whitish-ocher background: Fore femur with or without diffusive reddish marking on anterior side of proximal half, with orange apical band, occupying distal 0.4 of femur, darker brown apically. Middle and hind femora with orange pre-apical band within apical 1/3 of femur.</paragraph>
<paragraph id="8B8136B0FFD884218CABF97FDEACF8BD" blockId="69.[151,1437,1545,2039]" pageId="69" pageNumber="70">
Fore and hind wings with dark reddish-brown macula on costal brace and bases of Sc and R; longitudinal and cross veins colorless. Costal cross veins proximad of bulla either absent, of few, colorless and very thin, so that visible only under high magnification (as in
<figureCitation id="13052A35FFD884218EEAF907D9D9F959" box="[646,743,1761,1787]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="69" pageNumber="70">Fig. 142</figureCitation>
). Pterostigmatic cross veins not dense, perpendicular to longitudinal veins or slightly oblique, non-branched (
<figureCitation id="13052A35FFD884218EDFF8E3D82BF882" box="[691,789,1797,1824]" captionStart="FIGURES 260–266" captionStartId="69.[151,264,1402,1427]" captionTargetBox="[151,1436,166,1382]" captionTargetId="figure@69.[151,1436,166,1382]" captionTargetPageId="69" captionText="FIGURES 260–266. Thraulodes quevedoensis. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736971" httpUri="https://zenodo.org/record/3736971/files/figure.png" pageId="69" pageNumber="70">Fig. 266</figureCitation>
). Hind wing with prominent costal projection (
<figureCitation id="13052A35FFD88421894EF8E3DEB8F8BD" box="[1314,1414,1797,1823]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="69" pageNumber="70">Fig. 272</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFD884218CABF8CFDBE6F855" blockId="69.[151,1437,1545,2039]" pageId="69" pageNumber="70">Abdominal hypodermal coloration: Tergum I brownish. Terga II–V ocher, translucent, tinged with orange medi- ally and on posterior margins; terga VI–VII orange; tergum VIII anteriorly-medially orange and posteriorly-laterally whitish; terga XI–X orange; some terga with a pair of small, indistinct midway spots; pleura with gray stigmatic spots. Abdominal sterna I–VI ocher, sterna VII–IX ocher with orange. Caudalii ocher, in proximal part with reddishbrown spot on each 2nd joining, in middle part with reddish-browns spot on each 4th joining, in distal half without spots.</paragraph>
<caption id="DF416638FFDB84228CFBF971D826F95A" ID-DOI="http://doi.org/10.5281/zenodo.3736973" ID-Zenodo-Dep="3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,264,1687,1712]" targetBox="[151,1424,174,1654]" targetPageId="70">
<paragraph id="8B8136B0FFDB84228CFBF971D826F95A" blockId="70.[151,1436,1687,1784]" pageId="70" pageNumber="71">
<emphasis id="B94AEAA2FFDB84228CFBF971DA53F912" bold="true" box="[151,365,1687,1712]" pageId="70" pageNumber="71">FIGURES 267–273.</emphasis>
<taxonomicName id="4C3E4D33FFDB84228D18F97ED948F912" authorityName="Flowers" authorityYear="2009" box="[372,630,1688,1712]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFDB84228D18F97ED948F912" box="[372,630,1688,1712]" italics="true" pageId="70" pageNumber="71">Thraulodes quevedoensis</emphasis>
</taxonomicName>
. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg.
</paragraph>
</caption>
<caption id="DF416638FFDA84238CFBF862D988F83F" ID-DOI="http://doi.org/10.5281/zenodo.3736975" ID-Zenodo-Dep="3736975" box="[151,694,1924,1949]" httpUri="https://zenodo.org/record/3736975/files/figure.png" pageId="71" pageNumber="72" startId="71.[151,264,1924,1949]" targetBox="[151,1398,166,1905]" targetPageId="71">
<paragraph id="8B8136B0FFDA84238CFBF862D988F83F" blockId="71.[151,694,1924,1949]" box="[151,694,1924,1949]" pageId="71" pageNumber="72">
<emphasis id="B94AEAA2FFDA84238CFBF862DA53F83F" bold="true" box="[151,365,1924,1949]" pageId="71" pageNumber="72">FIGURES 274–275.</emphasis>
<taxonomicName id="4C3E4D33FFDA84238D1FF863D94AF83F" authorityName="Flowers" authorityYear="2009" box="[371,628,1925,1949]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFDA84238D1FF863D94AF83F" box="[371,628,1925,1949]" italics="true" pageId="71" pageNumber="72">Thraulodes quevedoensis</emphasis>
</taxonomicName>
, eggs.
</paragraph>
</caption>
<paragraph id="8B8136B0FFD5842C8CABFF7FDA09FEE1" blockId="72.[151,1437,153,755]" pageId="72" pageNumber="73">
Genitalia (
<figureCitation id="13052A35FFD5842C8D50FF7FDAE4FF11" box="[316,474,153,179]" captionStart="FIGURES 260–266" captionStartId="69.[151,264,1402,1427]" captionTargetBox="[151,1436,166,1382]" captionTargetId="figure@69.[151,1436,166,1382]" captionTargetPageId="69" captionText="FIGURES 260–266. Thraulodes quevedoensis. 260–261, genitalia of male imago with focus on penis and on styliger; 262, right half of subimaginal exuviae of mesonotum; 263–264, male imago in ventral and lateral view; 265, abdominal terga and sterna of male imago; 266, apex of fore wing." figureDoi="http://doi.org/10.5281/zenodo.3736971" httpUri="https://zenodo.org/record/3736971/files/figure.png" pageId="72" pageNumber="73">Figs 260–261</figureCitation>
,
<figureCitation id="13052A35FFD5842C8D88FF7FD972FF16" box="[484,588,153,180]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="72" pageNumber="73">267–270</figureCitation>
): Styliger, gonostyli and penis ocher. Dorsal margin of styliger either roundish, without median extension (
<figureCitation id="13052A35FFD5842C8D94FF5BD964FF7A" box="[504,602,189,216]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="72" pageNumber="73">Fig. 269</figureCitation>
), or with poorly expressed extension (
<figureCitation id="13052A35FFD5842C886AFF5BDF57FF7A" box="[1030,1129,189,216]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="72" pageNumber="73">Fig. 268</figureCitation>
). Penis lobes narrowed and divergent apically, without lateral pouches; apices roundish, with ears not separated by emarginations. Telopenes in form of spear-like rolls, attached on dorsal side and directed medially-dorsally, curved, with groove opened dorsally-laterally.
</paragraph>
<paragraph id="8B8136B0FFD5842C8CABFEAADBEBFE29" blockId="72.[151,1437,153,755]" pageId="72" pageNumber="73">
<emphasis id="B94AEAA2FFD5842C8CABFEAADA46FEC5" bold="true" box="[199,376,332,359]" italics="true" pageId="72" pageNumber="73">Female imago.</emphasis>
Abdominal terga II–V brownish with ocher; in other respects coloration similar to that of male.
</paragraph>
<paragraph id="8B8136B0FFD5842C8CABFE73D8F5FE71" blockId="72.[151,1437,153,755]" pageId="72" pageNumber="73">
<emphasis id="B94AEAA2FFD5842C8CABFE73DA3FFE0D" bold="true" box="[199,257,405,431]" italics="true" pageId="72" pageNumber="73">Eggs</emphasis>
(
<figureCitation id="13052A35FFD5842C8D7CFE73DA8FFE0D" box="[272,433,404,431]" captionStart="FIGURES 274–275" captionStartId="71.[151,264,1924,1949]" captionTargetBox="[151,1398,166,1905]" captionTargetId="figure@71.[151,1398,166,1905]" captionTargetPageId="71" captionText="FIGURES 274–275. Thraulodes quevedoensis, eggs." figureDoi="http://doi.org/10.5281/zenodo.3736975" httpUri="https://zenodo.org/record/3736975/files/figure.png" pageId="72" pageNumber="73">Figs 274–275</figureCitation>
). Mostly barrel-shaped. Each KTC surrounded by ring-like cover. Protuberances densely located close to these rings; other chorion rugose, without protuberances.
</paragraph>
<paragraph id="8B8136B0FFD5842C8CABFE3AD8DEFE55" blockId="72.[151,1437,153,755]" box="[199,992,476,504]" pageId="72" pageNumber="73">
<emphasis id="B94AEAA2FFD5842C8CABFE3ADA70FE55" bold="true" box="[199,334,476,503]" pageId="72" pageNumber="73">Dimension.</emphasis>
Fore wing length (and approximate body length)
<quantity id="4CC69B55FFD5842C8F10FE3AD8E2FE55" box="[892,988,476,503]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="72" pageNumber="73" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFD5842C8CABFDE6DFAAFDBE" box="[199,1172,512,540]" pageId="72" pageNumber="73" type="distribution">
<paragraph id="8B8136B0FFD5842C8CABFDE6DFAAFDBE" blockId="72.[151,1437,153,755]" box="[199,1172,512,540]" pageId="72" pageNumber="73">
<emphasis id="B94AEAA2FFD5842C8CABFDE6DA5FFDB9" bold="true" box="[199,353,512,539]" pageId="72" pageNumber="73">Distribution.</emphasis>
<collectingCountry id="F3297620FFD5842C8D0BFDE7DAFAFDB9" box="[359,452,513,539]" name="Ecuador" pageId="72" pageNumber="73">Ecuador</collectingCountry>
(
<bibRefCitation id="EFAF4B41FFD5842C8DBFFDE7D94EFDB9" author="Flowers, R. W." box="[467,624,513,539]" pageId="72" pageNumber="73" pagination="55 - 61" refId="ref69757" refString="Flowers, R. W. (2009) A new species of Thraulodes (Ephemeroptera: Leptophlebiidae, Atalophlebiinae) from a highly altered river in western Ecuador. Zootaxa, 2052 (1), 55 - 61. https: // doi. org / 10.11646 / zootaxa. 2052.1.4" type="journal article" year="2009">Flowers 2009</bibRefCitation>
) and
<collectingCountry id="F3297620FFD5842C8EC2FDE7D9DFFDB9" box="[686,737,513,539]" name="Peru" pageId="72" pageNumber="73">Peru</collectingCountry>
(
<bibRefCitation id="EFAF4B41FFD5842C8E9CFDE7DF35FDB9" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[752,1035,513,539]" pageId="72" pageNumber="73" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds 1967</bibRefCitation>
; this study).
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFD5842C8CABFDC2DA54FD51" pageId="72" pageNumber="73" type="description">
<paragraph id="8B8136B0FFD5842C8CABFDC2DA54FD51" blockId="72.[151,1437,153,755]" pageId="72" pageNumber="73">
<emphasis id="B94AEAA2FFD5842C8CABFDC2DA5CFD9D" bold="true" box="[199,354,548,575]" pageId="72" pageNumber="73">Comparison.</emphasis>
<bibRefCitation id="EFAF4B41FFD5842C8D06FDC3D9A8FDE2" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[362,662,549,576]" pageId="72" pageNumber="73" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds (1967)</bibRefCitation>
reported that this species «bears strong resemblance to
<taxonomicName id="4C3E4D33FFD5842C897CFDC0DEA2FD9D" authorityName="&quot;: Traver &amp; Edmunds" authorityYear="1967" box="[1296,1436,549,575]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="B94AEAA2FFD5842C897CFDC0DEA2FD9D" box="[1296,1436,549,575]" italics="true" pageId="72" pageNumber="73">T. schlingeri</emphasis>
</taxonomicName>
in respect to form of eyes, once-banded femora, and tergum 6 similar in color to 7–10; faint midway spots, very tiny stigmatic dots».
<taxonomicName id="4C3E4D33FFD5842C8DEAFD8BD977FD25" authorityName="Flowers" authorityYear="2009" box="[390,585,621,647]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="quevedoensis">
<emphasis id="B94AEAA2FFD5842C8DEAFD8BD977FD25" box="[390,585,621,647]" italics="true" pageId="72" pageNumber="73">Th. quevedoensis</emphasis>
</taxonomicName>
well differs both from the sympatric species
<taxonomicName id="4C3E4D33FFD5842C8823FD8BDFD2FD25" authorityName="&quot;: Traver &amp; Edmunds" authorityYear="1967" box="[1103,1260,621,647]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="B94AEAA2FFD5842C8823FD8BDFD2FD25" box="[1103,1260,621,647]" italics="true" pageId="72" pageNumber="73">Th. schlingeri</emphasis>
</taxonomicName>
and the related Central American species
<taxonomicName id="4C3E4D33FFD5842C8DD6FD77D976FD09" authorityName="Chacon, Segnini &amp; Dominguez" authorityYear="1999" box="[442,584,657,683]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="marreroi">
<emphasis id="B94AEAA2FFD5842C8DD6FD77D976FD09" box="[442,584,657,683]" italics="true" pageId="72" pageNumber="73">Th. marreroi</emphasis>
</taxonomicName>
by penis structure, whose telopenes are not bent ventrally, but retain a dorsal position (
<figureCitation id="13052A35FFD5842C8D6DFD53DA5AFD72" box="[257,356,693,720]" captionStart="FIGURES 267–273" captionStartId="70.[151,264,1687,1712]" captionTargetBox="[151,1424,174,1654]" captionTargetId="figure@70.[151,1436,166,1668]" captionTargetPageId="70" captionText="FIGURES 267–273. Thraulodes quevedoensis. 267, genitalia of male imago in lateral view; 268, styliger and penis in ventral view; 269, genitalia of another specimen in ventral view; 270, the same, apex of penis in dorsal view; 271, larval protogonostyli and protopenis; 272, hind wing; 273, larval claw of middle leg." figureDoi="http://doi.org/10.5281/zenodo.3736973" httpUri="https://zenodo.org/record/3736973/files/figure.png" pageId="72" pageNumber="73">Fig. 267</figureCitation>
).
<bibRefCitation id="EFAF4B41FFD5842C8D1BFD53D99FFD72" author="Traver, J. R. &amp; Edmunds, G. F. Jr." box="[375,673,693,720]" pageId="72" pageNumber="73" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. &amp; Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver &amp; Edmunds (1967)</bibRefCitation>
correctly stated that «ova appear as in
<taxonomicName id="4C3E4D33FFD5842C8836FD50DFDBFD6D" authorityName="&quot;: Traver &amp; Edmunds" authorityYear="1967" box="[1114,1253,693,719]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="schlingeri">
<emphasis id="B94AEAA2FFD5842C8836FD50DFDBFD6D" box="[1114,1253,693,719]" italics="true" pageId="72" pageNumber="73">T. schlingeri</emphasis>
</taxonomicName>
» (compare
<figureCitation id="13052A35FFD5842C8900FD53DBC7FD51" captionStart="FIGURES 274–275" captionStartId="71.[151,264,1924,1949]" captionTargetBox="[151,1398,166,1905]" captionTargetId="figure@71.[151,1398,166,1905]" captionTargetPageId="71" captionText="FIGURES 274–275. Thraulodes quevedoensis, eggs." figureDoi="http://doi.org/10.5281/zenodo.3736975" httpUri="https://zenodo.org/record/3736975/files/figure.png" pageId="72" pageNumber="73">Figs 274–275</figureCitation>
and
<figureCitation id="13052A35FFD5842C8D5CFD3FDA60FD51" box="[304,350,729,755]" captionStart="FIGURE 207" captionStartId="59.[151,250,1418,1443]" captionTargetBox="[151,1436,258,1399]" captionTargetId="figure@59.[151,1436,258,1399]" captionTargetPageId="59" captionText="FIGURE 207. Thraulodes schlingeri, eggs." figureDoi="http://doi.org/10.5281/zenodo.3736957" httpUri="https://zenodo.org/record/3736957/files/figure.png" pageId="72" pageNumber="73">207</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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