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<document id="5DEDACB1E81E563464CC5686D843CB07" ID-CLB-Dataset="26396" ID-DOI="10.11646/zootaxa.4621.1.1" ID-GBIF-Dataset="a381190d-7f50-4a66-bc33-3a0417c9f89f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3257228" ID-ZooBank="3D77F329-9BB8-4231-90E4-6A9E0250FCDF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1561539959806" checkinUser="plazi" docAuthor="Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo &amp; Vieira, Juliana S." docDate="2019" docId="03978788B93CFF8CC1E36CE4AFA89C3D" docLanguage="en" docName="zootaxa.4621.1.1.pdf" docOrigin="Zootaxa 4621 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arhytodini Raffray 1890" docType="treatment" docVersion="13" lastPageNumber="74" masterDocId="FFAEFFF0B974FFC5C1746833AE7B997A" masterDocTitle="Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution" masterLastPageNumber="406" masterPageNumber="1" pageNumber="73" updateTime="1698737271145" updateUser="ExternalLinkService">
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<mods:title id="FEE384CF49E96C6841D53CC3FD7EF620">Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution</mods:title>
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<mods:namePart id="E0A609D853696C1A1BD71D5EC024AE57">Asenjo, Angélico</mods:namePart>
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<mods:namePart id="69618E6C9B34C970D518ABB2028CA244">Irmler, Ulrich</mods:namePart>
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<mods:namePart id="E66E79E2D10D9883B780A7C77EB05FA6">Klimaszewski, Jan</mods:namePart>
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<mods:namePart id="E2CFB60EEBEDCBC1C24ABD41251F9E76">Chandler, Donald S.</mods:namePart>
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<mods:namePart id="1A54F4228A39489B65D5F46006D8AEBB">Fierros-López, Hugo Eduardo</mods:namePart>
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<mods:namePart id="49C80A90F6FD3D3BEC65CC00341F375B">Vieira, Juliana S.</mods:namePart>
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<mods:title id="5C19EBED542737300FF47CA1105DC5ED">Zootaxa</mods:title>
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<mods:date id="46C6C24AF5CA3CEEE45730080B7D51DE">2019</mods:date>
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<mods:number id="7B9A838CE22A0BBAFDC1666821625E8B">4621</mods:number>
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<treatment id="03978788B93CFF8CC1E36CE4AFA89C3D" ID-GBIF-Taxon="158388844" LSID="urn:lsid:plazi:treatment:03978788B93CFF8CC1E36CE4AFA89C3D" httpUri="http://treatment.plazi.org/id/03978788B93CFF8CC1E36CE4AFA89C3D" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<subSubSection id="C3246515B93CFF8DC1E36CE4AC939C3A" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="8B81369EB93CFF8DC1E36CE4AC559D8B" blockId="72.[151,558,1239,1265]" box="[151,558,1239,1265]" pageId="72" pageNumber="73">
<heading id="D0C981F2B93CFF8DC1E36CE4AC559D8B" bold="true" box="[151,558,1239,1265]" fontSize="11" level="1" pageId="72" pageNumber="73" reason="1">
<emphasis id="B94AEA8CB93CFF8DC1E36CE4AC559D8B" bold="true" box="[151,558,1239,1265]" pageId="72" pageNumber="73">
Tribe
<taxonomicName id="4C3E4D1DB93CFF8DC1966CE4AC559D8B" ID-CoL="8XHKP" authority="Raffray, 1890 b" authorityName="Raffray" authorityYear="1890" box="[226,558,1239,1265]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="tribe" tribe="Arhytodini">
Arhytodini
<bibRefCitation id="EFAF4B6FB93CFF8DC0006CE4AC559D8B" author="Raffray, A." box="[372,558,1239,1265]" pageId="72" pageNumber="73" pagination="81 - 172" refId="ref228502" refString="Raffray, A. (1890 b) Etude sur les Pselaphides. V. Tableaux synoptiques. - Notes et synonymie. Revue d'Entomologie, 9, 81 - 172." type="journal article" year="1890">Raffray, 1890b</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36D14AC939C3A" blockId="72.[151,888,1319,1383]" box="[151,744,1319,1344]" pageId="72" pageNumber="73">
<emphasis id="B94AEA8CB93CFF8DC1E36D14AEAD9C3A" bold="true" box="[151,214,1319,1344]" pageId="72" pageNumber="73">0242.</emphasis>
<taxonomicName id="4C3E4D1DB93CFF8DC1A96D14AC939C3A" authority="Chandler &amp; Wolda, 1986: 521" authorityName="Chandler &amp; Wolda" authorityPageNumber="521" authorityYear="1986" box="[221,744,1319,1344]" class="Insecta" family="Staphylinidae" genus="Caccoplectinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93CFF8DC1A96D14AFF69C3A" bold="true" box="[221,397,1319,1344]" italics="true" pageId="72" pageNumber="73">Caccoplectinus</emphasis>
<bibRefCitation id="EFAF4B6FB93CFF8DC0E06D14AC939C3A" author="Chandler, D. S. &amp; Wolda, H." box="[404,744,1319,1344]" pageId="72" pageNumber="73" pagination="469 - 524" refId="ref184146" refString="Chandler, D. S. &amp; Wolda, H. (1986) Seasonality and diversity of Caccoplectus, with a review of the genus and description of a new genus, Caccoplectinus (Coleoptera: Pselaphidae). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 113, 469 - 524." type="journal article" year="1986">Chandler &amp; Wolda, 1986: 521</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B93CFF8CC1E36D7CACD89BF0" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8B81369EB93CFF8DC1E36D7CAD039C1D" blockId="72.[151,888,1319,1383]" box="[151,888,1358,1383]" pageId="72" pageNumber="73">
<typeStatus id="5485883CB93CFF8DC1E36D7CAEAA9C1D" box="[151,209,1359,1383]" pageId="72" pageNumber="73">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93CFF8DC0436D7DAD039C1D" authority="Chandler &amp; Wolda, 1886" authorityName="Chandler &amp; Wolda" authorityYear="1886" box="[311,888,1358,1383]" class="Insecta" family="Staphylinidae" genus="Caccoplectinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="afoveatus">
<emphasis id="B94AEA8CB93CFF8DC0436D7DAC2D9C1C" box="[311,598,1358,1383]" italics="true" pageId="72" pageNumber="73">Caccoplectinus afoveatus</emphasis>
Chandler &amp; Wolda, 1886
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36DAFADF69CCF" blockId="72.[151,909,1436,1578]" box="[151,909,1436,1461]" pageId="72" pageNumber="73">Only one species is known from and is restricted to South America.</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36DF7AD589CA6" blockId="72.[151,909,1436,1578]" box="[151,803,1475,1500]" pageId="72" pageNumber="73">
Key:
<bibRefCitation id="EFAF4B6FB93CFF8DC1A06DF0AC5A9CA6" author="Chandler, D. S. &amp; Wolda, H." box="[212,545,1475,1500]" pageId="72" pageNumber="73" pagination="469 - 524" refId="ref184146" refString="Chandler, D. S. &amp; Wolda, H. (1986) Seasonality and diversity of Caccoplectus, with a review of the genus and description of a new genus, Caccoplectinus (Coleoptera: Pselaphidae). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 113, 469 - 524." type="journal article" year="1986">Chandler &amp; Wolda 1986: 480</bibRefCitation>
(species of the genus).
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36DD9AF5E9F78" blockId="72.[151,909,1436,1578]" box="[151,293,1514,1539]" pageId="72" pageNumber="73">Distribution.</paragraph>
<paragraph id="8B81369EB93CFF8DC1DE6E22AC749F50" blockId="72.[151,909,1436,1578]" box="[170,527,1553,1578]" pageId="72" pageNumber="73">
South America:
<collectingCountry id="F329760EB93CFF8DC0156E22AC719F50" box="[353,522,1553,1578]" name="French Guiana" pageId="72" pageNumber="73">French Guiana</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36E6CAC479F0D" blockId="72.[151,691,1631,1695]" box="[151,572,1631,1656]" pageId="72" pageNumber="73">
<emphasis id="B94AEA8CB93CFF8DC1E36E6CAEAD9F02" bold="true" box="[151,214,1631,1656]" pageId="72" pageNumber="73">0243.</emphasis>
<taxonomicName id="4C3E4D1DB93CFF8DC1A96E6CAC479F0D" authority="Sharp, 1887 b: 22" authorityName="Sharp" authorityPageNumber="22" authorityYear="1887" box="[221,572,1631,1656]" class="Insecta" family="Staphylinidae" genus="Caccoplectus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93CFF8DC1A96E6CAF0E9F02" bold="true" box="[221,373,1631,1656]" italics="true" pageId="72" pageNumber="73">Caccoplectus</emphasis>
<bibRefCitation id="EFAF4B6FB93CFF8DC0086E6CAC479F0D" author="Sharp, D. S." box="[380,572,1631,1656]" pageId="72" pageNumber="73" pagination="1 - 71" refId="ref238527" refString="Sharp, D. S. (1887 b) Biologia Centrali-Americana, Insecta, Coleoptera, Pselaphidae, Scydmaenidae. 2 (1), 1 - 71, pls. 1 - 2." type="journal article" year="1887">Sharp, 1887b: 22</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36EB4ACC89FE5" blockId="72.[151,691,1631,1695]" box="[151,691,1670,1695]" pageId="72" pageNumber="73">
<typeStatus id="5485883CB93CFF8DC1E36EB4AEAA9FE5" box="[151,209,1671,1695]" pageId="72" pageNumber="73">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93CFF8DC0436EB5ACC89FE5" authority="Sharp, 1887" authorityName="Sharp" authorityYear="1887" box="[311,691,1670,1695]" class="Insecta" family="Staphylinidae" genus="Caccoplectus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="B94AEA8CB93CFF8DC0436EB5AC5F9FE5" box="[311,548,1670,1695]" italics="true" pageId="72" pageNumber="73">Caccoplectus celatus</emphasis>
Sharp, 1887
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36EE6AA629F97" blockId="72.[151,1436,1748,2007]" box="[151,1049,1748,1773]" pageId="72" pageNumber="73">
34 species (
<quantity id="4CC69B7BB93CFF8DC0696EE6AF3D9F97" box="[285,326,1748,1773]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="72" pageNumber="73" unit="in" value="1.0">1 in</quantity>
Mexican amber) are included in this genus from Latin America.
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36EC8AB0D9E41" blockId="72.[151,1436,1748,2007]" pageId="72" pageNumber="73">
Characters/Description:
<bibRefCitation id="EFAF4B6FB93CFF8DC0DF6EC8ACF19E6E" author="Raffray, A." box="[427,650,1787,1812]" pageId="72" pageNumber="73" pagination="1 - 400" refId="ref229028" refString="Raffray, A. (1904 a) Genera et catalogue des Pselaphides [continued]. Annales de la Societe Entomologique de France, 73, 1 - 400." type="journal article" year="1904">Raffray 1904a: 316</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB93CFF8DC20F6EC8AA219E6E" author="Raffray, A." box="[891,1114,1787,1812]" pageId="72" pageNumber="73" pagination="1 - 487" refId="ref229141" refString="Raffray, A. (1908 a) Coleoptera. Fam. Pselaphidae, pp. 1 - 487, 9 pls. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles." type="book chapter" year="1908">Raffray 1908a: 316</bibRefCitation>
(characters); O.
<bibRefCitation id="EFAF4B6FB93CFF8DC46B6EC8AE829E40" author="Park, O." pageId="72" pageNumber="73" pagination="227 - 267" refId="ref225424" refString="Park, O. (1944) New and little known Pselaphidae (Coleoptera) from Brazil, Colombia and Mexico, with keys to Mexican genera and species. Bulletin of the Chicago Academy of Sciences, 7 (4), 227 - 267." type="journal article" year="1944">Park 1944: 242</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB93CFF8DC0946F11AD579E40" author="Chandler, D. S. &amp; Wolda, H." box="[480,812,1826,1851]" pageId="72" pageNumber="73" pagination="469 - 524" refId="ref184146" refString="Chandler, D. S. &amp; Wolda, H. (1986) Seasonality and diversity of Caccoplectus, with a review of the genus and description of a new genus, Caccoplectinus (Coleoptera: Pselaphidae). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 113, 469 - 524." type="journal article" year="1986">Chandler &amp; Wolda 1986: 480</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB93CFF8DC2B26F11AAEC9E41" author="Chandler, D. S." box="[966,1175,1826,1851]" pageId="72" pageNumber="73" pagination="49 - 102" refId="ref183879" refString="Chandler, D. S. (2002) 5. Subfamilia Pselaphinae Latreille, 1802. In: Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (Eds.), Guia illustrada para los generos de Staphylinidae (Coleoptera) de Mexico [Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico]. Universidad de Guadalajara &amp; Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO). Mexico, pp. 49 - 102." type="book chapter" year="2002">Chandler 2002: 50</bibRefCitation>
(characters in key).
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36F79AA5B9EF2" blockId="72.[151,1436,1748,2007]" pageId="72" pageNumber="73">
Key: O.
<bibRefCitation id="EFAF4B6FB93CFF8DC1896F7AAFD49E18" author="Park, O." box="[253,431,1865,1890]" pageId="72" pageNumber="73" pagination="227 - 267" refId="ref225424" refString="Park, O. (1944) New and little known Pselaphidae (Coleoptera) from Brazil, Colombia and Mexico, with keys to Mexican genera and species. Bulletin of the Chicago Academy of Sciences, 7 (4), 227 - 267." type="journal article" year="1944">Park 1944: 251</bibRefCitation>
(Mexican species);
<bibRefCitation id="EFAF4B6FB93CFF8DC3E86F7AADF59E1B" author="Chandler, D. S." box="[668,910,1865,1890]" pageId="72" pageNumber="73" pagination="301 - 316" refId="ref183350" refString="Chandler, D. S. (1976 a) A revision of the genus Caccoplectus (Coleoptera: Pselaphidae). The Coleopterists Bulletin, 29 (4), 301 - 316. [Note: nominal year of publication 1975]" type="journal article" year="1976">Chandler 1976a: 304</bibRefCitation>
(species of the genus);
<bibRefCitation id="EFAF4B6FB93CFF8DC5DD6F7AABE79E18" author="Chandler, D. S." box="[1193,1436,1865,1890]" pageId="72" pageNumber="73" pagination="154 - 158" refId="ref183394" refString="Chandler, D. S. (1976 b) New species of Caccoplectus (Coleoptera: Pselaphidae). The Pan-Pacific Entomologist, 52, 154 - 158." type="journal article" year="1976">Chandler 1976b: 154</bibRefCitation>
(species of the genus);
<bibRefCitation id="EFAF4B6FB93CFF8DC0A66F43AD649EF2" author="Chandler, D. S. &amp; Wolda, H." box="[466,799,1904,1929]" pageId="72" pageNumber="73" pagination="469 - 524" refId="ref184146" refString="Chandler, D. S. &amp; Wolda, H. (1986) Seasonality and diversity of Caccoplectus, with a review of the genus and description of a new genus, Caccoplectinus (Coleoptera: Pselaphidae). Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 113, 469 - 524." type="journal article" year="1986">Chandler &amp; Wolda 1986: 480</bibRefCitation>
(species of the genus).
</paragraph>
<paragraph id="8B81369EB93CFF8DC1E36FA4AF5E9ED5" blockId="72.[151,1436,1748,2007]" box="[151,293,1943,1968]" pageId="72" pageNumber="73">Distribution.</paragraph>
<paragraph id="8B81369EB93CFF8DC1DE6F8CACCD9EAD" blockId="72.[151,1436,1748,2007]" box="[170,694,1982,2007]" pageId="72" pageNumber="73">
West Indies:
<collectingCountry id="F329760EB93CFF8DC0496F8DAF009EAD" box="[317,379,1982,2007]" name="Cuba" pageId="72" pageNumber="73">Cuba</collectingCountry>
,
<collectingCountry id="F329760EB93CFF8DC0F26F8DAF989EAD" box="[390,483,1982,2007]" name="Jamaica" pageId="72" pageNumber="73">Jamaica</collectingCountry>
, Trinidad-Tobago.
</paragraph>
<paragraph id="8B81369EB93DFF8CC1DE68ADACC599CC" blockId="73.[170,716,158,261]" box="[170,702,158,183]" pageId="73" pageNumber="74">
Central America:
<collectingCountry id="F329760EB93DFF8CC00668ADAFB199CD" box="[370,458,158,183]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC0A268ADAC2999CD" box="[470,594,158,183]" name="Guatemala" pageId="73" pageNumber="74">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC32968ACACC299CC" box="[605,697,159,182]" name="Panama" pageId="73" pageNumber="74">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB93DFF8CC1DE68F6ACB799A4" blockId="73.[170,716,158,261]" box="[170,716,197,222]" pageId="73" pageNumber="74">
South America:
<collectingCountry id="F329760EB93DFF8CC01568F6AC7199A4" box="[353,522,197,222]" name="French Guiana" pageId="73" pageNumber="74">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC36168F6AC0E99A4" box="[533,629,197,222]" name="Ecuador" pageId="73" pageNumber="74">Ecuador</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC3F468F6ACB399A4" box="[640,712,197,222]" name="Brazil" pageId="73" pageNumber="74">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB93DFF8CC1DE68DFAFC2987F" blockId="73.[170,716,158,261]" box="[170,441,236,261]" pageId="73" pageNumber="74">Other regions: Nearctic.</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36909AC459828" blockId="73.[151,706,314,378]" box="[151,574,314,339]" pageId="73" pageNumber="74">
<emphasis id="B94AEA8CB93DFF8CC1E36909AEAD9829" bold="true" box="[151,214,314,339]" pageId="73" pageNumber="74">0244.</emphasis>
<taxonomicName id="4C3E4D1DB93DFF8CC1A96909AC459828" authority="Besuchet, 1987: 231" authorityName="Besuchet" authorityPageNumber="231" authorityYear="1987" box="[221,574,314,339]" class="Insecta" family="Staphylinidae" genus="Pachacuti" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC1A96909AF2A9829" bold="true" box="[221,337,314,339]" italics="true" pageId="73" pageNumber="74">Pachacuti</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC02C6909AC459828" author="Besuchet, C." box="[344,574,314,339]" pageId="73" pageNumber="74" pagination="231 - 240" refId="ref174661" refString="Besuchet, C. (1987) Pselaphini neotropicaux nouveaux (Coleoptera: Pselaphidae). Archives des Sciences (Geneve), 40 (2), 231 - 240." type="journal article" year="1987">Besuchet, 1987: 231</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36951ACB99803" blockId="73.[151,706,314,378]" box="[151,706,353,378]" pageId="73" pageNumber="74">
<typeStatus id="5485883CB93DFF8CC1E36951AEAA9800" box="[151,209,354,378]" pageId="73" pageNumber="74">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93DFF8CC0436952ACB99803" authority="Besuchet, 1987" authorityName="Besuchet" authorityYear="1987" box="[311,706,353,378]" class="Insecta" family="Staphylinidae" genus="Pachacuti" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="huggerti">
<emphasis id="B94AEA8CB93DFF8CC0436952AC759800" box="[311,526,353,378]" italics="true" pageId="73" pageNumber="74">Pachacuti huggerti</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC3616952ACB99803" author="Besuchet, C." box="[533,706,353,378]" pageId="73" pageNumber="74" pagination="231 - 240" refId="ref174661" refString="Besuchet, C. (1987) Pselaphini neotropicaux nouveaux (Coleoptera: Pselaphidae). Archives des Sciences (Geneve), 40 (2), 231 - 240." type="journal article" year="1987">Besuchet, 1987</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E3699CAD2898B2" blockId="73.[151,851,431,534]" box="[151,851,431,456]" pageId="73" pageNumber="74">2 species are known from and are restricted to South America.</paragraph>
<paragraph id="8B81369EB93DFF8CC1E369E5AF5E9894" blockId="73.[151,851,431,534]" box="[151,293,470,495]" pageId="73" pageNumber="74">Distribution.</paragraph>
<paragraph id="8B81369EB93DFF8CC1DE69CEAC5C9B6C" blockId="73.[151,851,431,534]" box="[170,551,509,534]" pageId="73" pageNumber="74">
South America:
<collectingCountry id="F329760EB93DFF8CC01569CEAFBA9B6C" box="[353,449,509,534]" name="Ecuador" pageId="73" pageNumber="74">Ecuador</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC0B969CEAC589B6C" box="[461,547,509,534]" name="Bolivia" pageId="73" pageNumber="74">Bolivia</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36A78AC509B1E" blockId="73.[151,1122,587,729]" box="[151,555,587,612]" pageId="73" pageNumber="74">
<emphasis id="B94AEA8CB93DFF8CC1E36A78AEAD9B1E" bold="true" box="[151,214,587,612]" pageId="73" pageNumber="74">0245.</emphasis>
<taxonomicName id="4C3E4D1DB93DFF8CC1A96A78AC509B1E" authority="Westwood, 1870: 126" authorityName="Westwood" authorityPageNumber="126" authorityYear="1870" box="[221,555,587,612]" class="Insecta" family="Staphylinidae" genus="Rhytus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC1A96A78AF559B1E" bold="true" box="[221,302,587,612]" italics="true" pageId="73" pageNumber="74">Rhytus</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC0416A78AC509B1E" author="Westwood, J. O." box="[309,555,587,612]" pageId="73" pageNumber="74" pagination="125 - 132" refId="ref244187" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the coleopterous family Pselaphidae. Transactions of the Entomological Society of London, 1870 (2), 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870: 126</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36A40ACD89BF0" blockId="73.[151,1122,587,729]" box="[151,675,626,651]" pageId="73" pageNumber="74">
<typeStatus id="5485883CB93DFF8CC1E36A40AEAA9BF1" box="[151,209,627,651]" pageId="73" pageNumber="74">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93DFF8CC0436A41ACD89BF0" authority="Westwood, 1870" authorityName="Westwood" authorityYear="1870" box="[311,675,626,651]" class="Insecta" family="Staphylinidae" genus="Rhytus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B94AEA8CB93DFF8CC0436A41AFA59BF0" box="[311,478,626,651]" italics="true" pageId="73" pageNumber="74">Rhytus vestitus</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC0916A41ACD89BF0" author="Westwood, J. O." box="[485,675,626,651]" pageId="73" pageNumber="74" pagination="125 - 132" refId="ref244187" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the coleopterous family Pselaphidae. Transactions of the Entomological Society of London, 1870 (2), 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B93DFF8CC1BB6AAAAA199BC8" box="[207,1122,665,690]" pageId="73" pageNumber="74" type="reference_group">
<paragraph id="8B81369EB93DFF8CC1BB6AAAAA199BC8" blockId="73.[151,1122,587,729]" box="[207,1122,665,690]" pageId="73" pageNumber="74">
<treatmentCitationGroup id="AB2E11B0B93DFF8CC1BB6AAAAA199BC8" box="[207,1122,665,690]" pageId="73" pageNumber="74">
<taxonomicName id="4C3E4D1DB93DFF8CC1BB6AAAAC5B9BCB" ID-CoL="8XJQQ" authority="Reitter, 1882 b: 209" authorityName="Reitter" authorityPageNumber="209" authorityYear="1882" box="[207,544,665,690]" class="Insecta" family="Staphylinidae" genus="Arhytodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC1BB6AAAAF3A9BC8" box="[207,321,665,690]" italics="true" pageId="73" pageNumber="74">Arhytodes</emphasis>
<treatmentCitation id="0A9F108FB93DFF8CC0336AAAAC5B9BCB" author="Reitter, E." box="[327,544,665,690]" page="209" pageId="73" pageNumber="74" year="1882">
<bibRefCitation id="EFAF4B6FB93DFF8CC0336AAAAC5B9BCB" author="Reitter, E." box="[327,544,665,690]" pageId="73" pageNumber="74" pagination="177 - 211" refId="ref230040" refString="Reitter, E. (1882 b) Versuch einer systematischen Eintheilung der Clavigeriden und Pselaphiden. Verhandlungen des Naturforschenden Vereines in Brunn, 20, 177 - 211. [Note: nominal year of publication 1881]" type="journal article" year="1882">Reitter, 1882b: 209</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB93DFF8CC3846AA9AA4A9BCB" author="Newton, A. F. &amp; Chandler, D. S." box="[752,1073,665,690]" page="7" pageId="73" pageNumber="74" year="1989">
<bibRefCitation id="EFAF4B6FB93DFF8CC3846AA9AA4A9BCB" author="Newton, A. F. &amp; Chandler, D. S." box="[752,1073,665,690]" pageId="73" pageNumber="74" pagination="1 - 93" refId="ref220430" refString="Newton, A. F. &amp; Chandler, D. S. (1989) World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana Zoology (New Series), 53, 1 - 93." type="journal article" year="1989">Newton &amp; Chandler 1989: 7</bibRefCitation>
</treatmentCitation>
, 54]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B93DFF8CC0736AF2AFA89C3D" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="8B81369EB93DFF8CC0736AF2AD689BA2" blockId="73.[151,1122,587,729]" box="[263,787,704,729]" pageId="73" pageNumber="74">
<typeStatus id="5485883CB93DFF8CC0736AF2AF3A9BA3" box="[263,321,705,729]" pageId="73" pageNumber="74">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93DFF8CC0D36AF3AD689BA2" authority="Westwood, 1870" authorityName="Westwood" authorityYear="1870" box="[423,787,704,729]" class="Insecta" family="Staphylinidae" genus="Rhytus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B94AEA8CB93DFF8CC0D36AF3AC359BA2" box="[423,590,704,729]" italics="true" pageId="73" pageNumber="74">Rhytus vestitus</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC3216AF3AD689BA2" author="Westwood, J. O." box="[597,787,704,729]" pageId="73" pageNumber="74" pagination="125 - 132" refId="ref244187" refString="Westwood, J. O. (1870) Descriptions of twelve new exotic species of the coleopterous family Pselaphidae. Transactions of the Entomological Society of London, 1870 (2), 125 - 132. https: // doi. org / 10.1111 / j. 1365 - 2311.1870. tb 01868. x" type="journal article" year="1870">Westwood, 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36B3EAD299D4D" blockId="73.[151,1436,781,1079]" pageId="73" pageNumber="74">
15 species are known from and are restricted to Central and South America. Characters/Description:
<bibRefCitation id="EFAF4B6FB93DFF8CC0DF6B07ACF19A37" author="Raffray, A." box="[427,650,820,845]" pageId="73" pageNumber="74" pagination="1 - 487" refId="ref229141" refString="Raffray, A. (1908 a) Coleoptera. Fam. Pselaphidae, pp. 1 - 487, 9 pls. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 64. V. Verteneuil &amp; L. Desmet, Bruxelles." type="book chapter" year="1908">Raffray 1908a: 359</bibRefCitation>
(characters in key), 412 (characters as
<taxonomicName id="4C3E4D1DB93DFF8CC5276B07AABE9A37" authorityName="Reitter" authorityYear="1882" box="[1107,1221,820,845]" class="Insecta" family="Staphylinidae" genus="Arhytodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC5276B07AABE9A37" box="[1107,1221,820,845]" italics="true" pageId="73" pageNumber="74">Arhytodes</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAF4B6FB93DFF8CC5AB6B07AE829A0E" author="Chandler, D. S." pageId="73" pageNumber="74" pagination="339 - 344" refId="ref183603" refString="Chandler, D. S. (1992 a) Short-winged Mould Beetles of the tribe Arhytodini of Panama, with description of new taxa (Coleoptera: Pselaphidae: Pselaphinae). In: Quinter, D. &amp; Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, pp. 339 - 344." type="book chapter" year="1992">Chandler 1992a: 339</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB93DFF8CC0E76B6FAC1F9A0F" author="Chandler, D. S." box="[403,612,860,885]" pageId="73" pageNumber="74" pagination="49 - 102" refId="ref183879" refString="Chandler, D. S. (2002) 5. Subfamilia Pselaphinae Latreille, 1802. In: Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (Eds.), Guia illustrada para los generos de Staphylinidae (Coleoptera) de Mexico [Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico]. Universidad de Guadalajara &amp; Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO). Mexico, pp. 49 - 102." type="book chapter" year="2002">Chandler 2002: 50</bibRefCitation>
(characters in key). Key:
<bibRefCitation id="EFAF4B6FB93DFF8CC1A16BB0AFDF9AE1" author="Raffray, A." box="[213,420,899,924]" pageId="73" pageNumber="74" pagination="401 - 476" refId="ref229096" refString="Raffray, A. (1905) Genera et catalogue des Pselaphides [continued]. Annales de la Societe Entomologique de France, 73, 401 - 476, pl. 1 - 3. [Note: nominal year of publication 1904]" type="journal article" year="1905">Raffray 1905: 411</bibRefCitation>
(species group);
<bibRefCitation id="EFAF4B6FB93DFF8CC3136BB0AD539AE1" author="Raffray, A." box="[615,808,899,924]" pageId="73" pageNumber="74" pagination="15 - 52" refId="ref229224" refString="Raffray, A. (1909) Nouvelles especes de Pselaphides. Annales de la Societe Entomologique de France, 78, 15 - 52." type="journal article" year="1909">Raffray 1909: 47</bibRefCitation>
(species of the genus as
<taxonomicName id="4C3E4D1DB93DFF8CC5366BB0AACF9AE6" authorityName="Reitter" authorityYear="1882" box="[1090,1204,899,924]" class="Insecta" family="Staphylinidae" genus="Arhytodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC5366BB0AACF9AE6" box="[1090,1204,899,924]" italics="true" pageId="73" pageNumber="74">Arhytodes</emphasis>
</taxonomicName>
); O.
<bibRefCitation id="EFAF4B6FB93DFF8CC59B6BB0ABE79AE1" author="Park, O." box="[1263,1436,899,924]" pageId="73" pageNumber="74" refId="ref225339" refString="Park, O. (1942) A study in neotropical Pselaphidae. Northwestern University Studies in the Biological Sciences and Medicine, Number 1, 1, Northwestern University, Evanston and Chicago. x + 403 pp, 21 pls." type="book" year="1942">Park 1942: 344</bibRefCitation>
(species of the genus as
<taxonomicName id="4C3E4D1DB93DFF8CC0AB6B99AC2A9AB9" authorityName="Reitter" authorityYear="1882" box="[479,593,938,963]" class="Insecta" family="Staphylinidae" genus="Arhytodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC0AB6B99AC2A9AB9" box="[479,593,938,963]" italics="true" pageId="73" pageNumber="74">Arhytodes</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAF4B6FB93DFF8CC31C6B99AD289AB8" author="Chandler, D. S." box="[616,851,938,963]" pageId="73" pageNumber="74" pagination="339 - 344" refId="ref183603" refString="Chandler, D. S. (1992 a) Short-winged Mould Beetles of the tribe Arhytodini of Panama, with description of new taxa (Coleoptera: Pselaphidae: Pselaphinae). In: Quinter, D. &amp; Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, pp. 339 - 344." type="book chapter" year="1992">Chandler 1992a: 340</bibRefCitation>
(species of
<collectingCountry id="F329760EB93DFF8CC2AE6B99AA439AB8" box="[986,1080,938,962]" name="Panama" pageId="73" pageNumber="74">Panama</collectingCountry>
). Distribution. Central America:
<collectingCountry id="F329760EB93DFF8CC0066BCBAFB19D6B" box="[370,458,1016,1041]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC0A26BCAAC499D6A" box="[470,562,1017,1040]" name="Panama" pageId="73" pageNumber="74">Panama</collectingCountry>
. South America:
<collectingCountry id="F329760EB93DFF8CC0156C2DAFA19D4D" box="[353,474,1054,1079]" name="Venezuela" pageId="73" pageNumber="74">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC0926C13AC679D4D" box="[486,540,1056,1079]" name="Peru" pageId="73" pageNumber="74">Peru</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC3536C2DAC079D4D" box="[551,636,1054,1079]" name="Bolivia" pageId="73" pageNumber="74">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC3F36C2DACB59D4D" box="[647,718,1054,1079]" name="Brazil" pageId="73" pageNumber="74">Brazil</collectingCountry>
,
<collectingCountry id="F329760EB93DFF8CC3AE6C2DAD359D4D" box="[730,846,1054,1079]" name="Argentina" pageId="73" pageNumber="74">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36C5FAC319DFF" blockId="73.[151,729,1132,1197]" box="[151,586,1132,1157]" pageId="73" pageNumber="74">
<emphasis id="B94AEA8CB93DFF8CC1E36C5FAEAD9DFF" bold="true" box="[151,214,1132,1157]" pageId="73" pageNumber="74">0246.</emphasis>
<taxonomicName id="4C3E4D1DB93DFF8CC1A96C5FAC319DFF" authority="Chandler, 1992 a: 342" authorityName="Chandler" authorityPageNumber="342" authorityYear="1992" box="[221,586,1132,1157]" class="Insecta" family="Staphylinidae" genus="Woldenka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB93DFF8CC1A96C5FAF299DFF" bold="true" box="[221,338,1132,1157]" italics="true" pageId="73" pageNumber="74">Woldenka</emphasis>
<bibRefCitation id="EFAF4B6FB93DFF8CC02C6C5FAC319DFF" author="Chandler, D. S." box="[344,586,1132,1157]" pageId="73" pageNumber="74" pagination="339 - 344" refId="ref183603" refString="Chandler, D. S. (1992 a) Short-winged Mould Beetles of the tribe Arhytodini of Panama, with description of new taxa (Coleoptera: Pselaphidae: Pselaphinae). In: Quinter, D. &amp; Aiello, A. (Eds.), Insects of Panama and Mesoamerica: Selected Studies. Oxford University Press, Oxford, pp. 339 - 344." type="book chapter" year="1992">Chandler, 1992a: 342</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36CA6ACA29DD6" blockId="73.[151,729,1132,1197]" box="[151,729,1172,1197]" pageId="73" pageNumber="74">
<typeStatus id="5485883CB93DFF8CC1E36CA6AEAA9DD7" box="[151,209,1173,1197]" pageId="73" pageNumber="74">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB93DFF8CC0436CA7ACA29DD6" authority="Chandler, 1992" authorityName="Chandler" authorityYear="1992" box="[311,729,1172,1197]" class="Insecta" family="Staphylinidae" genus="Woldenka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="barroensis">
<emphasis id="B94AEA8CB93DFF8CC0436CA7AC5D9DD7" box="[311,550,1172,1197]" italics="true" pageId="73" pageNumber="74">Woldenka barroensis</emphasis>
Chandler, 1992
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36CD1ADE59D81" blockId="73.[151,926,1250,1352]" box="[151,926,1250,1275]" pageId="73" pageNumber="74">Only one species is known from and is restricted to Central America.</paragraph>
<paragraph id="8B81369EB93DFF8CC1E36D3AAF5E9C5B" blockId="73.[151,926,1250,1352]" box="[151,293,1289,1314]" pageId="73" pageNumber="74">Distribution.</paragraph>
<paragraph id="8B81369EB93DFF8CC1DE6D1CAFA89C3D" blockId="73.[151,926,1250,1352]" box="[170,467,1327,1352]" pageId="73" pageNumber="74">
Central America:
<collectingCountry id="F329760EB93DFF8CC0066D03AFB59C3D" box="[370,462,1328,1351]" name="Panama" pageId="73" pageNumber="74">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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