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<mods:title>Systematic revision of the genus Peronia Fleming, 1822 (Gastropoda, Euthyneura, Pulmonata, Onchidiidae)</mods:title>
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<mods:namePart>Aslam, Sadar</mods:namePart>
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<mods:namePart>Tan, Shau Hwai</mods:namePart>
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<treatment ID-GBIF-Taxon="168209567" LSID="urn:lsid:plazi:treatment:8E4FA523FB6C5383A3C9FEE739D8A786" httpUri="http://treatment.plazi.org/id/8E4FA523FB6C5383A3C9FEE739D8A786" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="8E4FA523-FB6C-5383-A3C9-FEE739D8A786" authority="(Cuvier, 1804)" baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">Peronia peronii (Cuvier, 1804)</taxonomicName>
<figureCitation captionStart="Figure 7" captionStartId="F9" captionText="Figure 7. Habitat, Peronia peronii, Papua New Guinea, Madang, limestone rocky intertidal (st PM 12)." figureDoi="10.3897/zookeys.972.52853.figure7" httpUri="https://binary.pensoft.net/fig/458307" pageId="0" pageNumber="1">Figs 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F10" captionText="Figure 8. Live animals, Peronia peronii, Mauritius A dorsal view, 140 mm long [5872] (MNHN-IM- 2019 - 1605) B dorsal view, 125 mm long [5874] (MNHN-IM- 2019 - 1605) C ventral view, same as A; D ventral view, same as B." figureDoi="10.3897/zookeys.972.52853.figure8" httpUri="https://binary.pensoft.net/fig/458308" pageId="0" pageNumber="1">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F12" captionText="Figure 10. Digestive system, dorsal view, Peronia peronii. The arrow indicates the orientation of the transitional loop A Mauritius [5874] (MNHN-IM- 2019 - 1605) B Papua New Guinea, Madang [5472] (MNHN-IM- 2013 - 14052) C Guam [5840] (MNHN-IM- 2019 - 1609). Scale bars: 10 mm (A-C). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure10" httpUri="https://binary.pensoft.net/fig/458310" pageId="0" pageNumber="1">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F13" captionText="Figure 11. Radula, Peronia peronii A-C Papua New Guinea [5476] (MNHN-IM- 2013 - 16260) D-H Guam [5840] (MNHN-IM- 2019 - 1609) A lateral teeth B rachidian and innermost lateral teeth C rachidian and innermost lateral teeth D rachidian and innermost lateral teeth E rachidian and lateral teeth F lateral teeth G lateral teeth, frontal view H outermost lateral teeth. Scale bars: 100 μm (A), 40 μm (B, D), 80 μm (C), 200 μm (E, F), 100 μm (G, H)." figureDoi="10.3897/zookeys.972.52853.figure11" httpUri="https://binary.pensoft.net/fig/458311" pageId="0" pageNumber="1">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F14" captionText="Figure 12. Radula, Peronia peronii, Mauritius [5872] (MNHN-IM- 2019 - 1605) A rachidian and innermost lateral teeth B rachidian and lateral teeth C outermost lateral teeth D lateral teeth. Scale bars: 40 μm (A), 100 μm (B), 20 μm (C), 200 μm (D)." figureDoi="10.3897/zookeys.972.52853.figure12" httpUri="https://binary.pensoft.net/fig/458312" pageId="0" pageNumber="1">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F15" captionText="Figure 13. Posterior, hermaphroditic (female) reproductive system, Peronia peronii A Mauritius [3605] (MNHN-IM- 2019 - 1606) B Papua New Guinea, Madang [5472] (MNHN-IM- 2013 - 14052). Scale bars: 10 mm (A), 5 mm (B). Abbreviations: dd deferent duct, fgm female gland mass, hg hermaphroditic gland, ov oviduct, rs receptaculum seminis, sp spermatheca." figureDoi="10.3897/zookeys.972.52853.figure13" httpUri="https://binary.pensoft.net/fig/458313" pageId="0" pageNumber="1">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F16" captionText="Figure 14. Anterior, male, copulatory apparatus, Peronia peronii A Mauritius [3605] (MNHN-IM- 2019 - 1606) B Papua New Guinea, Madang [5472] (MNHN-IM- 2013 - 14052). Scale bars: 10 mm (A), 5 mm (B). Abbreviations: ag accessory penial gland, dd deferent duct, ms muscular sac, ps penial sheath, rm retractor muscle, v vestibule." figureDoi="10.3897/zookeys.972.52853.figure14" httpUri="https://binary.pensoft.net/fig/458314" pageId="0" pageNumber="1">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F17" captionText="Figure 15. Penial hooks, Peronia peronii A Mauritius [5872] (MNHN-IM- 2019 - 1605) B Mauritius [3605] (MNHN-IM- 2019 - 1606) C, D Guam [5840] (MNHN-IM- 2019 - 1609). Scale bars: 6 μm (A, B, D), 40 μm (C)." figureDoi="10.3897/zookeys.972.52853.figure15" httpUri="https://binary.pensoft.net/fig/458315" pageId="0" pageNumber="1">, 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F18" captionText="Figure 16. Accessory penial gland spine, Peronia peronii A, B, F, G Mauritius C, D, H Papua New Guinea E, I Guam A [3605] (MNHN-IM- 2019 - 1606) B [5872] (MNHN-IM- 2019 - 1605) C [5471] (MNHN-IM- 2013 - 12500) D [5472] (MNHN-IM- 2013 - 14052) E [5840] (MNHN-IM- 2019 - 1609) F same as A; G same as B; H same as C; I same as E. Scale bars: 800 μm (A-D), 1 mm (E), 40 μm (F-I)." figureDoi="10.3897/zookeys.972.52853.figure16" httpUri="https://binary.pensoft.net/fig/458316" pageId="0" pageNumber="1">, 16</figureCitation>
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<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">Onchidium peronii</taxonomicName>
Cuvier, 1804: 37-51, pl. 6, figs 1-9;
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/1844622" refId="B44" refString="Cuvier, G, 1816. Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee (Vol. 2). Deterville, Paris, xviii + 532 pp.[according to Roux (1976), all four volumes were published by the end of December 1816, even although the year 1817 is printed on the title page] https://www.biodiversitylibrary.org/page/1844622" title="Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee (Vol. 2). Deterville, Paris, xviii + 532 pp. [according to Roux (1976), all four volumes were published by the end of December 1816, even although the year 1817 is printed on the title page]" url="https://www.biodiversitylibrary.org/page/1844622" year="1816">Cuvier 1816</bibRefCitation>
: 411;
<bibRefCitation author="Lamarck, JB Monet de" journalOrPublisher="Published by the Author, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/13181853" refId="B112" refString="Lamarck, JB Monet de, 1822. Histoire naturelle des animaux sans vertebres. Tome sixieme, 2eme partie. Suite des Gasteropodes. Published by the Author, Paris, https://www.biodiversitylibrary.org/page/13181853" title="Histoire naturelle des animaux sans vertebres. Tome sixieme, 2 eme partie. Suite des Gasteropodes." url="https://www.biodiversitylibrary.org/page/13181853" year="1822">Lamarck 1822</bibRefCitation>
: 46;
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/33372853" refId="B45" refString="Cuvier, G, 1830. Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX] https://www.biodiversitylibrary.org/page/33372853" title="Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX]" url="https://www.biodiversitylibrary.org/page/33372853" year="1830">Cuvier 1830</bibRefCitation>
: 46;
<bibRefCitation author="Voigt, FS" journalOrPublisher="Brodhaus, Leipzig" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/19175572" refId="B188" refString="Voigt, FS, 1834. Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken. Brodhaus, Leipzig, https://www.biodiversitylibrary.org/page/19175572" title="Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken." url="https://www.biodiversitylibrary.org/page/19175572" year="1834">Voigt 1834</bibRefCitation>
: 101;
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Deterville, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41057481" refId="B61" refString="Deshayes, GP, 1836-1845. Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1-139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot;Le regne animal illustre&quot; or the &quot;Disciples' edition&quot;] https://www.biodiversitylibrary.org/page/41057481" title="Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1 - 139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot; Le regne animal illustre &quot; or the &quot; Disciples' edition &quot;]" url="https://www.biodiversitylibrary.org/page/41057481" year="1836 - 1845">Deshayes 1836-1845</bibRefCitation>
: pl. 26, fig. 2;
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41706785" refId="B63" refString="Deshayes, GP, Milne-Edwards, H, 1836. Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J.B.P.A. de Lamarck. Deuxieme edition, Tome septieme. Histoire des Mollusques. J.-B. Bailliere, Paris, https://www.biodiversitylibrary.org/page/41706785" title="Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J. B. P. A. de Lamarck. Deuxieme edition, Tome septieme. Histoire des Mollusques. J. - B." url="https://www.biodiversitylibrary.org/page/41706785" year="1836">Deshayes and Milne-Edwards 1836</bibRefCitation>
: 709; JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: 117; ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: pl. 181, fig. 7;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge 1855</bibRefCitation>
: 124, pl. 16, fig. 8;
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate 1893</bibRefCitation>
: 172-173, pl. 12, figs 85, 87, 91;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.789" author="Odhner, NJ" journalOrPublisher="Arkiv foer Zoologi" pageId="0" pageNumber="1" pagination="1 - 52" refId="B136" refString="Odhner, NJ, 1919. Contribution a la faune malacologique de Madagascar. Arkiv foer Zoologi 12: 1 - 52, DOI: https://doi.org/10.5962/bhl.part.789" title="Contribution a la faune malacologique de Madagascar." url="https://doi.org/10.5962/bhl.part.789" volume="12" year="1919">Odhner 1919</bibRefCitation>
: 42;
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
: 44-45, 71-72 [in part only].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">Peronia peronii</taxonomicName>
(Cuvier, 1804):
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming 1822a</bibRefCitation>
: 574;
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">Fleming 1822b</bibRefCitation>
: 463;
<bibRefCitation author="Keferstein, WM" editor="Bronn, HG" journalOrPublisher="Bd. 3 (2). Winter, Leipzig &amp; Heidelberg" pageId="0" pageNumber="1" pagination="1160 - 1306" publicationUrl="https://www.biodiversitylibrary.org/page/35742549" refId="B102" refString="Keferstein, WM, 1865a. Lungenschnecken: Pulmonata. In: Bronn, HG, Ed., Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild. Bd. 3(2). Winter, Leipzig &amp; Heidelberg: 1160 - 1306, https://www.biodiversitylibrary.org/page/35742549" title="Lungenschnecken: Pulmonata." url="https://www.biodiversitylibrary.org/page/35742549" volumeTitle="Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild." year="1865 a">Keferstein 1865a</bibRefCitation>
: pl. CIII, fig. 1;
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
: 190-191 [in part only];
<bibRefCitation author="Marcus, Er" journalOrPublisher="Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse" pageId="0" pageNumber="1" pagination="873 - 934" refId="B122" refString="Marcus, Er, Marcus, Ev, 1960. Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse 12: 873 - 934" title="Opisthobranchia aus dem Roten Meer und von den Malediven." volume="12" year="1960">Marcus and Marcus 1960</bibRefCitation>
: 877;
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus 1970</bibRefCitation>
: 213 [in part only];
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. 2011</bibRefCitation>
: 428;
<bibRefCitation DOI="https://doi.org/10.1186/1471-2148-11-295" author="White, TR" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="1" pagination="1 - 295" refId="B196" refString="White, TR, Conrad, M, Tseng, R, Golding, R, Gomes, S, de Frias Martins, AM, Dayrat, B, 2011. Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications. BMC Evolutionary Biology 11: 1 - 295, DOI: https://doi.org/10.1186/1471-2148-11-295" title="Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications." url="https://doi.org/10.1186/1471-2148-11-295" volume="11" year="2011">White et al. 2011</bibRefCitation>
: 4.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchis peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">Onchis peronii</taxonomicName>
(Cuvier, 1804):
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
1822: xxxi.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">Peronia mauritiana</taxonomicName>
Blainville, 1824: 281; Adams and Adams 1855: 235.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">Onchidium tonganum</taxonomicName>
Quoy &amp; Gaimard, 1832: 210-211, pl. 15, figs 17, 18; Semper 1880: 258-260, pl. XIX, figs 2, 9, pl. XXII, figs 1, 2, 10 [in part only];
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh 1884a</bibRefCitation>
: 142-148, pl. VI, fig. 19, pl. VII, figs 1-6.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia tongana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongana">Peronia tongana</taxonomicName>
(Quoy &amp; Gaimard, 1832):
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/27509779" refId="B139" refString="Oken, L, 1834a. Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283-310. https://biodiversitylibrary.org/page/27509779" title="Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283 - 310." url="https://biodiversitylibrary.org/page/27509779" year="1834 a">Oken 1834a</bibRefCitation>
: 287; JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: 117; ME Gray, 1850: pl. 182, fig. 1, as
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="tongensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongensis">tongensis</taxonomicName>
; Adams and Adams 1855: 235, pl. LXXXI, fig. 3;
<bibRefCitation author="Keferstein, WM" editor="Bronn, HG" journalOrPublisher="Bd. 3 (2). Winter, Leipzig &amp; Heidelberg" pageId="0" pageNumber="1" pagination="1160 - 1306" publicationUrl="https://www.biodiversitylibrary.org/page/35742549" refId="B102" refString="Keferstein, WM, 1865a. Lungenschnecken: Pulmonata. In: Bronn, HG, Ed., Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild. Bd. 3(2). Winter, Leipzig &amp; Heidelberg: 1160 - 1306, https://www.biodiversitylibrary.org/page/35742549" title="Lungenschnecken: Pulmonata." url="https://www.biodiversitylibrary.org/page/35742549" volumeTitle="Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild." year="1865 a">Keferstein 1865a</bibRefCitation>
: pl. CII, fig. 20;
<bibRefCitation author="Tapparone Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="5 - 313" publicationUrl="https://www.biodiversitylibrary.org/page/10812598" refId="B183" refString="Tapparone Canefri, C, 1883. Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini. Annali del Museo Civico di Storia Naturale di Genova 19: 5 - 313, https://www.biodiversitylibrary.org/page/10812598" title="Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini." url="https://www.biodiversitylibrary.org/page/10812598" volume="19" year="1883">Tapparone Canefri 1883</bibRefCitation>
: 214 [in part only];
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
: 191-192, figs 4-7 [in part only].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">Onchidium punctatum</taxonomicName>
Quoy &amp; Gaimard, 1832: 215-216, pl. 15, figs 27, 28. Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">Peronia punctata</taxonomicName>
(Quoy &amp; Gaimard, 1832):
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/27509779" refId="B139" refString="Oken, L, 1834a. Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283-310. https://biodiversitylibrary.org/page/27509779" title="Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283 - 310." url="https://biodiversitylibrary.org/page/27509779" year="1834 a">Oken 1834a</bibRefCitation>
: 287; JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: 117; ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray 1850</bibRefCitation>
: pl. 183, fig. 3; Adams and Adams 1855: 235;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.119392" author="Chenu, JC" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="1" refId="B32" refString="Chenu, JC, 1859. Manuel de Conchyliologie et de Paleontologie Conchyliologique (Vol. 1). Masson, Paris, vii + 508 pp. https://doi.org/10.5962/bhl.title.119392" title="Manuel de Conchyliologie et de Paleontologie Conchyliologique (Vol. 1). Masson, Paris, vii + 508 pp." url="https://doi.org/10.5962/bhl.title.119392" year="1859">Chenu 1859</bibRefCitation>
: 474, fig. 3505;
<bibRefCitation author="Tapparone Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="5 - 313" publicationUrl="https://www.biodiversitylibrary.org/page/10812598" refId="B183" refString="Tapparone Canefri, C, 1883. Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini. Annali del Museo Civico di Storia Naturale di Genova 19: 5 - 313, https://www.biodiversitylibrary.org/page/10812598" title="Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini." url="https://www.biodiversitylibrary.org/page/10812598" volume="19" year="1883">Tapparone Canefri 1883</bibRefCitation>
: 214.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">Onchidium melanopneumon</taxonomicName>
Bergh, 1884a: 129-142, pl. IV, figs 25-27, pl. V, figs 1-27, pl. VI, figs 5-18, 20-21;
<bibRefCitation author="Joyeux-Laffuie, J" journalOrPublisher="Journal of the Society for the Bibliography of Natural History" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/44109240" refId="B100" refString="Joyeux-Laffuie, J, 1885. Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884b)] https://www.biodiversitylibrary.org/page/44109240" title="Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884 b)]" url="https://www.biodiversitylibrary.org/page/44109240" year="1885">Joyeux-Laffuie 1885</bibRefCitation>
: viii-xi.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">Paraperonia fidjiensis</taxonomicName>
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, 1934a: 197-198, figs 9-11. Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">Peronia verruculata</taxonomicName>
:
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
1872a
</bibRefCitation>
: 28;
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
1872b
</bibRefCitation>
: 325, as
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="vermiculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vermiculata">vermiculata</taxonomicName>
[non
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">Peronia verruculata</taxonomicName>
(Cuvier, 1830)].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="1">paralectotype</emphasis>
</emphasis>
(
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
). Mauritius • lectotype, hereby designated, by means of
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) anatomical drawings. Timor • 1 paralectotype, 60/40 mm; F
<normalizedToken originalValue="Péron">Peron</normalizedToken>
leg.; MNHN-IM-2000-22938. The fact that the specimen illustrated by Cuvier cannot be located does not invalidate the lectotype designation (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 74.4). That individual, according to Cuvier, measured approximately 140 mm long (preserved).
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) detailed anatomical drawings are exclusively based on the individual collected by
<normalizedToken originalValue="Péron">Peron</normalizedToken>
in Mauritius. Note that, although
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) illustrations are truly remarkable, they are flipped at 180° because, for instance, the heart and the male anterior parts are on the left. Something must have happened during the engraving or the printing.
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 71) referred to Mauritius as the
<normalizedToken originalValue="“Typ-Lokalität”">&quot;Typ-Lokalitaet&quot;</normalizedToken>
of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
but did not formally designate a lectotype for
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. In case of syntypes, &quot;the place of origin of the lectotype becomes the type locality of the nominal species-group taxon, despite any previously published statement of the type locality.&quot; (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 76.2)
</paragraph>
<paragraph pageId="0" pageNumber="1">
The original description of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
was based on two specimens collected by
<normalizedToken originalValue="Péron">Peron</normalizedToken>
: the lectotype from Mauritius, of which the internal anatomy was illustrated in detail by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6), could not be located and is likely lost; the paralectotype from Timor (MNHN-IM-2000-22938) was very briefly mentioned by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: 39) who merely wrote that another specimen was brought from Timor by
<normalizedToken originalValue="Péron">Peron</normalizedToken>
and that
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
is present &quot;at the two extreme ends of the Indian Ocean.&quot; The paralectotype (60/40 mm) is well preserved even though dorsal papillae with eyes cannot be counted because their color faded. It is obvious that Cuvier did not actually use it for his detailed anatomical description and illustrations on plate 6, because it was never opened prior to the present study, except for a tiny cut near the lung. It was carefully opened on its side to draw a dorsal view of its intestinal loops of type I (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9A</figureCitation>
) and measure the length (4.5 mm) of the spine of the accessory penial gland (by transparency, so that the male copulatory apparatus was not dissected).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
(
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
). Mauritius • lectotype, hereby designated, by means of
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) anatomical drawings. The species name
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation author="Blainville, HMD de" journalOrPublisher="Levrault, Strasbourg" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/25299228" refId="B23" refString="Blainville, HMD de, 1824. Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G. Levrault, Strasbourg, https://www.biodiversitylibrary.org/page/25299228" title="Mollusques. In: Cuvier F (Ed) Dictionnaire des Sciences Naturelles (Vol. 32). F. G." url="https://www.biodiversitylibrary.org/page/25299228" year="1824">Blainville (1824</bibRefCitation>
: 281) for a species originally illustrated by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6) in the
<emphasis italics="true" pageId="0" pageNumber="1">
Annales du
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
<normalizedToken originalValue="dHistoire">d'Histoire</normalizedToken>
naturelle
</emphasis>
and which Cuvier named
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
.
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
reference to
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804)
</bibRefCitation>
plate 6 (&quot;La
<normalizedToken originalValue="Péronie">Peronie</normalizedToken>
de
<normalizedToken originalValue="lIsle-de-France">l'Isle-de-France</normalizedToken>
[Mauritius].
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
. Blainv., Cuv., Ann. du Mus., 5, pl. 6.&quot;) serves as an indication, and
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
is an available binomen (
<bibRefCitation author="ICZN" journalOrPublisher="Bulletin of Zoological Nomenclature" pageId="0" pageNumber="1" refId="B97" refString="ICZN, 1999. International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." title="International code of zoological nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature, 306 pp." year="1999">ICZN 1999</bibRefCitation>
: Article 12.2.1). However,
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
original description of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
was based on two specimens, a lectotype from Mauritius and a paralectotype from Timor, but
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) plate of anatomical drawings exclusively illustrates the lectotype from Mauritius (see above). Because the lectotype of
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
also is the lectotype of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
remains what it always was, i.e., a junior objective synonym of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Blainville also mentioned the name
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
in his
<emphasis italics="true" pageId="0" pageNumber="1">Manuel de Malacologie et de Conchyliologie</emphasis>
(Blainville 1825: 490) and in the article
<normalizedToken originalValue="“Péronie”">&quot;Peronie&quot;</normalizedToken>
of the
<emphasis italics="true" pageId="0" pageNumber="1">Dictionnaire des Sciences Naturelles</emphasis>
(
<bibRefCitation author="Blainville, HMD de" editor="Cuvier, F" journalOrPublisher="38). F. G. Levrault, Strasbourg" pageId="0" pageNumber="1" pagination="519 - 524" publicationUrl="https://www.biodiversitylibrary.org/page/25302418" refId="B24" refString="Blainville, HMD de, 1826. Peronie. In: Cuvier, F, Ed., Dictionnaire des sciences naturelles (Vol. 38). F. G. Levrault, Strasbourg: 519 - 524, https://www.biodiversitylibrary.org/page/25302418" title="Peronie." url="https://www.biodiversitylibrary.org/page/25302418" volumeTitle="Dictionnaire des sciences naturelles (Vol." year="1826">Blainville 1826</bibRefCitation>
: 523). The illustration published by Blainville (1827: pl. 46, fig. 7) in the
<emphasis italics="true" pageId="0" pageNumber="1">Atlas</emphasis>
of the
<emphasis italics="true" pageId="0" pageNumber="1">Manuel</emphasis>
differs from that published by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6, fig. 1). The specimen used by Blainville for that illustration could not be located, which does not matter much since it does not have any name-bearing function. However, it also means that, because there are two species of
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
in Mauritius,
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
(1827: pl. 46, fig. 7) illustration may or may not refer to
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
(
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
). Tonga • lectotype, hereby designated, 100/60 mm; Panhi-Motou [possibly the small island of Pangaimotu]; MNHN-IM-2000-22937. It is unclear how many specimens Quoy and Gaimard (1832: 210-211, pl. 15, figs 17, 18) examined for the original description of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
. They may have examined more than one individual. Regardless, it is clear that
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
applies to a
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species because the notum of the lectotype bears gills which were also illustrated in the original description. Its notum also bears fifteen dorsal papillae with eyes but others probably faded. The lectotype was dissected prior to the present study. The accessory penial gland and the penial apparatus are missing (pieces of the deferent duct remain). The intestinal loops are of type I with a transitional loop between 2 and 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9B</figureCitation>
). Quoy and Gaimard (1832: 216) briefly mentioned the presence of
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
in Manokwari, West Papua, Indonesia, but that record could not be confirmed (although
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is known to be present there because Manokwari is the type locality of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="1">paralectotypes</emphasis>
</emphasis>
(
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
). Indonesia • lectotype, hereby designated, 70/60 mm; dans le port de Dorey [Manokwari harbor, West Papua]; 1829; JRC Quoy and JP Gaimard leg.; MNHN-IM-2000-22966. • 2 paralectotypes, 35/25 and 32/30 mm; same collection data as for the lectotype; MNHN-IM-2000-33701. An old label of the lectotype says &quot;
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">Onchidium punctatum</taxonomicName>
, Q. G, Ast. pl. 15, fig. 27, de la Nouvelle
<normalizedToken originalValue="Guinée">Guinee</normalizedToken>
, Quoy et Gaimard 1829.&quot; That old label does not say
<normalizedToken originalValue="“Dorey”">&quot;Dorey&quot;</normalizedToken>
(for the locality), which is only mentioned in the original description, but it clearly indicates that the lectotype was part of the type series of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
. The lectotype bears dorsal gills, as illustrated by Quoy and Gaimard (1832: pl. 15, figs 27, 28). It was dissected prior to the present study, likely by
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 203-204) and its penis is missing but its intestinal loops are of type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9C</figureCitation>
). Its spine of the accessory penial gland, still in place in the animal, is 3.7 mm long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
A second jar was found with two paralectotypes (MNHN-IM-2000-33701). An old label for that second jar says &quot;
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
<normalizedToken originalValue="piquetée">piquetee</normalizedToken>
, Q G. MM Quoy Gaimard, 1829&quot; with no locality data. The name &quot;
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
&quot; was added on the label. The number
<normalizedToken originalValue="“51”">&quot;51&quot;</normalizedToken>
also appears on another old label, which corresponds to an unknown numbering system. There also is a more recent label saying &quot;
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia picta" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="picta">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia picta</emphasis>
</taxonomicName>
QG, M. Quoy et Gaimard, 1829.&quot; Quoy and Gaimard did not describe any onchidiid species with the specific name
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="picta" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="picta">
<emphasis italics="true" pageId="0" pageNumber="1">picta</emphasis>
</taxonomicName>
. However, the French vernacular name of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
in Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832: 215) original description is &quot;
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Onchidium</taxonomicName>
<normalizedToken originalValue="piquetée.”">piquetee.&quot;</normalizedToken>
So, it is likely that these two additional specimens were part of the type series of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
. Both paralectotypes (35/25 and 32/30 mm) bear dorsal gills. The largest paralectotype was dissected prior to the present study, possibly by
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 203, 204), and its penis is missing but its accessory penial gland remains. The small paralectotype was not dissected.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 203) listed three individuals from Port-Dorey which he (implicitly) regarded as part of the original series of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
.
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
gave the measurements for only two individuals: &quot;
<emphasis italics="true" pageId="0" pageNumber="1">a</emphasis>
&quot; (35/25 mm), likely the largest paralectotype; &quot;
<emphasis italics="true" pageId="0" pageNumber="1">b</emphasis>
&quot; (77/56 mm), likely the lectotype. In addition, in his re-description of
<taxonomicName baseAuthorityName="Quoy &amp; Gaimard" baseAuthorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis punctata</emphasis>
</taxonomicName>
,
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 204-205) mentioned two individuals identified as
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
and collected by Quoy and Gaimard in 1829, from an unknown locality. Those two individuals are likely within another jar found at the MNHN with the old number
<normalizedToken originalValue="“48”">&quot;48&quot;</normalizedToken>
and a label saying &quot;
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
M. Quoy et Gaimard 1829.&quot; There is no reason to consider that those two unidentified individuals from the collection of Quoy and Gaimard were part of the type series of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
. Finally, there is no other old material at the MNHN which could be assigned to the type series of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
. There are only three other old specimens from Port Dorey at the MNHN: the two syntypes (MNHN-IM-2000-22950) of
<taxonomicName baseAuthorityName="Lesson" baseAuthorityYear="1831" class="Gastropoda" family="Onchidiidae" genus="Wallaconchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wallaconchis ater" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ater">
<emphasis italics="true" pageId="0" pageNumber="1">Wallaconchis ater</emphasis>
</taxonomicName>
(Goulding et al. 2018: 63), and one specimen collected by Raffray in 1878 (with numbers
<normalizedToken originalValue="“22”">&quot;22&quot;</normalizedToken>
and
<normalizedToken originalValue="“75”">&quot;75&quot;</normalizedToken>
on the label).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
(
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium melanopneumon</emphasis>
</taxonomicName>
). Fiji • holotype, by monotypy, 65/40 mm; Kandavu [Kadavu]; Aug 1874; HMS Challenger leg.; NHMUK 1888.5.30.39. The holotype was entirely dissected by Bergh and is now empty. Given the presence of dorsal gills,
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium melanopneumon</emphasis>
</taxonomicName>
clearly applies to a
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="1">paralectotypes</emphasis>
</emphasis>
(
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
). Fiji • lectotype, hereby designated, 60/50 mm; 1876; Filhol leg.; MNHN-IM-2000-33692. No jar clearly labeled as the type material of
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
was found at the MNHN, but the lectotype could be traced, and six paralectotypes could not be found at the MNHN.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 197-198, figs 9-11) described
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
based on seven individuals from Fiji (&quot;Iles Fidji&quot;) collected by Filhol (Henri Filhol [1843-1902]) in 1876 and with the following sizes: 75/50 mm for six &quot;
<emphasis italics="true" pageId="0" pageNumber="1">a</emphasis>
&quot; individuals and 70/50 mm for a seventh &quot;
<emphasis italics="true" pageId="0" pageNumber="1">b</emphasis>
&quot; individual. Two jars of material collected in Fiji by Filhol in 1876 were found at the MNHN. The first jar, labeled as &quot;
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Peronia</taxonomicName>
[written over
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Oncidium</taxonomicName>
] I. Fidji Mr. Filhol n°11 1876&quot; and
<normalizedToken originalValue="“71,”">&quot;71,&quot;</normalizedToken>
contains a single
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
specimen which, given its size (60/50 mm), very likely is part of the type series of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
, and which is designated as the lectotype (MNHN-IM-2000-33692). Its radula and all reproductive parts are missing. Its intestinal loops are clearly of type I, with a transitional loop at ~ 1
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9E</figureCitation>
). The second jar, labeled as &quot;
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Oncidiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidiella" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Oncidiella</taxonomicName>
I. Fidji Mr. Filhol n°11 1876&quot; and
<normalizedToken originalValue="“101,”">&quot;101,&quot;</normalizedToken>
contains four poorly-preserved specimens which do not even appear to belong to
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
, with a size (less than 30 mm) not compatible with the original description of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
, and which, therefore, cannot be regarded as part of the type series.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
Mauritius • 2 specimens 140/100 mm [5872] and 125/75 mm [5874]; La Mivoie;
<geoCoordinate degrees="20" direction="south" minutes="20.659" orientation="latitude" precision="1" value="-20.344316">20°20.659'S</geoCoordinate>
,
<geoCoordinate degrees="57" direction="east" minutes="21.763" orientation="longitude" precision="1" value="57.362717">57°21.763'E</geoCoordinate>
; 11 Jun 2014; TC Goulding leg.; st 177, basalt rocks, at night; MNHN-IM-2019-1605. • 1 specimen 110/100 mm [3605]; Mahebourg, waterfront;
<geoCoordinate degrees="20" direction="south" minutes="24.317" orientation="latitude" precision="1" value="-20.405283">20°24.317'S</geoCoordinate>
,
<geoCoordinate degrees="57" direction="east" minutes="42.605" orientation="longitude" precision="1" value="57.710083">57°42.605'E</geoCoordinate>
; 13 Jun 2014; TC Goulding leg.; st 178, rocky intertidal, with algae, just before sunrise; MNHN-IM-2019-1606. • 1 specimen 100/90 mm [1553]; Grand Port, east side of
<normalizedToken originalValue="île">ile</normalizedToken>
Marianne;
<geoCoordinate degrees="20" direction="south" minutes="22.828" orientation="latitude" precision="1" value="-20.380466">20°22.828'S</geoCoordinate>
,
<geoCoordinate degrees="57" direction="east" minutes="47.220" orientation="longitude" precision="1" value="57.787">57°47.220'E</geoCoordinate>
; May 2003; O Griffiths leg.; A2518, out of water on limestone platform; MNHN-IM-2019-1607.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Mariana Islands • 1 specimen 115/80 mm [443]; Guam Island, Bile Bay;
<geoCoordinate degrees="13" direction="north" minutes="17.124" orientation="latitude" precision="1" value="13.2854">13°17.124'N</geoCoordinate>
,
<geoCoordinate degrees="144" direction="east" minutes="39.742" orientation="longitude" precision="1" value="144.66237">144°39.742'E</geoCoordinate>
; 23 Mar 2007; C Carlson leg.; reef margin; CASIZ 180486. • 1 specimen 85/70 mm [5840]; Guam Island, Bile Bay;
<geoCoordinate degrees="13" direction="north" minutes="16.582" orientation="latitude" precision="1" value="13.276366">13°16.582'N</geoCoordinate>
,
<geoCoordinate degrees="144" direction="east" minutes="39.752" orientation="longitude" precision="1" value="144.66254">144°39.752'E</geoCoordinate>
; 27 Nov 2007; C Carlson leg.; shoreline; MNHN-IM-2019-1609.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Papua New Guinea -
<emphasis bold="true" pageId="0" pageNumber="1">Madang</emphasis>
• 1 specimen 70/60 mm [5476]; Wonad Island;
<geoCoordinate degrees="05" direction="south" minutes="08.1" orientation="latitude" precision="92" value="-5.135">05°08.1'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="49.3" orientation="longitude" precision="92" value="145.82167">145°49.3'E</geoCoordinate>
; 29 Nov 2012; MNHN Expedition Papua Niugini leg.; st PM43, night tide, sandy beach and intertidal rocks; MNHN-IM-2013-16260. • 1 specimen 65/45 mm [5477]; Wonad Island;
<geoCoordinate degrees="05" direction="south" minutes="08.1" orientation="latitude" precision="92" value="-5.135">05°08.1'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="49.3" orientation="longitude" precision="92" value="145.82167">145°49.3'E</geoCoordinate>
; 27 Nov &amp; 09 Dec 2012; MNHN Expedition Papua Niugini leg.; st PM41, sandy beach and intertidal rocks; MNHN-IM-2013-15872. • 1 specimen 55/40 mm [5474]; Rempi Area, Barag Island;
<geoCoordinate degrees="05" direction="south" minutes="01.1" orientation="latitude" precision="92" value="-5.0183334">05°01.1'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="47.9" orientation="longitude" precision="92" value="145.79834">145°47.9'E</geoCoordinate>
; 15 Nov 2012; MNHN Expedition Papua Niugini leg.; st PM25, fringing reef on narrow barrier island; MNHN-IM-2013-14054. • 1 specimen 80/60 mm [5472]; same collection data as for the preceding; MNHN-IM-2013-14052. • 1 specimen 80/70 mm [5471]; Rempi Area, South Dumduman Island;
<geoCoordinate degrees="05" direction="south" minutes="00.2" orientation="latitude" precision="92" value="-5.0033336">05°00.2'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="47.6" orientation="longitude" precision="92" value="145.79333">145°47.6'E</geoCoordinate>
; 9 Nov 2012; MNHN Expedition Papua Niugini leg.; st PM 12, limestone rocky intertidal; MNHN-IM-2013-12500. -
<emphasis bold="true" pageId="0" pageNumber="1">New Ireland</emphasis>
• 1 specimen 50/40 mm [6086]; Kavieng, Lemus Island;
<geoCoordinate degrees="02" direction="south" minutes="38" orientation="latitude" precision="925" value="-2.6333332">02°38'S</geoCoordinate>
,
<geoCoordinate degrees="150" direction="east" minutes="37.5" orientation="longitude" precision="92" value="150.625">150°37.5'E</geoCoordinate>
; 12-14 Jun 2014; MNHN Expedition Kavieng 2014 leg.; st KM24, mixed platform with seagrass; MNHN-IM-2013-53482.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(historical museum collections).</emphasis>
Chagos Archipelago • 1 specimen 95/65 mm; Ye Ye, Peros Banhos atoll; 24 Feb 1996; M Spalding (from N
<normalizedToken originalValue="Yonows">Yonow's</normalizedToken>
personal collection) leg.; exposed on shallow reef flat on rocks; MNHN-IM-2014-7992.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Fiji • 1 specimen 75/50 mm; Viti Isles; A Garrett leg.; ANSP 57967. • 1 specimen 28/25 mm; Viti Levu, Namuka;
<geoCoordinate degrees="18" direction="south" minutes="08" orientation="latitude" precision="925" value="-18.133333">18°08'S</geoCoordinate>
,
<geoCoordinate degrees="177" direction="east" minutes="23" orientation="longitude" precision="925" value="177.38333">177°23'E</geoCoordinate>
; 18 Apr 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; barrier reef; SMNH 180357. • 2 specimens 23/20 mm and 15/15 mm; Viti Levu, SW Suva, Namuka;
<geoCoordinate degrees="18" direction="south" minutes="08" orientation="latitude" precision="925" value="-18.133333">18°08'S</geoCoordinate>
,
<geoCoordinate degrees="177" direction="east" minutes="23" orientation="longitude" precision="925" value="177.38333">177°23'E</geoCoordinate>
; 16 Jun 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; barrier reef; SMNH 180374. • 1 specimen 37/30 mm; Viti Levu, Namuka;
<geoCoordinate degrees="18" direction="south" minutes="08" orientation="latitude" precision="925" value="-18.133333">18°08'S</geoCoordinate>
,
<geoCoordinate degrees="177" direction="east" minutes="23" orientation="longitude" precision="925" value="177.38333">177°23'E</geoCoordinate>
; 19 Jun 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; SMNH 180375. • 1 specimen 80/65 mm; Viti Levu, Bau Island;
<geoCoordinate degrees="17" direction="south" minutes="58" orientation="latitude" precision="925" value="-17.966667">17°58'S</geoCoordinate>
,
<geoCoordinate degrees="178" direction="east" minutes="36" orientation="longitude" precision="925" value="178.6">178°36'E</geoCoordinate>
; 2 Jul 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; reef; SMNH 180373.
</paragraph>
<paragraph pageId="0" pageNumber="1">India • 2 specimens 85/55 mm and 70/50 mm; Nicobar Islands, Pulo Milo, Little Nicobar; Reinhardt, Galathea 305 leg.; NHMD 613753.</paragraph>
<paragraph pageId="0" pageNumber="1">
Indonesia -
<emphasis bold="true" pageId="0" pageNumber="1">Java</emphasis>
• 1 specimen 90/55 mm; Batavia [Jakarta]; 1899; C Aurivillius leg.; SMNH 180355. -
<emphasis bold="true" pageId="0" pageNumber="1">Sumatra</emphasis>
• 1 specimen 100/70 mm; Sumatra; Deshayes leg.; MNHN-IM-2012-25150. • 1 specimen 65/50 mm; west coast of Sumatra, Pulo Pasu [or Pulu Pasu]; 1891; C Aurivillius leg.; SMNH 180354. -
<emphasis bold="true" pageId="0" pageNumber="1">Tanimbar</emphasis>
• 1 specimen 60/50 mm; Jamdena Straits, West side of Mitak Island;
<geoCoordinate degrees="07" direction="south" minutes="11" orientation="latitude" precision="925" value="-7.1833334">07°11'S</geoCoordinate>
,
<geoCoordinate degrees="131" direction="east" minutes="28" orientation="longitude" precision="925" value="131.46666">131°28'E</geoCoordinate>
; 22 Jun 1970; Mariel King Memorial Expedition Moluccas MV
<normalizedToken originalValue="“Pele”">&quot;Pele&quot;</normalizedToken>
1970 leg.; WAM S26723.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Kiribati • 1 specimen 35/35 mm; Gilbert Islands, Apaiang [Abaiang];
<geoCoordinate degrees="01" direction="north" minutes="49" orientation="latitude" precision="925" value="1.8166666">01°49'N</geoCoordinate>
,
<geoCoordinate degrees="172" direction="east" minutes="57" orientation="longitude" precision="925" value="172.95">172°57'E</geoCoordinate>
; 12 Aug 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef; SMNH 180353. • 1 specimen 70/65 mm; Gilbert Islands, Tarawa;
<geoCoordinate degrees="01" direction="north" minutes="26" orientation="latitude" precision="925" value="1.4333334">01°26'N</geoCoordinate>
, 173E; 16-20 Aug 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; reef; SMNH 180382. • 1 specimen 65/50 mm; Gilbert Islands, Aranuka; 00N, 174E; 6 Oct 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; reef; SMNH 180376. • 1 specimen 70/50 mm; Gilbert Islands, Aranuka; 00N, 174E; 22 Oct 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef; SMNH 180377. • 1 specimen 30/25 mm; Gilbert Islands, Aranuka; 00N, 174E; 1 Nov 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; SMNH 180378. • 1 specimen 15/15 mm; Gilbert Islands, Aranuka; 00N, 174E; 26 Oct 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef; SMNH 180383. • 2 specimens 20/15 mm and 17/13 mm; Gilbert Islands, Aranuka; 00N, 174E; 26 Oct 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef east; SMNH 180384. • 1 specimen 15/12 mm; Gilbert Islands, Aranuka;
<geoCoordinate degrees="00" direction="north" minutes="09" orientation="latitude" precision="925" value="0.15">00°09'N</geoCoordinate>
,
<geoCoordinate degrees="173" direction="east" minutes="35" orientation="longitude" precision="925" value="173.58333">173°35'E</geoCoordinate>
; 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef; SMNH 180478. • 1 specimen 65/50 mm; Gilbert Islands; Oct 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; outer reef; SMNH 180475. • 1 specimen 80/65 mm; Gilbert Islands, Apamama [Abemama];
<geoCoordinate degrees="00" direction="north" minutes="24" orientation="latitude" precision="925" value="0.4">00°24'N</geoCoordinate>
,
<geoCoordinate degrees="173" direction="east" minutes="55" orientation="longitude" precision="925" value="173.91667">173°55'E</geoCoordinate>
; 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; entrance reef; SMNH 180380. • 1 specimen 45/35 mm; Gilbert Islands, Apamama [Abemama]; 00N, 173E; 1917; S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; at low tide; SMNH 180379.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Madagascar • 1 specimen 65/50 mm; Tulear [Toliara];
<geoCoordinate degrees="23" direction="south" minutes="22" orientation="latitude" precision="925" value="-23.366667">23°22'S</geoCoordinate>
,
<geoCoordinate degrees="43" direction="east" minutes="39" orientation="longitude" precision="925" value="43.65">43°39'E</geoCoordinate>
; Feb 1913; K Afzelius leg.; coral reef; SMNH 180381.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Maldive Islands • 1 specimen 85/55 mm; Tiladummati Atoll, Faro Islet, on reef NW of Fildau Island;
<geoCoordinate degrees="06" direction="south" minutes="55.333" orientation="latitude" precision="1" value="-6.9222164">06°55.333'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="east" minutes="11.833" orientation="longitude" precision="1" value="73.19722">73°11.833'E</geoCoordinate>
; 30 &amp; 31 Mar 1964; R Robertson, International Indian Ocean Expedition leg.; st R021, intertidal, on dead coral rubble; ANSP 304860.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Marshall Islands • 1 specimen 37/30 mm; Jaluit; 06N, 170E; 20 Oct 1917; C Hessle, S
<normalizedToken originalValue="Bocks">Bock's</normalizedToken>
Pacific Expedition 1917-1918 leg.; west shore southeast of entrance; SMNH 180356.
</paragraph>
<paragraph pageId="0" pageNumber="1">Mauritius • 10 specimens up to 90/60 mm; probably Mauritius according to a new label (the original label was destroyed); 1929-1930; T Mortensen leg.; NHMD 613752.</paragraph>
<paragraph pageId="0" pageNumber="1">
New Caledonia • 1 specimen 100/60 mm; Touho, NW,
<normalizedToken originalValue="Koë">Koe</normalizedToken>
Reef, 2 mi. SSE; 16-20 Jan 1961; Kline &amp; Orr leg.; 0-4 feet, live and dead coral, sand, weed; ANSP 270221.
</paragraph>
<paragraph pageId="0" pageNumber="1">Palau • 1 specimen 80/65 mm; ANSP 203028.</paragraph>
<paragraph pageId="0" pageNumber="1">Seychelles • 1 specimen 90/70 mm; 1830; Dussumier leg.; MNHN-IM-2012-25149. • 1 specimen 85/60 mm; 1841; L. Rousseau leg.; MNHN-IM-2012-25148.</paragraph>
<paragraph pageId="0" pageNumber="1">
Tanzania • 1 specimen 65/50 mm; Zanzibar; 1902; C Eliot leg.; ANSP 84336. • 1 specimen 80/65 mm; west coast; Jun 1995; M Richmond &amp; M Toni (from N
<normalizedToken originalValue="Yonows">Yonow's</normalizedToken>
personal collection) leg.; sheltered, on limestone rock, intertidal exposed at low tide, common at night; MNHN-IM-2014-7991.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="genbank sequence">
<paragraph pageId="0" pageNumber="1">GenBank sequence.</paragraph>
<paragraph pageId="0" pageNumber="1">
One COI sequence was obtained from GenBank (LC390402) for an individual identified as
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
sp. and collected from Okinawa, Japan (
<bibRefCitation author="Takagi, M" journalOrPublisher="Part Second, The Birds. John Murray, London" pageId="0" pageNumber="1" publicationUrl="http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" refId="B181" refString="Takagi, M, Takao, Y, Mizuno, K, Ieyama, H, 2019. Genetic diversity of Onchidiidae in Japan Fauna Ryukyuana 49: 23-37. http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" title="Genetic diversity of Onchidiidae in Japan Fauna Ryukyuana 49: 23 - 37." url="http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" year="2019">Takagi et al. 2019</bibRefCitation>
), which is the northernmost confirmed locality for
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). Given that our fresh molecular samples of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
from the West Pacific (Guam, Papua New Guinea) are conspecific with those from Mauritius, it is assumed here that all individuals with a long spine of the accessory penial gland belong to the same species. Strictly speaking, however, the presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
from places like Zanzibar, the Maldives, Nicobar Islands, West Papua, Timor, Palau, New Caledonia, and Tonga, would still need to be validated with fresh material.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Interestingly, but for unclear reasons,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
seems to be only recorded from relatively small islands, the largest ones being Timor, New Caledonia, and Fiji. Even in Papua New Guinea, it was found on small islands close to the mainland but not on the mainland.
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
seems to be transported across vast distances from the western Pacific Ocean to the western Indian Ocean, but which does not seem to settle on the coasts of large land masses. We did not find it in any of the many localities we visited in the Philippines, Vietnam, Malaysia, Borneo, Sulawesi, Halmahera, Sumatra, etc. It is possible that we occasionally missed it in a few places (obviously we missed it in Timor and New Caledonia where it is present), but it is unlikely that we missed it everywhere.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is confirmed in the following locations (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
): Chagos Archipelago (new record); Fiji (type locality of
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
; Hoffmann, 1928; present study); India, Nicobar Islands (
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
1872a
</bibRefCitation>
,
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">b</bibRefCitation>
: 325, as
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
; Bergh, 1884a, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
; Hoffmann, 1928; present study); Indonesia, Java (
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
; present study), Sumatra (
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
;
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
; present study), Tanimbar (new record), Timor (
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier 1804</bibRefCitation>
, paralectotype of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
), West Papua (type locality of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
Quoy &amp; Gaimard, 1832); Japan, Okinawa (
<bibRefCitation author="Takagi, M" journalOrPublisher="Part Second, The Birds. John Murray, London" pageId="0" pageNumber="1" publicationUrl="http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" refId="B181" refString="Takagi, M, Takao, Y, Mizuno, K, Ieyama, H, 2019. Genetic diversity of Onchidiidae in Japan Fauna Ryukyuana 49: 23-37. http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" title="Genetic diversity of Onchidiidae in Japan Fauna Ryukyuana 49: 23 - 37." url="http://w3.u-ryukyu.ac.jp/naruse/lab/2019E_files/49-3_Takagi_etal.pdf" year="2019">Takagi et al. 2019</bibRefCitation>
, as onchidiids of &quot;Group II&quot;; new record); Kiribati, Gilbert Islands (
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
; present study); Madagascar (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.789" author="Odhner, NJ" journalOrPublisher="Arkiv foer Zoologi" pageId="0" pageNumber="1" pagination="1 - 52" refId="B136" refString="Odhner, NJ, 1919. Contribution a la faune malacologique de Madagascar. Arkiv foer Zoologi 12: 1 - 52, DOI: https://doi.org/10.5962/bhl.part.789" title="Contribution a la faune malacologique de Madagascar." url="https://doi.org/10.5962/bhl.part.789" volume="12" year="1919">Odhner 1919</bibRefCitation>
; present study); Maldive Islands (
<bibRefCitation author="Marcus, Er" journalOrPublisher="Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse" pageId="0" pageNumber="1" pagination="873 - 934" refId="B122" refString="Marcus, Er, Marcus, Ev, 1960. Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse 12: 873 - 934" title="Opisthobranchia aus dem Roten Meer und von den Malediven." volume="12" year="1960">Marcus and Marcus 1960</bibRefCitation>
,
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">1970</bibRefCitation>
; present study); Mariana Islands, Guam (
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1186/1471-2148-11-295" author="White, TR" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="1" pagination="1 - 295" refId="B196" refString="White, TR, Conrad, M, Tseng, R, Golding, R, Gomes, S, de Frias Martins, AM, Dayrat, B, 2011. Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications. BMC Evolutionary Biology 11: 1 - 295, DOI: https://doi.org/10.1186/1471-2148-11-295" title="Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications." url="https://doi.org/10.1186/1471-2148-11-295" volume="11" year="2011">White et al. 2011</bibRefCitation>
; present study); Marshall Islands (
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
; present study); Mauritius (type locality of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
; Semper 1880;
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh 1884a</bibRefCitation>
, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
;
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate 1893</bibRefCitation>
; present study); New Caledonia (new record); Palau (new record); Papua New Guinea, Madang (new record), New Ireland (new record); Seychelles (
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
; present study); Tanzania, Zanzibar (new record); and Tonga (type locality of
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
Quoy &amp; Gaimard, 1832). The most western records of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
are Zanzibar and southwestern Madagascar; its most eastern records are Okinawa, Guam, Kiribati, and Tonga. Note that
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is most likely also present in Tokara Islands (
<bibRefCitation DOI="https://doi.org/10.2307/2520262" author="Baba, K" journalOrPublisher="Zoological Memoirs, University of Bombay" pageId="0" pageNumber="1" refId="B12" refString="Baba, K, 1958. Seashore life of Japan [in Japanese]. Hokuryukan, Tokyo, 148 pp. [5 pls] https://doi.org/10.2307/2520262" title="Seashore life of Japan [in Japanese]. Hokuryukan, Tokyo, 148 pp. [5 pls]" url="https://doi.org/10.2307/2520262" year="1958">Baba 1958</bibRefCitation>
: 144, as
<taxonomicName lsidName="O. verruculatum" pageId="0" pageNumber="1" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. verruculatum</emphasis>
</taxonomicName>
), just south Kyushu, ca. 30N, which would be its most northern record.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The following records from the literature are not confirmed here, because authors did not provide enough information supporting the identification: Djibouti (
<bibRefCitation author="Vayssiere, A" journalOrPublisher="Annales de la Faculte des Sciences de l'Universite de Marseille, Supplement" pageId="0" pageNumber="1" pagination="5 - 157" publicationUrl="https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" refId="B187" refString="Vayssiere, A, 1912. Recherches zoologiques et anatomiques sur les Opisthobranches de la Mer Rouge et du Golfe d'Aden. Deuxieme Partie. Annales de la Faculte des Sciences de l'Universite de Marseille, Supplement 20: 5 - 157, https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" title="Recherches zoologiques et anatomiques sur les Opisthobranches de la Mer Rouge et du Golfe d'Aden. Deuxieme Partie." url="https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" volume="20" year="1912">
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
1912
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" author="O'Donoghue, CH" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" pagination="713 - 841" refId="B137" refString="O'Donoghue, CH, 1929. Report on the Opisthobranchiata. Zoological Results of the Cambridge Expedition to the Suez Canal. Transactions of the Zoological Society of London 22: 713 - 841, DOI: https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" title="Report on the Opisthobranchiata. Zoological Results of the Cambridge Expedition to the Suez Canal." url="https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" volume="22" year="1929">
<normalizedToken originalValue="ODonoghue">O'Donoghue</normalizedToken>
1929
</bibRefCitation>
;
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
); India, Nicobar (
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Annual Reviews of Ecology, Evolution and Systematics" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/30983429" refId="B78" refString="Godwin-Austen, HH, 1895. List and Distribution of the Land-Mollusca of the Andaman and Nicobar Islands, with Descriptions of some supposed new Species. Proceedings of the Zoological Society of London [1895]: 438-456. https://www.biodiversitylibrary.org/page/30983429" title="List and Distribution of the Land-Mollusca of the Andaman and Nicobar Islands, with Descriptions of some supposed new Species. Proceedings of the Zoological Society of London [1895]: 438 - 456." url="https://www.biodiversitylibrary.org/page/30983429" year="1895">Godwin-Austen 1895</bibRefCitation>
, as
<taxonomicName lsidName="O. mauritianum" pageId="0" pageNumber="1" rank="species" species="mauritianum">
<emphasis italics="true" pageId="0" pageNumber="1">O. mauritianum</emphasis>
</taxonomicName>
; Patil &amp; Kulkarni, 2013); Indonesia, West Papua (Quoy and Gaimard 1832, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
); Japan (
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)73859-7" author="Arimoto, H" journalOrPublisher="Tetrahedron Letters" pageId="0" pageNumber="1" pagination="5781 - 5784" refId="B8" refString="Arimoto, H, Cheng, JF, Nishiyama, S, Yamamura, S, 1993. Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II. Tetrahedron Letters 34 (36): 5781 - 5784, DOI: https://doi.org/10.1016/S0040-4039(00)73859-7" title="Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II." url="https://doi.org/10.1016/S0040-4039(00)73859-7" volume="34" year="1993">Arimoto et al. 1993</bibRefCitation>
); Kenya (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.46994" author="Martens, E von" journalOrPublisher="Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien" pageId="0" pageNumber="1" pagination="1 - 332" refId="B125" refString="Martens, E von, 1897. Suess- und brackwasser-mollusken des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien 4: 1 - 332, DOI: https://doi.org/10.5962/bhl.title.46994" title="Suess- und brackwasser-mollusken des Indischen Archipels." url="https://doi.org/10.5962/bhl.title.46994" volume="4" year="1897">Martens 1897</bibRefCitation>
); Madagascar (
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus 1970</bibRefCitation>
); Mariana Islands, Guam (
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)98720-3" author="Biskupiak, JE" journalOrPublisher="Tetrahedon Letters" pageId="0" pageNumber="1" pagination="4307 - 4310" refId="B19" refString="Biskupiak, JE, Ireland, CM, 1985. Cytotoxic metabolites from the mollusc Peronia peronii. Tetrahedon Letters 26: 4307 - 4310, DOI: https://doi.org/10.1016/S0040-4039(00)98720-3" title="Cytotoxic metabolites from the mollusc Peronia peronii." url="https://doi.org/10.1016/S0040-4039(00)98720-3" volume="26" year="1985">Biskupiak and Ireland 1985</bibRefCitation>
); Mozambique (
<bibRefCitation author="Martens, E von" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/36341972" refId="B124" refString="Martens, E von, 1879. Uebersicht der von W. Peters von 1843 bis 1847 in Mossambique gesammelten Mollusca. Monatsberichte der koeniglich Preussischen Akademie der Wissenschaften zu Berlin [1879]: 727-749. https://biodiversitylibrary.org/page/36341972" title="Uebersicht der von W. Peters von 1843 bis 1847 in Mossambique gesammelten Mollusca. Monatsberichte der koeniglich Preussischen Akademie der Wissenschaften zu Berlin [1879]: 727 - 749." url="https://biodiversitylibrary.org/page/36341972" year="1879">Martens 1879</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly 1912</bibRefCitation>
,
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">1939</bibRefCitation>
;
<bibRefCitation author="Macnae, W" journalOrPublisher="Mitochondrial DNA" pageId="0" pageNumber="1" refId="B118" refString="Macnae, W, Kalk, M [Eds], 1958. A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." title="A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." year="1958">Macnae and Kalk 1958</bibRefCitation>
); Papua New Guinea, New Ireland (
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
1934a
</bibRefCitation>
, as
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
); Persian Gulf (
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a064590" author="White, KM" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="241 - 253" refId="B195" refString="White, KM, 1951. On a collection of Molluscs, mainly Nudibranchs from the Red Sea. Proceedings of the Malacological Society of London 28 (6): 241 - 253, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a064590" title="On a collection of Molluscs, mainly Nudibranchs from the Red Sea." url="https://doi.org/10.1093/oxfordjournals.mollus.a064590" volume="28" year="1951">White 1951</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.13005/bbra/1870" author="Bitaab, MA" journalOrPublisher="Biosciences Biotechnology Research Asia" pageId="0" pageNumber="1" pagination="2023 - 2035" refId="B20" refString="Bitaab, MA, Ranaei Siadat, SO, Pazooki, J, Sefidbakht, Y, 2015. Antibacterial and molecular dynamics study of the Dolabellanin B2 isolated from sea slug, Peronia peronii. Biosciences Biotechnology Research Asia 12 (3): 2023 - 2035, DOI: https://doi.org/10.13005/bbra/1870" title="Antibacterial and molecular dynamics study of the Dolabellanin B 2 isolated from sea slug, Peronia peronii." url="https://doi.org/10.13005/bbra/1870" volume="12" year="2015">Bitaab et al. 2015</bibRefCitation>
); Philippines (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12547" author="Casto de Elera, RPF" journalOrPublisher="Colegio de Santo Tomas, Manila" pageId="0" pageNumber="1" refId="B30" refString="Casto de Elera, RPF, 1896. Catalogo sistematico de toda la Fauna de Filipinas conocida hasta el presente, y a la vez el de la coleccion zoologica del museo de PP. Dominicos del Colegio-Universidad de Sto. Tomas de Manila (Vol. 3). Colegio de Santo Tomas, Manila, DOI: https://doi.org/10.5962/bhl.title.12547" title="Catalogo sistematico de toda la Fauna de Filipinas conocida hasta el presente, y a la vez el de la coleccion zoologica del museo de PP. Dominicos del Colegio-Universidad de Sto. Tomas de Manila (Vol. 3)." url="https://doi.org/10.5962/bhl.title.12547" year="1896">Casto de Elera 1896</bibRefCitation>
, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
), Bohol (Semper 1880, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
); Red Sea (
<bibRefCitation author="Sturany, R" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe" pageId="0" pageNumber="1" pagination="209 - 283" publicationUrl="https://biodiversitylibrary.org/page/6903441" refId="B175" refString="Sturany, R, 1904. Expeditionen S.M. Schiff &quot;Pola&quot; in das Rote Meer. Noerdliche und suedliche Haelfte. 1895/96-1897/98. Zoologische Ergebnisse. XXIII. Gastropoden des Rothen Meeres. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 74: 209 - 283, https://biodiversitylibrary.org/page/6903441" title="Expeditionen S. M. Schiff &quot; Pola &quot; in das Rote Meer. Noerdliche und suedliche Haelfte. 1895 / 96 - 1897 / 98. Zoologische Ergebnisse. XXIII. Gastropoden des Rothen Meeres." url="https://biodiversitylibrary.org/page/6903441" volume="74" year="1904">Sturany 1904</bibRefCitation>
); Samoa (Semper 1880, as
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
); South Africa, Natal (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13936" author="Krauss, F" journalOrPublisher="Ebner &amp; Seubert, Stuttgart" pageId="0" pageNumber="1" refId="B106" refString="Krauss, F, 1848. Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten. Ebner &amp; Seubert, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.13936" title="Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten." url="https://doi.org/10.5962/bhl.title.13936" year="1848">Krauss 1848</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12875" author="Sturany, R" journalOrPublisher="Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien" pageId="0" pageNumber="1" refId="B174" refString="Sturany, R, 1898. Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales. Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien, DOI: https://doi.org/10.5962/bhl.title.12875" title="Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales." url="https://doi.org/10.5962/bhl.title.12875" year="1898">Sturany 1898</bibRefCitation>
;
<bibRefCitation author="Collinge, WE" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="159 - 174" publicationUrl="https://www.biodiversitylibrary.org/page/6322951" refId="B36" refString="Collinge, WE, 1910. The slugs of Natal. Annals of the Natal Museum 2: 159 - 174, https://www.biodiversitylibrary.org/page/6322951" title="The slugs of Natal." url="https://www.biodiversitylibrary.org/page/6322951" volume="2" year="1910">Collinge 1910</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly 1912</bibRefCitation>
,
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">1939</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1201/EBK1439821169-2" author="Morrisey, D" editor="Gibson, RN" journalOrPublisher="(Vol. 48). CRC Press, Boca Raton" pageId="0" pageNumber="1" pagination="43 - 160" refId="B132" refString="Morrisey, D, Swales, A, Ditttmann, S, Morrison, MA, Lovelock, CE, Beard, CM, 2010. The ecology and management of temperate mangroves. In: Gibson, RN, Atkinson, RJA, Gordon, JDM, Eds., Oceanography and Marine Biology: an Annual Review. (Vol. 48). CRC Press, Boca Raton: 43 - 160, DOI: https://doi.org/10.1201/EBK1439821169-2" title="The ecology and management of temperate mangroves." url="https://doi.org/10.1201/EBK1439821169-2" volumeTitle="Oceanography and Marine Biology: an Annual Review." year="2010">Morrisey et al. 2010</bibRefCitation>
); Australia, Lord Howe Island (
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
;
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
), Torres Strait (
<bibRefCitation author="Smith, EA" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://www.biodiversitylibrary.org/item/63054" refId="B165" refString="Smith, EA, 1884. Mollusca. In: [no named editor] Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881-2. Part I. The collections from Melanesia; Part II. Collections from the western Indian Ocean. The Trustees of the British Museum, London, 34-116, 487-508. http://www.biodiversitylibrary.org/item/63054" title="Mollusca. In: [no named editor] Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 2. Part I. The collections from Melanesia; Part II. Collections from the western Indian Ocean. The Trustees of the British Museum, London, 34 - 116, 487 - 508." url="http://www.biodiversitylibrary.org/item/63054" year="1884">Smith 1884</bibRefCitation>
, as
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
), Western Australia (
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
was named after
<normalizedToken originalValue="François">Francois</normalizedToken>
<normalizedToken originalValue="Péron">Peron</normalizedToken>
[1775-1810] who collected the two slugs described by Cuvier during the Baudin expedition [1800-1803].
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
, and
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
were named after type localities.
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
was named after the speckled (
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">punctatum</emphasis>
</taxonomicName>
in Latin) dorsal notum of live animals.
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium melanopneumon</emphasis>
</taxonomicName>
was named after the black (
<emphasis italics="true" pageId="0" pageNumber="1">melas</emphasis>
in Greek) lung (
<emphasis italics="true" pageId="0" pageNumber="1">pneumon</emphasis>
in Greek) tissue of the holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F9" captionText="Figure 7. Habitat, Peronia peronii, Papua New Guinea, Madang, limestone rocky intertidal (st PM 12)." figureDoi="10.3897/zookeys.972.52853.figure7" httpUri="https://binary.pensoft.net/fig/458307" pageId="0" pageNumber="1">7</figureCitation>
). Live slugs of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
are found in the rocky intertidal, like most other
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
slugs. Many of our specimens were collected at night or just before sunrise, suggesting that
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is, at least partly, a nocturnal species. This could explain why we missed it at some localities where we only collected during the day.
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
is not rare, but it is definitely not as common as some other species. The fact that collecting it at night seems necessary, at least in some localities, might explain why collections of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
are not as abundant as collections of
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure7" httpUri="https://binary.pensoft.net/fig/458307" pageId="0" pageNumber="1" start="Figure 7" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
Habitat,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, Papua New Guinea, Madang, limestone rocky intertidal (st PM 12).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="color and morphology of live animals">
<paragraph pageId="0" pageNumber="1">Color and morphology of live animals</paragraph>
<paragraph pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F10" captionText="Figure 8. Live animals, Peronia peronii, Mauritius A dorsal view, 140 mm long [5872] (MNHN-IM- 2019 - 1605) B dorsal view, 125 mm long [5874] (MNHN-IM- 2019 - 1605) C ventral view, same as A; D ventral view, same as B." figureDoi="10.3897/zookeys.972.52853.figure8" httpUri="https://binary.pensoft.net/fig/458308" pageId="0" pageNumber="1">8</figureCitation>
). No picture of live animals was available for individuals from the West Pacific (Guam and Papua New Guinea). The description of the color of live animals is based on the Mauritius individuals. The dorsal notum is brown, with a greenish hue, light to dark, mottled with darker and lighter areas. The color of the dorsal papillae varies a s that of the background itself. The ventral surface (foot and hyponotum) is yellowish-greenish and can change rapidly in any given individual. The ocular tentacles are brown-grey, like the head. The dorsal notum of live animals is covered by dozens of papillae of various sizes. Dorsal papillae can be particularly tall (easily up to 4 mm), even in preserved specimens, and are evenly distributed over the entire notum. Preserved, they are difficult to distinguish from retracted dorsal gills in the posterior half of the notum. Some papillae bear black dorsal eyes at their tip. The number of papillae with dorsal eyes is variable (15-20). The longest animals are 140 mm long in Mauritius and 115 mm long in the West Pacific.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure8" httpUri="https://binary.pensoft.net/fig/458308" pageId="0" pageNumber="1" start="Figure 8" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
Live animals,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, Mauritius
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
dorsal view, 140 mm long [5872] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
dorsal view, 125 mm long [5874] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
ventral view, same as
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
;
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
ventral view, same as
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="digestive system">
<paragraph pageId="0" pageNumber="1">Digestive system</paragraph>
<paragraph pageId="0" pageNumber="1">
(Figs
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9</figureCitation>
-
<figureCitation captionStart="Figure 12" captionStartId="F14" captionText="Figure 12. Radula, Peronia peronii, Mauritius [5872] (MNHN-IM- 2019 - 1605) A rachidian and innermost lateral teeth B rachidian and lateral teeth C outermost lateral teeth D lateral teeth. Scale bars: 40 μm (A), 100 μm (B), 20 μm (C), 200 μm (D)." figureDoi="10.3897/zookeys.972.52853.figure12" httpUri="https://binary.pensoft.net/fig/458312" pageId="0" pageNumber="1">12</figureCitation>
). Examples of radular formulae are presented in Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Radular formulae in Peronia species. Each formula follows the same format: number of rows x number of lateral teeth per left half row - 1 (rachidian tooth) - number of lateral teeth per right half row. Each DNA extraction number corresponds to one individual. The letter H next to an extraction number indicates a holotype." httpUri="http://table.plazi.org/id/F6A603768F442AB5CA201CDCEC46CA6A" pageId="0" pageNumber="1" tableUuid="F6A603768F442AB5CA201CDCEC46CA6A">5</tableCitation>
. The median cusp of the rachidian teeth is approximately 75
<normalizedToken originalValue="μm">μm</normalizedToken>
long. The hook of the lateral teeth is approximately 160-200
<normalizedToken originalValue="μm">μm</normalizedToken>
long. The intestinal loops are of type I, with a transitional loop oriented between 12 to 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1" start="Figure 9" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
Digestive system, dorsal view,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, type specimens. The arrow indicates the orientation of the transitional loop
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
paralectotype,
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
, Timor (MNHN-IM-2000-22938)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
lectotype,
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
, Tonga (MNHN-IM-2000-22937)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
lectotype,
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
, Indonesia, West Papua (MNHN-IM-2000-22966)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
possible paralectotype,
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia gondwanae" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gondwanae">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia gondwanae</emphasis>
</taxonomicName>
, Mauritius (MNHN-IM-2000-33686)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
lectotype,
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
, Fiji (MNHN-IM-2000-33692). Scale bars: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, D</emphasis>
), 10 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B, C, E</emphasis>
). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure10" httpUri="https://binary.pensoft.net/fig/458310" pageId="0" pageNumber="1" start="Figure 10" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
Digestive system, dorsal view,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
. The arrow indicates the orientation of the transitional loop
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Mauritius [5874] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Papua New Guinea, Madang [5472] (MNHN-IM-2013-14052)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Guam [5840] (MNHN-IM-2019-1609). Scale bars: 10 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure11" httpUri="https://binary.pensoft.net/fig/458311" pageId="0" pageNumber="1" start="Figure 11" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
Radula,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
Papua New Guinea [5476] (MNHN-IM-2013-16260)
<emphasis bold="true" pageId="0" pageNumber="1">D-H</emphasis>
Guam [5840] (MNHN-IM-2019-1609)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
rachidian and innermost lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
rachidian and innermost lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
rachidian and innermost lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
rachidian and lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
lateral teeth, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
outermost lateral teeth. Scale bars: 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 40
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B, D</emphasis>
), 80
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
), 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">G, H</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure12" httpUri="https://binary.pensoft.net/fig/458312" pageId="0" pageNumber="1" start="Figure 12" startId="F14">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
Radula,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, Mauritius [5872] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
rachidian and innermost lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
rachidian and lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
outermost lateral teeth
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral teeth. Scale bars: 40
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
), 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), 200
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
).
</paragraph>
</caption>
<caption ID-Table-UUID="F6A603768F442AB5CA201CDCEC46CA6A" httpUri="http://table.plazi.org/id/F6A603768F442AB5CA201CDCEC46CA6A" pageId="0" pageNumber="1" start="Table 5" startId="T5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 5.</emphasis>
Radular formulae in
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species. Each formula follows the same format: number of rows
<normalizedToken originalValue="×">x</normalizedToken>
number of lateral teeth per left half row - 1 (rachidian tooth) - number of lateral teeth per right half row. Each DNA extraction number corresponds to one individual. The letter H next to an extraction number indicates a holotype.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Radular formula</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Spm length (mm)</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Locality</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">DNA extraction number</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="9">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #1)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
63
<normalizedToken originalValue="×">x</normalizedToken>
70-1-70
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00170</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Sulawesi</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2127</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
73
<normalizedToken originalValue="×">x</normalizedToken>
92-1-92
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ITBZC IM 00021</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Vietnam</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5621</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
56
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MTQ</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Queensland</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2622</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
74
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00166</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Halmahera</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5068</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
52
<normalizedToken originalValue="×">x</normalizedToken>
57-1-57
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00162</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Ambon</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2729</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
60
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00168</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Lombok</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2987</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
72-1-72
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00169</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Seram</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2870</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-12010</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5469</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
56
<normalizedToken originalValue="×">x</normalizedToken>
56-1-56
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-62393</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Vanuatu</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5481</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="4">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #2)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00178</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Sumatra</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1746</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
78-1-78
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00180</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Sumatra</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1797</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
60
<normalizedToken originalValue="×">x</normalizedToken>
68-1-68
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00180</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Sumatra</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1795</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
64
<normalizedToken originalValue="×">x</normalizedToken>
70-1-70
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00180</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Sumatra</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1796</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="4">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #3)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
68
<normalizedToken originalValue="×">x</normalizedToken>
78-1-78
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">USMMC 00051</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Peninsular Malaysia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">976</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
92-1-92
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">USMMC 00065</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Peninsular Malaysia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2547</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
65
<normalizedToken originalValue="×">x</normalizedToken>
85-1-85
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZRC.MOL.10496</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Singapore</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">990</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
72
<normalizedToken originalValue="×">x</normalizedToken>
86-1-86
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">USMMC 00064</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Peninsular Malaysia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">975</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #4)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
73
<normalizedToken originalValue="×">x</normalizedToken>
85-1-85
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1384</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pakistan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6164</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
75
<normalizedToken originalValue="×">x</normalizedToken>
95-1-95
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1385</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pakistan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6165</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
75
<normalizedToken originalValue="×">x</normalizedToken>
92-1-92
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1386</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Pakistan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6166</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
65
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1610</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Madagascar</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3231</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
55
<normalizedToken originalValue="×">x</normalizedToken>
63-1-63
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1611</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Madagascar</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3144</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
55
<normalizedToken originalValue="×">x</normalizedToken>
65-1-65
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-62398</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mozambique</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5510</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="4">
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(Red Sea)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
66
<normalizedToken originalValue="×">x</normalizedToken>
86-1-86
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZMH 27472</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Red Sea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">#1</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
80
<normalizedToken originalValue="×">x</normalizedToken>
95-1-95
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZMH 27472</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Red Sea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">#2</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
67
<normalizedToken originalValue="×">x</normalizedToken>
80-1-80
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZMH 27472</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Red Sea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">#3</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
80-1-80
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZMH 27472</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Red Sea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">#4</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="4">
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
85
<normalizedToken originalValue="×">x</normalizedToken>
90-1-90
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2009-16392</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Madagascar</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5501</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
78
<normalizedToken originalValue="×">x</normalizedToken>
90-1-90
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2009-16412</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Madagascar</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5504</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
47
<normalizedToken originalValue="×">x</normalizedToken>
50-1-50
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 332088</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Oman</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">703</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
72-1-72
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NMSA W7547</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">South Africa</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5841</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="5">
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
90
<normalizedToken originalValue="×">x</normalizedToken>
115-1-115
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">110</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1606</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mauritius</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3605</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
90
<normalizedToken originalValue="×">x</normalizedToken>
105-1-105
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">140</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1605</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mauritius</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5872</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
95
<normalizedToken originalValue="×">x</normalizedToken>
105-1-105
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">80</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-14052</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5472</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
90
<normalizedToken originalValue="×">x</normalizedToken>
100-1-100
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-16260</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5476</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
100
<normalizedToken originalValue="×">x</normalizedToken>
110-1-110
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1609</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Guam</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5840</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="6">
<taxonomicName lsidName="P. platei" pageId="0" pageNumber="1" rank="species" species="platei">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. platei</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
75
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 303653</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Hawaii</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">706</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
65
<normalizedToken originalValue="×">x</normalizedToken>
78-1-78
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 303653</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Hawaii</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5380</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
90-1-90
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">BPBM 284527</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Hawaii</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6160</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
80-1-80
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">BPBM 284528</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Hawaii</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6161</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
67
<normalizedToken originalValue="×">x</normalizedToken>
75-1-75
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-13762</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5405</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
72-1-72
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2013-13351</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Papua New Guinea</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5412</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="7">
<taxonomicName lsidName="P. sydneyensis" pageId="0" pageNumber="1" rank="species" species="sydneyensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. sydneyensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
56
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">AM C468916.001</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">New South Wales</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1516 H</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
51
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">AM C468915.001</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">New South Wales</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1517</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
47
<normalizedToken originalValue="×">x</normalizedToken>
45-1-45
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MTQ</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Queensland</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2667</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
58
<normalizedToken originalValue="×">x</normalizedToken>
70-1-70
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MTQ</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Queensland</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2680</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
35
<normalizedToken originalValue="×">x</normalizedToken>
35-1-35
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1594</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">New Caledonia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6189</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
55
<normalizedToken originalValue="×">x</normalizedToken>
70-1-70
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">41</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1595</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">New Caledonia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6195</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
45
<normalizedToken originalValue="×">x</normalizedToken>
46-1-46
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1598</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">New Caledonia</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6220</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">
<taxonomicName lsidName="P. okinawensis" pageId="0" pageNumber="1" rank="species" species="okinawensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. okinawensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
60
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 352288</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Okinawa, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">696-2</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
60
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 352288</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Okinawa, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">696-3</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
65
<normalizedToken originalValue="×">x</normalizedToken>
65-1-65
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">27</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UF 352288</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Okinawa, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">696-4 H</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">
<taxonomicName lsidName="P. setoensis" pageId="0" pageNumber="1" rank="species" species="setoensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. setoensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
50-1-50
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NSMT-Mo 78985</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Wakayama, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">5383 H</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
55
<normalizedToken originalValue="×">x</normalizedToken>
55-1-55
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NSMT-Mo 78987</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Wakayama, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3753</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
50-1-50
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NSMT-Mo 78987</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Wakayama, Japan</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3754</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="6">
<taxonomicName lsidName="P. griffithsi" pageId="0" pageNumber="1" rank="species" species="griffithsi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. griffithsi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
45-1-45
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00177</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Kei</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2934</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
50-1-50
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">17</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00176</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Kei</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2936</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
45-1-45
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">UMIZ 00177</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Kei</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3566</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
45
<normalizedToken originalValue="×">x</normalizedToken>
52-1-52
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1608</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mauritius</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3153</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
52-1-52
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2019-1608</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mauritius</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3156</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
50
<normalizedToken originalValue="×">x</normalizedToken>
50-1-50
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MNHN-IM-2000-35265</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Mauritius</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3157 H</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="4">
<taxonomicName lsidName="P. willani" pageId="0" pageNumber="1" rank="species" species="willani">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">P. willani</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
58
<normalizedToken originalValue="×">x</normalizedToken>
80-1-80
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NTM P.57625</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Northern Territory</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1628 H</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
53
<normalizedToken originalValue="×">x</normalizedToken>
60-1-60
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NTM P.57627</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Northern Territory</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1668</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
65
<normalizedToken originalValue="×">x</normalizedToken>
80-1-80
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NTM P.57627</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Northern Territory</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1626</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
70
<normalizedToken originalValue="×">x</normalizedToken>
90-1-90
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">NTM P.57626</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Northern Territory</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1620</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reproductive system">
<paragraph pageId="0" pageNumber="1">Reproductive system</paragraph>
<paragraph pageId="0" pageNumber="1">
(Figs
<figureCitation captionStart="Figure 13" captionStartId="F15" captionText="Figure 13. Posterior, hermaphroditic (female) reproductive system, Peronia peronii A Mauritius [3605] (MNHN-IM- 2019 - 1606) B Papua New Guinea, Madang [5472] (MNHN-IM- 2013 - 14052). Scale bars: 10 mm (A), 5 mm (B). Abbreviations: dd deferent duct, fgm female gland mass, hg hermaphroditic gland, ov oviduct, rs receptaculum seminis, sp spermatheca." figureDoi="10.3897/zookeys.972.52853.figure13" httpUri="https://binary.pensoft.net/fig/458313" pageId="0" pageNumber="1">13</figureCitation>
-
<figureCitation captionStart="Figure 16" captionStartId="F18" captionText="Figure 16. Accessory penial gland spine, Peronia peronii A, B, F, G Mauritius C, D, H Papua New Guinea E, I Guam A [3605] (MNHN-IM- 2019 - 1606) B [5872] (MNHN-IM- 2019 - 1605) C [5471] (MNHN-IM- 2013 - 12500) D [5472] (MNHN-IM- 2013 - 14052) E [5840] (MNHN-IM- 2019 - 1609) F same as A; G same as B; H same as C; I same as E. Scale bars: 800 μm (A-D), 1 mm (E), 40 μm (F-I)." figureDoi="10.3897/zookeys.972.52853.figure16" httpUri="https://binary.pensoft.net/fig/458316" pageId="0" pageNumber="1">16</figureCitation>
). In the anterior (male) parts, the muscular sac of the accessory penial gland is at least 30 mm long in specimens from Mauritius and at least 25 mm long in specimens from the West Pacific (Guam &amp; Papua New Guinea). Note that, in some additional museum specimens, the muscular sac was only 20 mm long, and, even exceptionally 17 mm long (see remarks below). The hollow spine of the accessory penial gland is narrow, elongated, and straight or slightly curved, and its shape (including at its tip) varies between individuals. Its length ranges from 3.4 mm ([5872] MNHN-IM-2019-1605) to 3.6 mm ([3605] MNHN-IM-2019-1606) in Mauritius, and from 3.5 mm ([5472] MNHN-IM-2013-14052) to 4 mm ([5471] MNHN-IM-2013-12500) in the West Pacific (Guam and Papua New Guinea). Its diameter at the conical base is approximately 400
<normalizedToken originalValue="μm">μm</normalizedToken>
in specimens from Mauritius and between 400 and 500
<normalizedToken originalValue="μm">μm</normalizedToken>
in specimens from the West Pacific (Guam and Papua New Guinea). Its diameter at the tip measures 160-170
<normalizedToken originalValue="μm">μm</normalizedToken>
in specimens from the West Pacific, and from 180 to 200
<normalizedToken originalValue="μm">μm</normalizedToken>
in specimens from Mauritius. Note that, in some additional museum specimens, the spine was only 3 mm long (see remarks below).
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure13" httpUri="https://binary.pensoft.net/fig/458313" pageId="0" pageNumber="1" start="Figure 13" startId="F15">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
Posterior, hermaphroditic (female) reproductive system,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Mauritius [3605] (MNHN-IM-2019-1606)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Papua New Guinea, Madang [5472] (MNHN-IM-2013-14052). Scale bars: 10 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
). Abbreviations: dd deferent duct, fgm female gland mass, hg hermaphroditic gland, ov oviduct, rs receptaculum seminis, sp spermatheca.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure14" httpUri="https://binary.pensoft.net/fig/458314" pageId="0" pageNumber="1" start="Figure 14" startId="F16">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
Anterior, male, copulatory apparatus,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Mauritius [3605] (MNHN-IM-2019-1606)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Papua New Guinea, Madang [5472] (MNHN-IM-2013-14052). Scale bars: 10 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
), 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
). Abbreviations: ag accessory penial gland, dd deferent duct, ms muscular sac, ps penial sheath, rm retractor muscle, v vestibule.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
The retractor muscle is shorter or longer than the penial sheath and inserts near the heart. Exceptionally, the retractor muscle can even be vestigial ([5472] MNHN-IM-2013-14052). Inside the penial sheath, the penis is a narrow, elongated, soft, hollow tube. Its distal end bears conical hooks which are less than 50
<normalizedToken originalValue="μm">μm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
<paragraph pageId="0" pageNumber="1">Diagnostic features</paragraph>
<paragraph pageId="0" pageNumber="1">
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
).
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
is the only
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species which is easy to identify anatomically. Indeed, it is characterized by a very long spine (at least 3 mm) of the accessory penial gland, which is distinctive and easily accessible (one just needs to pull on the flagellum of the penial gland or, even, in some cases, measure the spine by transparency). The two longest spines were found in the lectotype of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
(MNHN-IM-2000-33692) from Fiji (5 mm), and in an old historical specimen (ANSP 304860) from the Maldives (4.8 mm).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
is additionally characterized by a unique combination of anatomical traits: muscular sac longer than 20 mm, intestinal loops of type I (with a transitional loop oriented between 12 and 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
), retractor muscle inserting near the heart. Also, no individual larger than 80 mm was found in any other
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species so far. Animal size can be useful when several
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species are found at the same site. For instance, the two individuals of
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #1) found at the station PM 12 (near Madang, Papua New Guinea) are 35 and 38 mm long while the individual of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
from the same station is 80 mm long. The type I of its intestinal loops (with a transitional loop oriented between 12 and 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
) is only shared by
<taxonomicName lsidName="P. okinawensis" pageId="0" pageNumber="1" rank="species" species="okinawensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. okinawensis</emphasis>
</taxonomicName>
, a species endemic to Japan with which it is most closely related.
</paragraph>
<caption doi="10.3897/zookeys.972.52853.figure15" httpUri="https://binary.pensoft.net/fig/458315" pageId="0" pageNumber="1" start="Figure 15" startId="F17">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 15.</emphasis>
Penial hooks,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Mauritius [5872] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Mauritius [3605] (MNHN-IM-2019-1606)
<emphasis bold="true" pageId="0" pageNumber="1">C, D</emphasis>
Guam [5840] (MNHN-IM-2019-1609). Scale bars: 6
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A, B, D</emphasis>
), 40
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.972.52853.figure16" httpUri="https://binary.pensoft.net/fig/458316" pageId="0" pageNumber="1" start="Figure 16" startId="F18">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
Accessory penial gland spine,
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A, B, F, G</emphasis>
Mauritius
<emphasis bold="true" pageId="0" pageNumber="1">C, D, H</emphasis>
Papua New Guinea
<emphasis bold="true" pageId="0" pageNumber="1">E, I</emphasis>
Guam
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
[3605] (MNHN-IM-2019-1606)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
[5872] (MNHN-IM-2019-1605)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
[5471] (MNHN-IM-2013-12500)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
[5472] (MNHN-IM-2013-14052)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
[5840] (MNHN-IM-2019-1609)
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
same as
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
;
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
same as
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
;
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
same as
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
;
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
same as
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
. Scale bars: 800
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
), 1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
), 40
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">F-I</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Synonymies.</emphasis>
There is no doubt that
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804)
</bibRefCitation>
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
applies to the species described here, just based on animal size alone. According to Cuvier, the lectotype from Mauritius measured approximately 140 mm long and our molecular data show that all individuals of that size from Mauritius belong to a single species (Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
).
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6) detailed anatomical description and drawings are exclusively based on the lectotype (he did not dissect the paralectotype from Timor).
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: 48, pl. 6, fig. 8) described the spine of the accessory penial gland as a &quot;very sharp, brown spike&quot; but unfortunately did not provide its length. However,
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6, fig. 4) illustration of the intestinal loops is identical to some of our Mauritius individuals here: intestinal loops of type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
. The paralectotype of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Timor (MNHN-IM-2000- 22938) is only briefly mentioned by Cuvier in the original description. The length (4.5 mm) of the spine of its accessory penial gland (checked for the present study) indicates that it also belongs to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. Its intestinal loops are also identical to those of the lectotype (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
is a junior objective synonym of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
because they share the same name-bearing type.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
was described by Quoy and Gaimard (1832: 210-211, pl. 15, figs 17, 18) from
<normalizedToken originalValue="“Panhi-Motou,”">&quot;Panhi-Motou,&quot;</normalizedToken>
possibly the small island of Pangaimotu, Tonga, based on an unspecified number of individuals. The illustrations of the dorsal gills by Quoy and Gaimard (1832: pl. 15, figs 17, 18) and their presence on the notum of the lectotype (MNHN-IM-2000-22937) clearly indicate that
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
belongs to
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
. The lectotype was dissected prior to the present study and most of the male copulatory parts are missing (only the deferent duct remains). As a result, the length of the spine of the accessory penial gland, which is diagnostic of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, cannot be checked.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 191) listed the lectotype in the material he examined for his re-description of
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
, but he did not point out that it was part of the type material of
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
and he did not describe it anatomically. It is possible but not certain that
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
dissected the lectotype. At any rate, its intestinal loops are of type I with a transitional loop between 2 and 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9B</figureCitation>
). Both the length (100 mm) of the lectotype as well as its intestinal loops indicate that
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia tongana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia tongana</emphasis>
</taxonomicName>
is a junior synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
is regarded here as a junior synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
because the length (3.7 mm) of the spine of the accessory penial gland of the lectotype (MNHN-IM-2000-22966) is only compatible with
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). The length of the lectotype (70 mm, preserved) is also far more compatible with
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
than with
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
, another species found in West Papua. Our many individuals of
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
are all less than 60 mm long (alive), except a single individual from New Caledonia (73 mm alive). Given their small size, the two paralectotypes (MNHN-IM-2000-33701) likely belong to
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #1) instead of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, which would not be surprising at all because the type locality of
<taxonomicName lsidName="O. ferrugineum" pageId="0" pageNumber="1" rank="species" species="ferrugineum">
<emphasis italics="true" pageId="0" pageNumber="1">O. ferrugineum</emphasis>
</taxonomicName>
(a junior synonym of
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
) is the same as that of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
(Manokwari, West Papua, Indonesia). At the end of the description of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
, Quoy &amp; Gaimard (1832: 216) also mention in passing that they also found
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
in Port Dorey (i.e., Manokwari, West Papua, Indonesia) and they even point out that local inhabitants know how to distinguish both species. Both
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
are regarded here as junior synonyms of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. However, it remains true that there are two sympatric
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species in West Papua,
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, which can be distinguished in the field based on animal length (except, of course, for individuals measuring less than 60 mm long).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh (1884a</bibRefCitation>
: 129-142, pl. IV, figs 25-27, pl. V, figs 1-27, pl. VI, figs 5-18, 20, 21) described
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium melanopneumon</emphasis>
</taxonomicName>
from a single individual (65/40 mm) from Fiji. This specimen was completely dissected by Bergh and is now empty (NHMUK 1888.5.30.39).
<taxonomicName authorityName="Bergh" authorityYear="1884" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium melanopneumon" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium melanopneumon</emphasis>
</taxonomicName>
applies to a
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species due to the presence of dorsal gills, and the length (4 mm) of the spine of the accessory penial gland indicates that it applies to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Its intestinal loops (
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh 1884a</bibRefCitation>
: pl. V, fig. 27) are also similar to those found in
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, although the transitional loop is slightly past the 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
limit. As a result,
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
is regarded as a junior synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.1080/00222938409459805" author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" refId="B17" refString="Bergh, R, 1884b. On the affinities of the Onchidia. Annals and Magazine of Natural History 14(5th series): 259-266. [This is a translation of Bergh (1885)] https://doi.org/10.1080/00222938409459805" title="On the affinities of the Onchidia. Annals and Magazine of Natural History 14 (5 th series): 259 - 266. [This is a translation of Bergh (1885)]" url="https://doi.org/10.1080/00222938409459805" year="1884 b">Bergh (1884b</bibRefCitation>
: 263;
<bibRefCitation author="Bergh, R" journalOrPublisher="Morphologisches Jahrbuch" pageId="0" pageNumber="1" pagination="172 - 181" publicationUrl="https://biodiversitylibrary.org/page/13028090" refId="B18" refString="Bergh, R, 1885. Ueber die Verwandtschaftsbeziehungen der Onchidien. Morphologisches Jahrbuch 10: 172 - 181, https://biodiversitylibrary.org/page/13028090" title="Ueber die Verwandtschaftsbeziehungen der Onchidien." url="https://biodiversitylibrary.org/page/13028090" volume="10" year="1885">1885</bibRefCitation>
: 176) briefly mentioned again
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
in a comparative study on the affinities of onchidiids.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 197-198, figs 9-11) described
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
based on seven individuals from Fiji, one of which could be found and is designated as the lectotype (MNHN-IM-2000-33692). Because all reproductive parts are missing, the length of the spine of the accessory penial gland cannot be checked. However, according to
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 197, fig. 10), the spine of the accessory penial gland is 5 mm long, which is only compatible with
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
), and is the longest spine known in
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. The intestinal loops of the lectotype of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
are clearly of type I, with a transitional loop oriented at ~ 1
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F11" captionText="Figure 9. Digestive system, dorsal view, Peronia peronii, type specimens. The arrow indicates the orientation of the transitional loop A paralectotype, Onchidium peronii, Timor (MNHN-IM- 2000 - 22938) B lectotype, Onchidium tonganum, Tonga (MNHN-IM- 2000 - 22937) C lectotype, Onchidium punctatum, Indonesia, West Papua (MNHN-IM- 2000 - 22966) D possible paralectotype, Paraperonia gondwanae, Mauritius (MNHN-IM- 2000 - 33686) E lectotype, Paraperonia fidjiensis, Fiji (MNHN-IM- 2000 - 33692). Scale bars: 5 mm (A, D), 10 mm (B, C, E). Abbreviations: ddg dorsal digestive gland, i intestine, pdg posterior digestive gland, st stomach." figureDoi="10.3897/zookeys.972.52853.figure9" httpUri="https://binary.pensoft.net/fig/458309" pageId="0" pageNumber="1">9E</figureCitation>
), even though
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 197) erroneously described them a type V, which is a mistake he often made. Given the length of the lectotype (60 mm) and, most importantly, the length of the spine of the accessory penial gland,
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
is regarded as a junior synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Secondary literature</emphasis>
.
</emphasis>
Several early authors mentioned
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
without any new material (
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/1844622" refId="B44" refString="Cuvier, G, 1816. Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee (Vol. 2). Deterville, Paris, xviii + 532 pp.[according to Roux (1976), all four volumes were published by the end of December 1816, even although the year 1817 is printed on the title page] https://www.biodiversitylibrary.org/page/1844622" title="Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee (Vol. 2). Deterville, Paris, xviii + 532 pp. [according to Roux (1976), all four volumes were published by the end of December 1816, even although the year 1817 is printed on the title page]" url="https://www.biodiversitylibrary.org/page/1844622" year="1816">Cuvier 1816</bibRefCitation>
: 411;
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/33372853" refId="B45" refString="Cuvier, G, 1830. Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX] https://www.biodiversitylibrary.org/page/33372853" title="Le regne animal distribue d'apres son organisation (Vol. 3). Deterville, Paris, xvi + 504 pp. [pls I-XX]" url="https://www.biodiversitylibrary.org/page/33372853" year="1830">Cuvier 1830</bibRefCitation>
: 46;
<normalizedToken originalValue="Férussac">Ferussac</normalizedToken>
1822: xxxi;
<bibRefCitation author="Fleming, J" editor="Napier, M" journalOrPublisher="5). Archibald Constable, Edinburgh" pageId="0" pageNumber="1" pagination="567 - 584" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" refId="B72" refString="Fleming, J, 1822a. Mollusca. In: Napier, M, Ed., Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol. 5). Archibald Constable, Edinburgh: 567 - 584, https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" title="Mollusca." url="https://babel.hathitrust.org/cgi/pt?id=gri.ark:/13960/t8jd7m43f&amp;view=1up&amp;seq=7" volumeTitle="Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica (Vol." year="1822 a">Fleming 1822a</bibRefCitation>
: 574;
<bibRefCitation author="Fleming, J" journalOrPublisher="Archibald Constable, Edinburgh" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/28230157" refId="B73" refString="Fleming, J, 1822b. The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2). Archibald Constable, Edinburgh, https://www.biodiversitylibrary.org/page/28230157" title="The Philosophy of Zoology: or a General view of the Structure, Functions and Classification of Animals (Vol. 2)." url="https://www.biodiversitylibrary.org/page/28230157" year="1822 b">Fleming 1822b</bibRefCitation>
: 463;
<bibRefCitation author="Lamarck, JB Monet de" journalOrPublisher="Published by the Author, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/13181853" refId="B112" refString="Lamarck, JB Monet de, 1822. Histoire naturelle des animaux sans vertebres. Tome sixieme, 2eme partie. Suite des Gasteropodes. Published by the Author, Paris, https://www.biodiversitylibrary.org/page/13181853" title="Histoire naturelle des animaux sans vertebres. Tome sixieme, 2 eme partie. Suite des Gasteropodes." url="https://www.biodiversitylibrary.org/page/13181853" year="1822">Lamarck 1822</bibRefCitation>
: 46;
<bibRefCitation author="Voigt, FS" journalOrPublisher="Brodhaus, Leipzig" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/19175572" refId="B188" refString="Voigt, FS, 1834. Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken. Brodhaus, Leipzig, https://www.biodiversitylibrary.org/page/19175572" title="Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken." url="https://www.biodiversitylibrary.org/page/19175572" year="1834">Voigt 1834</bibRefCitation>
: 101).
<bibRefCitation author="Oken, L" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/27509779" refId="B139" refString="Oken, L, 1834a. Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283-310. https://biodiversitylibrary.org/page/27509779" title="Voyage de la Corvette l'Astrolabe sous le Commandement de Dumont Durville. Isis von Oken [1834]: 283 - 310." url="https://biodiversitylibrary.org/page/27509779" year="1834 a">Oken (1834a</bibRefCitation>
: 287) transferred Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832)
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
but with no justification.
</paragraph>
<paragraph pageId="0" pageNumber="1">
In the seventh volume of the second edition of
<normalizedToken originalValue="Lamarcks">Lamarck's</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">
Histoire naturelle des animaux sans
<normalizedToken originalValue="vertèbres">vertebres</normalizedToken>
</emphasis>
, which was revised by
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41706785" refId="B63" refString="Deshayes, GP, Milne-Edwards, H, 1836. Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J.B.P.A. de Lamarck. Deuxieme edition, Tome septieme. Histoire des Mollusques. J.-B. Bailliere, Paris, https://www.biodiversitylibrary.org/page/41706785" title="Histoire Naturelle des Animaux sans Vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent, par J. B. P. A. de Lamarck. Deuxieme edition, Tome septieme. Histoire des Mollusques. J. - B." url="https://www.biodiversitylibrary.org/page/41706785" year="1836">Deshayes and Milne-Edwards (1836)</bibRefCitation>
,
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
is proposed as a synonym of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
. However, as a reference for
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
, the authors mentioned the illustration published by Blainville (1827: pl. 46, fig. 7) in the
<emphasis italics="true" pageId="0" pageNumber="1">Atlas</emphasis>
of his
<emphasis italics="true" pageId="0" pageNumber="1">Manuel</emphasis>
which differs from that published by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6, fig. 1) and may or may not refer to
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
John Edward
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: 117) listed
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
and his wife Maria Emma
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: pl. 181, fig. 7) reproduced
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6, fig. 1) original figure of the dorsal notum of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
. As a result, JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: 117) and ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: pl. 181, fig. 7) are listed above as correct references of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. In the same work, JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: 117) regarded
<taxonomicName lsidName="P. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">P. punctata</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
(as spelling mistake
<taxonomicName lsidName="tongensis" pageId="0" pageNumber="1" rank="species" species="tongensis">
<emphasis italics="true" pageId="0" pageNumber="1">tongensis</emphasis>
</taxonomicName>
) as valid, and ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: pl. 182, fig. 1, pl. 183, fig. 3) reproduced the original illustrations by Quoy and Gaimard (1832: pl. 15, figs 17, 18, 27, 28). As a result, those names are also listed above as correct references. According to JE
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: 117),
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
(as
<emphasis italics="true" pageId="0" pageNumber="1">mauriciana</emphasis>
) was a valid
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species name but ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">Gray (1850</bibRefCitation>
: pl. 183, fig. 2) reproduced
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
(1827: pl. 46, fig. 7) illustration which differs from that published by
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">Cuvier (1804</bibRefCitation>
: pl. 6, fig. 1) and which may or may not refer to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
because there are two
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species in Mauritius (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). Therefore, ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">
<normalizedToken originalValue="Grays">Gray's</normalizedToken>
(1850
</bibRefCitation>
: pl. 183, fig. 2)
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
is not listed above as a correct reference of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. And, finally, ME
<bibRefCitation author="Gray, JE" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B86" refString="Gray, JE, 1850. Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." title="Explanation of plates and list of genera. In: Gray ME (Ed.) Figures of molluscous animals, selected from various authors. Etched for the use of students (Vol. 4). Longman, Brown, Green, and Longmans, London, iv + 124 pp." year="1850">
<normalizedToken originalValue="Grays">Gray's</normalizedToken>
(1850
</bibRefCitation>
: pl. 183, figs 4, 4a, 5) reproductions of
<bibRefCitation author="Savigny, JC" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" publicationUrl="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" refId="B157" refString="Savigny, JC, 1817. Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1-3] http://gallica.bnf.fr/ark:/12148/btv1b2300300m" title="Description de l'Egypte ou Recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'Armee francaise, publie par ordre du Gouvernement. Histoire Naturelle, Planches, Tome Deuxieme. Imprimerie Royale, Paris. [105 pls; gastropods on pls 1 - 3]" url="http://gallica.bnf.fr/ark:/12148/btv1b2300300m" year="1817">
<normalizedToken originalValue="Savignys">Savigny's</normalizedToken>
(1817
</bibRefCitation>
: pl. II, figs 3.1-3.3) illustrations of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from the Red Sea do not represent
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(see remarks on
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Adams and Adams (1855: 235) merely listed
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. punctata" pageId="0" pageNumber="1" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">P. punctata</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
as
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species names. Note that for
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, they refer to
<normalizedToken originalValue="Savignys">Savigny's</normalizedToken>
illustrations of individuals from the Red Sea misidentified as
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
by Audouin instead of
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
original description of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, which means that Adams and Adams refer to
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
instead of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(see remarks on
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
). Adams and Adams (1855: pl. LXXXI, fig. 3) also reproduced the original illustration of
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
by Quoy and Gaimard (1832: pl. 15, fig. 17).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge (1855</bibRefCitation>
: 124) mentioned &quot;
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
Cuv.&quot; from the East Indies and the Red Sea but with no new material or literature reference except for a German translation of
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Règne">Regne</normalizedToken>
Animal
</emphasis>
by
<bibRefCitation author="Voigt, FS" journalOrPublisher="Brodhaus, Leipzig" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/19175572" refId="B188" refString="Voigt, FS, 1834. Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken. Brodhaus, Leipzig, https://www.biodiversitylibrary.org/page/19175572" title="Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken." url="https://www.biodiversitylibrary.org/page/19175572" year="1834">Voigt (1834</bibRefCitation>
: 101) as well as &quot;Cuvier,
<normalizedToken originalValue="Règ">Reg</normalizedToken>
. anim. pl. 26, fig. 2.&quot;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge (1855</bibRefCitation>
: 124) followed
<bibRefCitation author="Voigt, FS" journalOrPublisher="Brodhaus, Leipzig" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/19175572" refId="B188" refString="Voigt, FS, 1834. Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken. Brodhaus, Leipzig, https://www.biodiversitylibrary.org/page/19175572" title="Das Thierreich, Geordnet nach Seiner Organisation, Als Grundlage der Naturgeschichte der Thiere und Einleitung in die vergleichende Anatomie. Vom Baron von Cuvier. Dritter Band, die Mollusken." url="https://www.biodiversitylibrary.org/page/19175572" year="1834">Voigt (1834</bibRefCitation>
: 101) and accepted
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
.
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
illustration (pl. 26, fig. 2) mentioned by Berge was actually published after
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
death in the
<normalizedToken originalValue="Disciples">Disciples'</normalizedToken>
edition of the
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Règne">Regne</normalizedToken>
Animal
</emphasis>
which was accompanied by beautiful illustrations. According to
<bibRefCitation DOI="https://doi.org/10.3366/jsbnh.1976.8.1.32" author="Cowan, CF" journalOrPublisher="Journal of the Society for the Bibliography of Natural History" pageId="0" pageNumber="1" pagination="32 - 64" refId="B39" refString="Cowan, CF, 1976. On the Disciples' Edition of Cuvier's Regne Animal. Journal of the Society for the Bibliography of Natural History 8: 32 - 64, DOI: https://doi.org/10.3366/jsbnh.1976.8.1.32" title="On the Disciples' Edition of Cuvier's Regne Animal." url="https://doi.org/10.3366/jsbnh.1976.8.1.32" volume="8" year="1976">Cowan (1976)</bibRefCitation>
, the authorship for the mollusks should be attributed to
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Deterville, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41057481" refId="B61" refString="Deshayes, GP, 1836-1845. Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1-139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot;Le regne animal illustre&quot; or the &quot;Disciples' edition&quot;] https://www.biodiversitylibrary.org/page/41057481" title="Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1 - 139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot; Le regne animal illustre &quot; or the &quot; Disciples' edition &quot;]" url="https://www.biodiversitylibrary.org/page/41057481" year="1836 - 1845">Deshayes (1836-1845)</bibRefCitation>
who prepared the volume of text and the atlas published in livraisons between 1836 and 1845. However, the exact dates of publication are still unknown for most pages and plates, including for the page 69 and the plate 26 where
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
is mentioned and illustrated. Note the spelling mistake
<taxonomicName genus="Unchidium" lsidName="Unchidium" pageId="0" pageNumber="1" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Unchidium</emphasis>
</taxonomicName>
on the figure caption of plate 26. The illustrations in both
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Deterville, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41057481" refId="B61" refString="Deshayes, GP, 1836-1845. Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1-139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot;Le regne animal illustre&quot; or the &quot;Disciples' edition&quot;] https://www.biodiversitylibrary.org/page/41057481" title="Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1 - 139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot; Le regne animal illustre &quot; or the &quot; Disciples' edition &quot;]" url="https://www.biodiversitylibrary.org/page/41057481" year="1836 - 1845">Deshayes (1836-1845</bibRefCitation>
: pl. 26, fig. 2) and
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge (1855</bibRefCitation>
: pl. 16, fig. 8) are mere reproductions of
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6, fig. 5) anatomical drawing of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge (1855</bibRefCitation>
: 124) also mentioned &quot;
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
Quoy&quot; from Australia (as Neuholland) but with no new material or literature reference except for another illustration (pl. 26, fig.1) from the
<normalizedToken originalValue="Disciples">Disciples'</normalizedToken>
edition of the
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Règne">Regne</normalizedToken>
Animal
</emphasis>
. Again, the illustrations in both
<bibRefCitation author="Deshayes, GP" journalOrPublisher="Deterville, Paris" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/41057481" refId="B61" refString="Deshayes, GP, 1836-1845. Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1-139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot;Le regne animal illustre&quot; or the &quot;Disciples' edition&quot;] https://www.biodiversitylibrary.org/page/41057481" title="Les Mollusques. In: Cuvier G (Ed.) Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de planches gravees, representant les types de tous les genres, les caracteres distinctifs des divers groupes et les modifications de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier, MM. Audouin, Blanchard, Deshayes, Alcide d'Orbigny, Doyere, Duges, Duvernoy, Laurillard, Milne Edwards, Roulin et Valenciennes. Fortin, Masson et Cie, Paris, xxxxv + 266 pp. [152 pls; plates are numbered 1 - 139 for an actual total of 152 plates; for a collation see Cowan (1976); often called &quot; Le regne animal illustre &quot; or the &quot; Disciples' edition &quot;]" url="https://www.biodiversitylibrary.org/page/41057481" year="1836 - 1845">Deshayes (1836-1845</bibRefCitation>
: pl. 26, fig. 1) and
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.16004" author="Berge, F" journalOrPublisher="Krais &amp; Hoffmann, Stuttgart" pageId="0" pageNumber="1" refId="B15" refString="Berge, F, 1855. Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren. Krais &amp; Hoffmann, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.16004" title="Conchylienbuch, oder, Allgemeine und besondere Naturgeschichte der Muscheln und Schnecken: nebst der Anweisung sie zu sammeln, zuzubereiten und aufzubewahren." url="https://doi.org/10.5962/bhl.title.16004" year="1855">Berge (1855</bibRefCitation>
: pl. 21, fig. 7) are mere reproductions of Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832: pl. 15, fig. 27) original illustration of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
. It cannot be determined which species Berge referred to exactly (because the localities mentioned by Berge are not the type localities).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The record of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Natal, South Africa (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.13936" author="Krauss, F" journalOrPublisher="Ebner &amp; Seubert, Stuttgart" pageId="0" pageNumber="1" refId="B106" refString="Krauss, F, 1848. Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten. Ebner &amp; Seubert, Stuttgart, DOI: https://doi.org/10.5962/bhl.title.13936" title="Die Suedafrikanischen Mollusken. Ein Beitrag zur Kenntniss der Mollusken des Kap- und Natallandes und zur Geographischen Verbreitung derselben, mit Beschreibung und Abbildung der neuen Arten." url="https://doi.org/10.5962/bhl.title.13936" year="1848">Krauss 1848</bibRefCitation>
: 72) likely is a record of
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, the only
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species known in South Africa so far (see remarks on
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
). However,
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5) could also be present in northeastern South Africa because its southernmost known locality is in Maputo, Mozambique (ca. 26°S). This record by Krauss was mentioned again by a few authors (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12875" author="Sturany, R" journalOrPublisher="Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien" pageId="0" pageNumber="1" refId="B174" refString="Sturany, R, 1898. Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales. Kaiserlich-Koeniglichen Hof- und Staatsdruckerei, Wien, DOI: https://doi.org/10.5962/bhl.title.12875" title="Catalog der bisher bekannt gewordenen Suedafrikanischen Land- und Suesswasser-Mollusken mit besonderer Beruecksichtigung des von Dr Penther gesammelten Materiales." url="https://doi.org/10.5962/bhl.title.12875" year="1898">Sturany 1898</bibRefCitation>
: 73;
<bibRefCitation author="Collinge, WE" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="159 - 174" publicationUrl="https://www.biodiversitylibrary.org/page/6322951" refId="B36" refString="Collinge, WE, 1910. The slugs of Natal. Annals of the Natal Museum 2: 159 - 174, https://www.biodiversitylibrary.org/page/6322951" title="The slugs of Natal." url="https://www.biodiversitylibrary.org/page/6322951" volume="2" year="1910">Collinge 1910</bibRefCitation>
: 171;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly 1912</bibRefCitation>
: 224-225;
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">Connolly 1939</bibRefCitation>
: 454).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.119392" author="Chenu, JC" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="1" refId="B32" refString="Chenu, JC, 1859. Manuel de Conchyliologie et de Paleontologie Conchyliologique (Vol. 1). Masson, Paris, vii + 508 pp. https://doi.org/10.5962/bhl.title.119392" title="Manuel de Conchyliologie et de Paleontologie Conchyliologique (Vol. 1). Masson, Paris, vii + 508 pp." url="https://doi.org/10.5962/bhl.title.119392" year="1859">Chenu (1859</bibRefCitation>
: 474, fig. 3505) mentioned
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia punctata</emphasis>
</taxonomicName>
with no additional material or records, and with a reference to Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832: pl. 15, fig. 27) original illustration of
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Keferstein, WM" editor="Bronn, HG" journalOrPublisher="Bd. 3 (2). Winter, Leipzig &amp; Heidelberg" pageId="0" pageNumber="1" pagination="1160 - 1306" publicationUrl="https://www.biodiversitylibrary.org/page/35742549" refId="B102" refString="Keferstein, WM, 1865a. Lungenschnecken: Pulmonata. In: Bronn, HG, Ed., Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild. Bd. 3(2). Winter, Leipzig &amp; Heidelberg: 1160 - 1306, https://www.biodiversitylibrary.org/page/35742549" title="Lungenschnecken: Pulmonata." url="https://www.biodiversitylibrary.org/page/35742549" volumeTitle="Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild." year="1865 a">Keferstein (1865a</bibRefCitation>
: pl. CII, fig. 20) reproduced Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832: pl. 15, fig. 17) original illustration of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
, which he classified in
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
<bibRefCitation author="Keferstein, WM" editor="Bronn, HG" journalOrPublisher="Bd. 3 (2). Winter, Leipzig &amp; Heidelberg" pageId="0" pageNumber="1" pagination="1160 - 1306" publicationUrl="https://www.biodiversitylibrary.org/page/35742549" refId="B102" refString="Keferstein, WM, 1865a. Lungenschnecken: Pulmonata. In: Bronn, HG, Ed., Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild. Bd. 3(2). Winter, Leipzig &amp; Heidelberg: 1160 - 1306, https://www.biodiversitylibrary.org/page/35742549" title="Lungenschnecken: Pulmonata." url="https://www.biodiversitylibrary.org/page/35742549" volumeTitle="Klassen und Ordnungen der Weichthiere (Malacozoa), wissenschaftlich dargestellt in Wort &amp; Bild." year="1865 a">Keferstein (1865a</bibRefCitation>
: pl. CIII, fig. 1) also reproduced
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804
</bibRefCitation>
: pl. 6, fig. 4) original illustration of the internal anatomy of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, which he also classified in
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on the collections of the Galathea Expedition preserved in Copenhagen,
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
(1872a
</bibRefCitation>
: 28;
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">1872b</bibRefCitation>
: 325) mentioned
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
(as spelling mistake
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="vermiculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vermiculata">
<emphasis italics="true" pageId="0" pageNumber="1">vermiculata</emphasis>
</taxonomicName>
in 1872b) from Pulo Milu [Pulo Milo, Little Nicobar] and Nancouri [Nancowry, Nicobar Islands], where he says it is common. Given the animal size (up to 133 mm long alive), we agree with
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 71) that
<normalizedToken originalValue="Mörchs">Moerch's</normalizedToken>
record is very likely a record of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. The preserved specimen (88/38 mm) reported by
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
(1872a
</bibRefCitation>
: 28;
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">1872b</bibRefCitation>
: 325) from the Galathea collections most likely is the specimen identified by Semper as
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
, described by
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh (1884a</bibRefCitation>
: 142-148, pl. VI, fig. 19, pl. VII, figs 1-6), examined by
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 44) for his description of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, and finally re-examined for the present study (NHMD 613753).
<bibRefCitation DOI="https://doi.org/10.1080/00222938409459805" author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" refId="B17" refString="Bergh, R, 1884b. On the affinities of the Onchidia. Annals and Magazine of Natural History 14(5th series): 259-266. [This is a translation of Bergh (1885)] https://doi.org/10.1080/00222938409459805" title="On the affinities of the Onchidia. Annals and Magazine of Natural History 14 (5 th series): 259 - 266. [This is a translation of Bergh (1885)]" url="https://doi.org/10.1080/00222938409459805" year="1884 b">Bergh (1884b</bibRefCitation>
: 264;
<bibRefCitation author="Bergh, R" journalOrPublisher="Morphologisches Jahrbuch" pageId="0" pageNumber="1" pagination="172 - 181" publicationUrl="https://biodiversitylibrary.org/page/13028090" refId="B18" refString="Bergh, R, 1885. Ueber die Verwandtschaftsbeziehungen der Onchidien. Morphologisches Jahrbuch 10: 172 - 181, https://biodiversitylibrary.org/page/13028090" title="Ueber die Verwandtschaftsbeziehungen der Onchidien." url="https://biodiversitylibrary.org/page/13028090" volume="10" year="1885">1885</bibRefCitation>
: 177) briefly mentioned
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
again in a comparative study on the affinities of onchidiids.
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
(1872a
</bibRefCitation>
: 28;
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">1872b</bibRefCitation>
: 325) mentioned
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
from Sambelong, Great Nicobar, Nicobar Islands, based on much smaller individuals from the collections of the Galathea Expedition in Copenhagen, which are a record for
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
(see remarks on that species).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The record of
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium mauritianum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritianum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium mauritianum</emphasis>
</taxonomicName>
from the Red Sea by
<bibRefCitation author="Pagenstecher, HA" journalOrPublisher="Engelmann, Leipzig" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/12426731" refId="B141" refString="Pagenstecher, HA, 1877. Mollusca. In: Kossmann R (Ed.) Zoologische Ergebnisse einer im Auftraege der Koeniglichen Academie der Wissenschaften zu Berlin ausgefuehrten Reise in die Kuestengebiete des Rothen Meeres, Erste Haelfte. W. Engelmann, Leipzig, https://biodiversitylibrary.org/page/12426731" title="Mollusca. In: Kossmann R (Ed.) Zoologische Ergebnisse einer im Auftraege der Koeniglichen Academie der Wissenschaften zu Berlin ausgefuehrten Reise in die Kuestengebiete des Rothen Meeres, Erste Haelfte. W." url="https://biodiversitylibrary.org/page/12426731" year="1877">Pagenstecher (1877</bibRefCitation>
: 62) refers to either
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
or
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The records of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Mozambique by
<bibRefCitation author="Martens, E von" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/36341972" refId="B124" refString="Martens, E von, 1879. Uebersicht der von W. Peters von 1843 bis 1847 in Mossambique gesammelten Mollusca. Monatsberichte der koeniglich Preussischen Akademie der Wissenschaften zu Berlin [1879]: 727-749. https://biodiversitylibrary.org/page/36341972" title="Uebersicht der von W. Peters von 1843 bis 1847 in Mossambique gesammelten Mollusca. Monatsberichte der koeniglich Preussischen Akademie der Wissenschaften zu Berlin [1879]: 727 - 749." url="https://biodiversitylibrary.org/page/36341972" year="1879">Martens (1879</bibRefCitation>
: 735) in Ibo Island (ca. 12°21'S) and Inhambane (ca. 23°52'S) are within the geographical range of both
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5) and
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). It is not possible to know to what species Martens was referring; this record by Martens was mentioned twice by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly (1912</bibRefCitation>
: 225;
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">1939</bibRefCitation>
: 454).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Semper (1880: 258-260, pl. XIX, figs 2, 9, pl. XXII, figs 1, 2, 10) referred to huge onchidiid slugs (from 50 to 105 mm, preserved) as Quoy and
<normalizedToken originalValue="Gaimards">Gaimard's</normalizedToken>
(1832)
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
and merely suggested, with a question mark, that
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
could refer to the same species. Semper (1880: 258) listed five geographical records for
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
: Tonga and West Papua (as Port Dorey), from Quoy and Gaimard (1832); Mauritius, based on some material from the Vienna and Kiel museums; Samoa, based on some material from the Museum Godeffroy; and Bohol, Philippines, based on his own collections. Semper (1880: 258) indicated that the specimens he examined were from 50 to 105 mm long, preserved, and that the smallest individual was found in Mauritius. His anatomical description perfectly matches the anatomy of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. In particular, a spine of an accessory penial gland measuring 4 mm long is only compatible with
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). However, he did not clearly indicate whether he observed a long spine in every specimen. It cannot be excluded that he measured the length of the accessory penial gland spine only in a specimen from Mauritius. Therefore, the records of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in Bohol and in Samoa are regarded here as questionable, even though it is very possible that
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
lives in Samoa, given that it is so close to Tonga (800 km) and Fiji (1100 km).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Semper (1882: 290) thought that
<bibRefCitation author="Cuvier, G" journalOrPublisher="Annales du Museum national d'Histoire naturelle" pageId="0" pageNumber="1" pagination="37 - 51" publicationUrl="https://www.biodiversitylibrary.org/page/48531138" refId="B43" refString="Cuvier, G, 1804. Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii. Annales du Museum national d'Histoire naturelle 5: 37 - 51, https://www.biodiversitylibrary.org/page/48531138" title="Memoire sur l'Oncidie, genre de mollusques nuds, voisin des limaces, et sur une espece nouvelle, Onchidium peronii." url="https://www.biodiversitylibrary.org/page/48531138" volume="5" year="1804">
<normalizedToken originalValue="Cuviers">Cuvier's</normalizedToken>
(1804)
</bibRefCitation>
original description of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
was problematic because his drawing of the dorsal notum did not match the internal anatomy. Because Semper was convinced that Cuvier used specimens that did not belong to the same species, he thought that the name
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
should not be used.
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893</bibRefCitation>
: 172-173) disagreed with Semper even without examining the type material of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. It is demonstrated here that the two type specimens described by Cuvier as
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
both belong to the same species (see above). Semper (1882: 268) was undecided about the nomenclatural status of what he called
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium mauritianum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritianum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium mauritianum</emphasis>
</taxonomicName>
(then a new combination), which he listed as one of the names for which a &quot;closer inspection of the originals&quot; was needed. Like most authors, he cited
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
(1827: pl. 46, fig. 7) illustration (which is not part of
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
original description) as a reference without realizing that it may or may not correspond to
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
, a junior objective synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. Semper (1882: 289) was also undecided about the status of
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
, for which he erroneously thought that the type locality was unknown. He suggested that it might refer to the same species as
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tumidum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tumidum</emphasis>
</taxonomicName>
, which is not possible because
<taxonomicName lsidName="O. tumidum" pageId="0" pageNumber="1" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tumidum</emphasis>
</taxonomicName>
was recently transferred to
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paromoionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paromoionchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paromoionchis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2019.500" author="Dayrat, B" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="1 - 77" refId="B55" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Bourke, AJ, Comendador, J, Tan, SH, 2019a. A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1 - 77, DOI: https://doi.org/10.5852/ejt.2019.500" title="A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." url="https://doi.org/10.5852/ejt.2019.500" volume="500" year="2019 a">Dayrat et al. 2019a</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Tapparone Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="5 - 313" publicationUrl="https://www.biodiversitylibrary.org/page/10812598" refId="B183" refString="Tapparone Canefri, C, 1883. Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini. Annali del Museo Civico di Storia Naturale di Genova 19: 5 - 313, https://www.biodiversitylibrary.org/page/10812598" title="Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini." url="https://www.biodiversitylibrary.org/page/10812598" volume="19" year="1883">Tapparone Canefri (1883</bibRefCitation>
: 214) listed all of
<normalizedToken originalValue="Sempers">Semper's</normalizedToken>
geographic records for
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia tongana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia tongana</emphasis>
</taxonomicName>
with no new material or anatomical observations (see above).
<bibRefCitation author="Tapparone Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="5 - 313" publicationUrl="https://www.biodiversitylibrary.org/page/10812598" refId="B183" refString="Tapparone Canefri, C, 1883. Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini. Annali del Museo Civico di Storia Naturale di Genova 19: 5 - 313, https://www.biodiversitylibrary.org/page/10812598" title="Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini." url="https://www.biodiversitylibrary.org/page/10812598" volume="19" year="1883">Tapparone Canefri (1883</bibRefCitation>
: 214) also regarded
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia punctata</emphasis>
</taxonomicName>
as a valid species name, but with no other reference or material than the original description by Quoy and Gaimard (1832). Tapparone
<normalizedToken originalValue="Canefris">Canefri's</normalizedToken>
suggestion that
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia punctata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia punctata</emphasis>
</taxonomicName>
could refer to the same species as
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tumidum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tumidum</emphasis>
</taxonomicName>
must be rejected because
<taxonomicName lsidName="O. tumidum" pageId="0" pageNumber="1" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tumidum</emphasis>
</taxonomicName>
was recently transferred to
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paromoionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paromoionchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paromoionchis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2019.500" author="Dayrat, B" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="1 - 77" refId="B55" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Bourke, AJ, Comendador, J, Tan, SH, 2019a. A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1 - 77, DOI: https://doi.org/10.5852/ejt.2019.500" title="A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." url="https://doi.org/10.5852/ejt.2019.500" volume="500" year="2019 a">Dayrat et al. 2019a</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Smith, EA" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://www.biodiversitylibrary.org/item/63054" refId="B165" refString="Smith, EA, 1884. Mollusca. In: [no named editor] Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881-2. Part I. The collections from Melanesia; Part II. Collections from the western Indian Ocean. The Trustees of the British Museum, London, 34-116, 487-508. http://www.biodiversitylibrary.org/item/63054" title="Mollusca. In: [no named editor] Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 2. Part I. The collections from Melanesia; Part II. Collections from the western Indian Ocean. The Trustees of the British Museum, London, 34 - 116, 487 - 508." url="http://www.biodiversitylibrary.org/item/63054" year="1884">Smith (1884</bibRefCitation>
: 92) mentioned
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium (Peronia) punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum" subGenus="Peronia">Onchidium (Peronia) punctatum</taxonomicName>
from Albany Island and Thursday Island, in the Torres Strait, without any description. This is likely a record of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
(unit #1), the only species thought to be present in the Torres Strait, although our study does not include any fresh material from the Torres Strait and
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
could also live there (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). Note that Thursday Island also happens to be the type locality of
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Scaphis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scaphis viridis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="viridis">
<emphasis italics="true" pageId="0" pageNumber="1">Scaphis viridis</emphasis>
</taxonomicName>
, a junior synonym of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">Bergh (1884a</bibRefCitation>
: 142-148, pl. VI, fig. 19, pl. VII, figs 1-6) described
<emphasis italics="true" pageId="0" pageNumber="1">
as
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">Onchidium tonganum</taxonomicName>
</emphasis>
a specimen from the collections of the Copenhagen Museum which was collected in the Nicobar Islands during the Galathea Expedition (station 305). That specimen (85/55 mm), dissected by Bergh, is in a jar (NHMD 613753) with a second specimen (70/50 mm) which is still entire and not dissected by Bergh. Both specimens were re-examined for the present study, although
<normalizedToken originalValue="Berghs">Bergh's</normalizedToken>
measurement of the penial gland spine in the largest specimen (4.25 mm) could not be checked because internal organs are missing. Given the specimen sizes, their intestinal loops (type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
in the second specimen), and the spine of their accessory penial glands (4.25 mm in the largest specimen according to Bergh, and 4 mm in the second specimen), those two specimens belong to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Joyeux-Laffuie, J" journalOrPublisher="Journal of the Society for the Bibliography of Natural History" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/44109240" refId="B100" refString="Joyeux-Laffuie, J, 1885. Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884b)] https://www.biodiversitylibrary.org/page/44109240" title="Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884 b)]" url="https://www.biodiversitylibrary.org/page/44109240" year="1885">Joyeux-Laffuie (1885</bibRefCitation>
: viii-xi) merely mentioned
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
in a summary of
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">
<normalizedToken originalValue="Berghs">Bergh's</normalizedToken>
(1884a)
</bibRefCitation>
work.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893</bibRefCitation>
: 172-173, pl. 12, figs 85, 87, 91) re-described
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
based on at least one specimen from Mauritius for which he did not provide any size. However, given the length of the spine of the accessory penial gland (7 mm long), there is no doubt that he examined
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. It is possible that he included a part of the duct of the accessory penial gland in that measurement because the longest spine observed in the present study was 5 mm, in the lectotype of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
(MNHN-IM-2000-33692). According to Plate, the retractor muscle inserts near the central nervous system, which does fit in the variation observed here (Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Plate listed several synonyms:
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
, and possibly (with a question mark)
<taxonomicName lsidName="P. corpulenta" pageId="0" pageNumber="1" rank="species" species="corpulenta">
<emphasis italics="true" pageId="0" pageNumber="1">P. corpulenta</emphasis>
</taxonomicName>
. Note that
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893</bibRefCitation>
: 172) rightly regarded
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
but for a weak reason (a similar pigmentation of the lung). These synonymies are all accepted here, except for
<taxonomicName lsidName="P. corpulenta" pageId="0" pageNumber="1" rank="species" species="corpulenta">
<emphasis italics="true" pageId="0" pageNumber="1">P. corpulenta</emphasis>
</taxonomicName>
which is regarded as a
<emphasis italics="true" pageId="0" pageNumber="1">nomen dubium</emphasis>
(see general discussion).
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893)</bibRefCitation>
did not comment on
<taxonomicName lsidName="O. punctatum" pageId="0" pageNumber="1" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. punctatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Annual Reviews of Ecology, Evolution and Systematics" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/30983429" refId="B78" refString="Godwin-Austen, HH, 1895. List and Distribution of the Land-Mollusca of the Andaman and Nicobar Islands, with Descriptions of some supposed new Species. Proceedings of the Zoological Society of London [1895]: 438-456. https://www.biodiversitylibrary.org/page/30983429" title="List and Distribution of the Land-Mollusca of the Andaman and Nicobar Islands, with Descriptions of some supposed new Species. Proceedings of the Zoological Society of London [1895]: 438 - 456." url="https://www.biodiversitylibrary.org/page/30983429" year="1895">Godwin-Austen (1895</bibRefCitation>
: 443) listed
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium mauritianum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritianum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium mauritianum</emphasis>
</taxonomicName>
in Little Nicobar. It is impossible to know what species was referred to. However,
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(of which
<emphasis italics="true" pageId="0" pageNumber="1">
P
<taxonomicName lsidName="mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">mauritiana</taxonomicName>
</emphasis>
is a junior synonym), is known to be present in Nicobar Islands.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12547" author="Casto de Elera, RPF" journalOrPublisher="Colegio de Santo Tomas, Manila" pageId="0" pageNumber="1" refId="B30" refString="Casto de Elera, RPF, 1896. Catalogo sistematico de toda la Fauna de Filipinas conocida hasta el presente, y a la vez el de la coleccion zoologica del museo de PP. Dominicos del Colegio-Universidad de Sto. Tomas de Manila (Vol. 3). Colegio de Santo Tomas, Manila, DOI: https://doi.org/10.5962/bhl.title.12547" title="Catalogo sistematico de toda la Fauna de Filipinas conocida hasta el presente, y a la vez el de la coleccion zoologica del museo de PP. Dominicos del Colegio-Universidad de Sto. Tomas de Manila (Vol. 3)." url="https://doi.org/10.5962/bhl.title.12547" year="1896">Casto de Elera (1896</bibRefCitation>
: 629) mentioned the presence of several species in the Philippines, without description or new material, mostly based on
<bibRefCitation author="Semper, C" editor="Semper, C" journalOrPublisher="Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251 - 264 [1880]" pageId="0" pageNumber="1" pagination="265 - 290" publicationUrl="https://biodiversitylibrary.org/" refId="B159" refString="Semper, C, 1880-1885. Dritte Familie, Onchididae [for Onchidiidae]. In: Semper, C, Ed., Reisen im Archipel der Philippinen, Zweiter Theil. Wissenschaftliche Resultate. Dritter Band. Landmollusken. CW Kreidel's Verlag, Wiesbaden, 251-264 [1880]: 265 - 290, https://biodiversitylibrary.org/" title="Dritte Familie, Onchididae [for Onchidiidae]." url="https://biodiversitylibrary.org/" volumeTitle="Reisen im Archipel der Philippinen, Zweiter Theil." year="1880 - 1885">
<normalizedToken originalValue="Sempers">Semper's</normalizedToken>
(1880-1885)
</bibRefCitation>
work:
<taxonomicName authorityName="Cuvier" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium verruculatum</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="O. savignyi" pageId="0" pageNumber="1" rank="species" species="savignyi">
<emphasis italics="true" pageId="0" pageNumber="1">O. savignyi</emphasis>
</taxonomicName>
(as
<emphasis italics="true" pageId="0" pageNumber="1">savigngi</emphasis>
). Our data suggest that
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
(unit #1) lives in the Philippines (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.46994" author="Martens, E von" journalOrPublisher="Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien" pageId="0" pageNumber="1" pagination="1 - 332" refId="B125" refString="Martens, E von, 1897. Suess- und brackwasser-mollusken des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien 4: 1 - 332, DOI: https://doi.org/10.5962/bhl.title.46994" title="Suess- und brackwasser-mollusken des Indischen Archipels." url="https://doi.org/10.5962/bhl.title.46994" volume="4" year="1897">Martens (1897</bibRefCitation>
: 126) listed
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
with a reference to
<normalizedToken originalValue="Blainvilles">Blainville's</normalizedToken>
(1827: pl. 46, fig. 7) illustration of
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
which may or may not refer to
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.46994" author="Martens, E von" journalOrPublisher="Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien" pageId="0" pageNumber="1" pagination="1 - 332" refId="B125" refString="Martens, E von, 1897. Suess- und brackwasser-mollusken des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien 4: 1 - 332, DOI: https://doi.org/10.5962/bhl.title.46994" title="Suess- und brackwasser-mollusken des Indischen Archipels." url="https://doi.org/10.5962/bhl.title.46994" volume="4" year="1897">Martens (1897</bibRefCitation>
: 126) also claimed that
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
, with a reference to the original description by Quoy and Gaimard (1832) as well as to
<normalizedToken originalValue="Sempers">Semper's</normalizedToken>
(1880: 258-260, pl. XIX, figs 2, 9, pl. XXII, figs 1, 2, 10) re-description, which may only partly correspond to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(see above). The record of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
in Mombasa (Mombas,
<normalizedToken originalValue="Ostküste">Ostkueste</normalizedToken>
Afrika) by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.46994" author="Martens, E von" journalOrPublisher="Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien" pageId="0" pageNumber="1" pagination="1 - 332" refId="B125" refString="Martens, E von, 1897. Suess- und brackwasser-mollusken des Indischen Archipels. Zoologische Ergebnisse einer Reise in Niederlaendisch Ost-Indien 4: 1 - 332, DOI: https://doi.org/10.5962/bhl.title.46994" title="Suess- und brackwasser-mollusken des Indischen Archipels." url="https://doi.org/10.5962/bhl.title.46994" volume="4" year="1897">Martens (1897</bibRefCitation>
: 126) is not supported by any description and is therefore not accepted here, even though it is possibly correct (
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is present in Zanzibar).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Collinge, WE" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="1 - 8" publicationUrl="https://www.biodiversitylibrary.org/page/30026709" refId="B35" refString="Collinge, WE, 1900. A collection of slugs from South Africa, with descriptions of some new species. Annals of the South African Museum 2: 1 - 8, https://www.biodiversitylibrary.org/page/30026709" title="A collection of slugs from South Africa, with descriptions of some new species." url="https://www.biodiversitylibrary.org/page/30026709" volume="2" year="1900">Collinge (1900</bibRefCitation>
: 7) and
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly (1912</bibRefCitation>
: 225) mentioned
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Green Point, Cape Peninsula, South Africa. Those specimens were later used by
<bibRefCitation author="Watson, H" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="237 - 308" publicationUrl="https://www.biodiversitylibrary.org/page/40651328" refId="B192" refString="Watson, H, 1925. The South African species of the molluscan genus Onchidella. Annals of the South African Museum 20: 237 - 308, https://www.biodiversitylibrary.org/page/40651328" title="The South African species of the molluscan genus Onchidella." url="https://www.biodiversitylibrary.org/page/40651328" volume="20" year="1925">Watson (1925</bibRefCitation>
: 283-284, pl. XXVII, figs 4-11, pl. XXVIII, figs 12, 14, pl. XXXI, fig. 58) to describe
<taxonomicName authorityName="H.M.Watson" authorityYear="1925" class="Gastropoda" family="Onchidiidae" genus="Onchidella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidella capensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="capensis">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidella capensis</emphasis>
</taxonomicName>
Watson, 1925.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Sturany, R" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe" pageId="0" pageNumber="1" pagination="209 - 283" publicationUrl="https://biodiversitylibrary.org/page/6903441" refId="B175" refString="Sturany, R, 1904. Expeditionen S.M. Schiff &quot;Pola&quot; in das Rote Meer. Noerdliche und suedliche Haelfte. 1895/96-1897/98. Zoologische Ergebnisse. XXIII. Gastropoden des Rothen Meeres. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Classe 74: 209 - 283, https://biodiversitylibrary.org/page/6903441" title="Expeditionen S. M. Schiff &quot; Pola &quot; in das Rote Meer. Noerdliche und suedliche Haelfte. 1895 / 96 - 1897 / 98. Zoologische Ergebnisse. XXIII. Gastropoden des Rothen Meeres." url="https://biodiversitylibrary.org/page/6903441" volume="74" year="1904">Sturany (1904</bibRefCitation>
: 269) mentioned the presence of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
in Massawa, Eritrea, Red Sea. This record most likely refers to either
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia verruculata" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia verruculata</emphasis>
</taxonomicName>
or
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
is one of the eight onchidiid species mentioned by
<bibRefCitation author="Hedley, C" journalOrPublisher="Report of the Australian Association for the advancement of Science" pageId="0" pageNumber="1" pagination="329 - 371" refId="B91" refString="Hedley, C, 1909. The marine fauna of Queensland. Report of the Australian Association for the advancement of Science 12: 329 - 371" title="The marine fauna of Queensland." volume="12" year="1909">Hedley (1909</bibRefCitation>
: 369) from Queensland, Australia, without any reference to any material. It is impossible to know what species Hedley refers to. Our data show that there are two
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species in Queensland (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The record of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Durban, Natal, South Africa by
<bibRefCitation author="Collinge, WE" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="159 - 174" publicationUrl="https://www.biodiversitylibrary.org/page/6322951" refId="B36" refString="Collinge, WE, 1910. The slugs of Natal. Annals of the Natal Museum 2: 159 - 174, https://www.biodiversitylibrary.org/page/6322951" title="The slugs of Natal." url="https://www.biodiversitylibrary.org/page/6322951" volume="2" year="1910">Collinge (1910</bibRefCitation>
: 171) likely is a record of
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, the only
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species known in South Africa so far (see remarks on
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
). However,
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5) could also be present in northeastern South Africa because its southernmost known locality is in Maputo, Mozambique (ca. 26°S). This record was mentioned again by
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly (1912</bibRefCitation>
: 225;
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">1939</bibRefCitation>
: 454).
</paragraph>
<paragraph pageId="0" pageNumber="1">
According to
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.11477" author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" pagination="59 - 306" refId="B37" refString="Connolly, M, 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306, DOI: https://doi.org/10.5962/bhl.title.11477" title="A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum." url="https://doi.org/10.5962/bhl.title.11477" volume="11" year="1912">Connolly (1912</bibRefCitation>
: 224-225),
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
is a valid name and
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
(as spelling mistake
<emphasis italics="true" pageId="0" pageNumber="1">mauritziana</emphasis>
) and
<taxonomicName lsidName="O. tonganum" pageId="0" pageNumber="1" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tonganum</emphasis>
</taxonomicName>
are its synonyms. The references listed by Connolly are all commented on above. Let us briefly emphasize here, however, that the localities of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
mentioned by Connolly in South Africa and Mozambique are problematic.
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">Connolly (1939</bibRefCitation>
: 454) later admitted that &quot;it is open to question (...) whether the true
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
Cuv. really exists in South Africa.&quot;
<bibRefCitation author="Connolly, M" journalOrPublisher="Annals of the South African Museum" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/40865351" refId="B38" refString="Connolly, M, 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1-660. [pls I-XIX] https://www.biodiversitylibrary.org/page/40865351" title="A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: i-iii, 1 - 660. [pls I-XIX]" url="https://www.biodiversitylibrary.org/page/40865351" year="1939">Connolly (1939</bibRefCitation>
:453), who did not cite
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbés">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
work, considered that
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
was a subgenus of
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium</emphasis>
</taxonomicName>
and should include onchidiid slugs with dorsal gills.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Vayssiere, A" journalOrPublisher="Annales de la Faculte des Sciences de l'Universite de Marseille, Supplement" pageId="0" pageNumber="1" pagination="5 - 157" publicationUrl="https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" refId="B187" refString="Vayssiere, A, 1912. Recherches zoologiques et anatomiques sur les Opisthobranches de la Mer Rouge et du Golfe d'Aden. Deuxieme Partie. Annales de la Faculte des Sciences de l'Universite de Marseille, Supplement 20: 5 - 157, https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" title="Recherches zoologiques et anatomiques sur les Opisthobranches de la Mer Rouge et du Golfe d'Aden. Deuxieme Partie." url="https://odyssee.univ-amu.fr/files/original/2/18/Annales-faculte-sc-Mrs_1912_T-20_Suppl.pdf" volume="20" year="1912">
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
(1912
</bibRefCitation>
: 125-129) recorded seven individuals of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
shipped to him from Moucha Islands (Djibouti) by Charles Gravier and
<normalizedToken originalValue="Félix">Felix</normalizedToken>
Pierre Jousseaume, two of the people who also collected many specimens studied by
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a)
</bibRefCitation>
.
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
mostly focused on the description of the radula, which is not useful to identify species.
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
reported a wide range of animal sizes (from 10 to 80 mm long and from 6 to 60 mm wide). Thus, it is very possible that he examined more than one species. Instead of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, which has never been positively recorded from Djibouti,
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
likely examined
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, or both (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). His specimens of large size most likely were
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
because
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
individuals rarely are longer than 60 mm (Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Note that the number of rows of teeth and the number of teeth per half row mentioned by
<normalizedToken originalValue="Vayssière">Vayssiere</normalizedToken>
(95 to 100 rows on average) are higher than what was observed here, although they are more compatible with
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
than
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Radular formulae in Peronia species. Each formula follows the same format: number of rows x number of lateral teeth per left half row - 1 (rachidian tooth) - number of lateral teeth per right half row. Each DNA extraction number corresponds to one individual. The letter H next to an extraction number indicates a holotype." httpUri="http://table.plazi.org/id/F6A603768F442AB5CA201CDCEC46CA6A" pageId="0" pageNumber="1" tableUuid="F6A603768F442AB5CA201CDCEC46CA6A">5</tableCitation>
), acknowledging that radular formulae are expected to vary.
</paragraph>
<paragraph pageId="0" pageNumber="1">
It is not possible to determine to what species
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.789" author="Odhner, NJ" journalOrPublisher="Arkiv foer Zoologi" pageId="0" pageNumber="1" pagination="1 - 52" refId="B136" refString="Odhner, NJ, 1919. Contribution a la faune malacologique de Madagascar. Arkiv foer Zoologi 12: 1 - 52, DOI: https://doi.org/10.5962/bhl.part.789" title="Contribution a la faune malacologique de Madagascar." url="https://doi.org/10.5962/bhl.part.789" volume="12" year="1919">Odhner (1919</bibRefCitation>
: 42) was referring solely based on his brief, external description of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Toliara, Madagascar. However, his material, dissected here, clearly belongs to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
: a single individual (65/50 mm) is characterized by intestinal loops of type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
, a spine of the accessory penial gland of 3 mm long, and a muscular sac of 25 mm long (SMNH 180381).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919)</bibRefCitation>
uncritically took for granted every species record ever published, without considering that species often are misidentified.
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919)</bibRefCitation>
accepted
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
as a valid name, with
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
as synonyms, and
<taxonomicName lsidName="P. corpulenta" pageId="0" pageNumber="1" rank="species" species="corpulenta">
<emphasis italics="true" pageId="0" pageNumber="1">P. corpulenta</emphasis>
</taxonomicName>
as a potential synonym (with a question mark). The references listed by
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919</bibRefCitation>
: 311-312) for
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
are all commented on above already. However,
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">
<normalizedToken originalValue="Bretnalls">Bretnall's</normalizedToken>
(1919
</bibRefCitation>
: 313) list of geographic records needs to be discussed, especially because Bretnall did not mention the key characters supporting a proper identification of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). The presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in Samoa, which Bretnall obtained from Semper (see above), should not be taken for granted, even if it is quite possible. The presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in the Buccaneer Archipelago, northern Western Australia (16S, 123E), based on specimens from the Australian Museum, should not be taken for granted, even though it is quite possible. The identification of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in the Santa Cruz Islands, Solomon Islands, based on specimens from the Australian Museum, should not be taken for granted (specimens may have been misidentified), even if the Santa Cruz Islands are within the known geographical range of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919</bibRefCitation>
: 315-316) also regarded
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
as a valid name, for which he cited
<bibRefCitation author="Bergh, R" journalOrPublisher="Kreidel, Wiesbaden" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/49395587" refId="B16" refString="Bergh, R, 1884a. Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H.M.S. Challenger during the years 1873-76. Zoology, Band 10, Theil 26. C.W. Kreidel, Wiesbaden, https://biodiversitylibrary.org/page/49395587" title="Report on Nudibranchiata. Report of the Scientific Results of the Voyage of the H. M. S. Challenger during the years 1873 - 76. Zoology, Band 10, Theil 26. C. W." url="https://biodiversitylibrary.org/page/49395587" year="1884 a">
<normalizedToken originalValue="Berghs">Bergh's</normalizedToken>
(1884a)
</bibRefCitation>
original description and its French summary by
<bibRefCitation author="Joyeux-Laffuie, J" journalOrPublisher="Journal of the Society for the Bibliography of Natural History" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/44109240" refId="B100" refString="Joyeux-Laffuie, J, 1885. Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884b)] https://www.biodiversitylibrary.org/page/44109240" title="Sur les affinites des Onchidies, par R. Bergh de Copenhague. Archives de Zoologie Experimentale et Generale 3: viii-x. [translation and summary of Bergh's (1884 b)]" url="https://www.biodiversitylibrary.org/page/44109240" year="1885">
<normalizedToken originalValue="Joyeux-Laffuies">Joyeux-Laffuie's</normalizedToken>
(1885)
</bibRefCitation>
, and indicated
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">
<normalizedToken originalValue="Plates">Plate's</normalizedToken>
(1893)
</bibRefCitation>
proposed synonymy (with
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
) with a question mark.
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919</bibRefCitation>
: 316) listed Lord Howe Island, off southeastern Australia (based on specimens from the Australian Museum), as a locality for
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
, but without description of key characters. Thus, the presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in Lord Howe Island, which is 1350 km south of the southernmost known locality of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(New Caledonia), is not taken for granted here. As for
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium punctatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="punctatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium punctatum</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919</bibRefCitation>
: 316-317) followed Semper (1882: 289) and
<bibRefCitation author="Tapparone Canefri, C" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="1" pagination="5 - 313" publicationUrl="https://www.biodiversitylibrary.org/page/10812598" refId="B183" refString="Tapparone Canefri, C, 1883. Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini. Annali del Museo Civico di Storia Naturale di Genova 19: 5 - 313, https://www.biodiversitylibrary.org/page/10812598" title="Fauna malacologica della Nuova Guinea e delle isole adiacenti. Parte I. Molluschi estramarini." url="https://www.biodiversitylibrary.org/page/10812598" volume="19" year="1883">Tapparone Canefri (1883</bibRefCitation>
: 214) who both thought that it could be a synonym of
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tumidum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tumidum</emphasis>
</taxonomicName>
(see above), which is not possible because
<taxonomicName lsidName="O. tumidum" pageId="0" pageNumber="1" rank="species" species="tumidum">
<emphasis italics="true" pageId="0" pageNumber="1">O. tumidum</emphasis>
</taxonomicName>
refers to a
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paromoionchis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paromoionchis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Paromoionchis</emphasis>
</taxonomicName>
species (
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2019.500" author="Dayrat, B" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="1" pagination="1 - 77" refId="B55" refString="Dayrat, B, Goulding, TC, Khalil, M, Apte, D, Bourke, AJ, Comendador, J, Tan, SH, 2019a. A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae). European Journal of Taxonomy 500: 1 - 77, DOI: https://doi.org/10.5852/ejt.2019.500" title="A new genus and three new species of mangrove slugs from the Indo-West Pacific (Gastropoda: Euthyneura: Onchidiidae)." url="https://doi.org/10.5852/ejt.2019.500" volume="500" year="2019 a">Dayrat et al. 2019a</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 71), following most of
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">
<normalizedToken originalValue="Plates">Plate's</normalizedToken>
(1893)
</bibRefCitation>
nomenclatural decisions, accepted
<taxonomicName authorityName="Blainville" authorityYear="1824" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia mauritiana" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia mauritiana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. corpulenta" pageId="0" pageNumber="1" rank="species" species="corpulenta">
<emphasis italics="true" pageId="0" pageNumber="1">P. corpulenta</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
as junior synonyms of
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. Hoffmann, like other authors, did not mention the key anatomical characters that allow a reliable identification of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
and uncritically accepted most geographical records published before him. As a result, his proposed distribution for
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
should not be taken for granted. For instance, the presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in Lord Howe Island, off southeastern Australia, obtained from
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall (1919)</bibRefCitation>
is questionable.
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 44) examined a specimen from the Nicobar Islands (NHMD 613753) which was originally mentioned by
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/35774183" refId="B130" refString="Moerch, OAL, 1872a. Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9-35. https://biodiversitylibrary.org/page/35774183" title="Synopsis Molluscorum a cl. J. Reinhardt lectorum in circumnavigatione orbis terrarum Danicae navis Galatheae dictae. Oversigt over de af Prof. J. Reinhardt paa Corvetten Galatheas Reise omkring Jorden indsamlede Bloeddyr. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjoebenhavn [1872]: 9 - 35." url="https://biodiversitylibrary.org/page/35774183" year="1872 a">
<normalizedToken originalValue="Mörch">Moerch</normalizedToken>
(1872a
</bibRefCitation>
: 28;
<bibRefCitation author="Moerch, OAL" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="303 - 327" publicationUrl="https://biodiversitylibrary.org/page/15684513" refId="B131" refString="Moerch, OAL, 1872b. Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale. Journal de Conchyliologie 20: 303 - 327, https://biodiversitylibrary.org/page/15684513" title="Catalogue de Mollusques terrestres et fluviatiles des anciennes colonies danoises du golfe du Bengale." url="https://biodiversitylibrary.org/page/15684513" volume="20" year="1872 b">1872b</bibRefCitation>
: 325; see above).
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann (1928</bibRefCitation>
: 44-45) also provided several geographical records (Sumatra, Java, Marshall Islands, Kiribati, Fiji) for
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
based on material preserved at the SMNH in Stockholm. His material was re-examined and all records are confirmed. Hoffmann only dissected two individuals, one from Sumatra (SMNH 180354) and one from Kiribati (SMNH 180379). The other eighteen specimens were dissected for the present study.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Eight large specimens (longer than 65 mm) examined by Hoffmann from Sumatra (SMNH 180354), Java (SMNH 180355), Kiribati (SMNH 180376, 180377, 180380, 180382, 180475), and Fiji (SMNH 180373) share the diagnostic characteristics of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
: a spine of the accessory penial gland between 3 and 4 mm long, intestinal loops of type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
, and a muscular sac between 20 and 25 mm long (exceptionally 17 mm, SMNH 180354). Seven smaller specimens (between 15 and 37 mm long) examined by Hoffmann from the Marshall Islands (SMNH 180356), Fiji (SMNH 180374), and Kiribati (SMNH 180353, 180383, 180384) are immature: the anterior male reproductive parts are barely developed, and, if present, the spine of the accessory penial gland is still soft (SMNH 180353). Given their intestinal loops (type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
), they are regarded as individuals of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. In other species, individuals of that size are already fully mature. Two smaller specimens (between 28 and 37 mm long) examined by Hoffmann from Fiji (SMNH 180357, 180375) belong to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
because of several characteristics (retractor muscle inserting near the heart, intestinal loops of type I with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
, a spine of 3 mm long). Their muscular sacs (11 and 15 mm) are shorter than in other specimens, suggesting that they likely are not fully mature. Two specimens from Kiribati (SMNH 180378, 180478), poorly preserved, could not be confidently identified. Finally, the male reproductive parts are missing in a specimen from Kiribati dissected by Hoffmann (SMNH 180379), but its intestinal loops (type I with a transitional loop oriented between 1 and 2
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
) confirm that it belongs to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Note that the locality of the specimen from Sumatra (SMNH 180354) is problematic. The label and
<normalizedToken originalValue="Hoffmanns">Hoffmann's</normalizedToken>
publication both say &quot;Pulu Pasu, west coast of Sumatra,&quot; but there is no such place on the west coast of Sumatra. There are two small islands off the west coast of Sumatra called Pulau Asu (Hinako Islands) and Pulau Pasumpahan (south of Padang). There also is a small island called Pulau Pasu in the Riau Islands, but that archipelago is located north of Sumatra, in the South China Sea. So, it is unclear where that specimen was collected exactly in Sumatra.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" author="O'Donoghue, CH" journalOrPublisher="Transactions of the Zoological Society of London" pageId="0" pageNumber="1" pagination="713 - 841" refId="B137" refString="O'Donoghue, CH, 1929. Report on the Opisthobranchiata. Zoological Results of the Cambridge Expedition to the Suez Canal. Transactions of the Zoological Society of London 22: 713 - 841, DOI: https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" title="Report on the Opisthobranchiata. Zoological Results of the Cambridge Expedition to the Suez Canal." url="https://doi.org/10.1111/j.1096-3642.1929.tb00212.x" volume="22" year="1929">
<normalizedToken originalValue="ODonoghue">O'Donoghue</normalizedToken>
(1929
</bibRefCitation>
: 833) reported one specimen (30/21 mm) of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
from Port Taufiq, Suez, Egypt. A radular formula (65
<normalizedToken originalValue="×">x</normalizedToken>
72-1-72) is not enough to identify a
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species, and he most likely examined
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
or
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Two names accepted as valid by
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a)
</bibRefCitation>
are regarded as synonyms of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
:
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 191) himself acknowledged that differences between
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
were weak. The traits that he mentioned (position of the pneumostome with respect to the anus, head longer than the foot) vary greatly due to preservation.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 197-198) did not compare
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Paraperonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraperonia fidjiensis" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">Paraperonia fidjiensis</emphasis>
</taxonomicName>
with
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
probably because he classified them in two distinct genera. However, there are no differences between the type material of
<taxonomicName lsidName="P. fidjiensis" pageId="0" pageNumber="1" rank="species" species="fidjiensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. fidjiensis</emphasis>
</taxonomicName>
and the type material of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 190) agreed with most authors that
<taxonomicName lsidName="P. mauritiana" pageId="0" pageNumber="1" rank="species" species="mauritiana">
<emphasis italics="true" pageId="0" pageNumber="1">P. mauritiana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. melanopneumon" pageId="0" pageNumber="1" rank="species" species="melanopneumon">
<emphasis italics="true" pageId="0" pageNumber="1">O. melanopneumon</emphasis>
</taxonomicName>
were synonyms of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. Like
<bibRefCitation author="Plate, L" journalOrPublisher="Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere" pageId="0" pageNumber="1" pagination="93 - 234" publicationUrl="https://biodiversitylibrary.org/page/11175213" refId="B146" refString="Plate, L, 1893. Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden. Zoologische Jahrbuecher, Anatomie und Ontogenie der Thiere 7: 93 - 234, https://biodiversitylibrary.org/page/11175213" title="Studien ueber opisthopneumone Lungenschnecken, II, Die Oncidiiden." url="https://biodiversitylibrary.org/page/11175213" volume="7" year="1893">Plate (1893)</bibRefCitation>
,
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(1934a
</bibRefCitation>
: 190) thought that
<taxonomicName lsidName="P. corpulenta" pageId="0" pageNumber="1" rank="species" species="corpulenta">
<emphasis italics="true" pageId="0" pageNumber="1">P. corpulenta</emphasis>
</taxonomicName>
was simply a potential synonym of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
but in fact it is a
<emphasis italics="true" pageId="0" pageNumber="1">nomen dubium</emphasis>
(see general discussion).
</paragraph>
<paragraph pageId="0" pageNumber="1">
All references cited by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
for
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
have been commented on above, but
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbés">Labbe's</normalizedToken>
(1934a)
</bibRefCitation>
proposed distribution ranges need additional clarification.
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbés">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 190-191) re-description of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
was based on one individual (100/70 mm) from Sumatra (MNHN-IM-2012-25150), one individual (90/70 mm) from the Seychelles (MNHN-IM-2012-25149), and ten individuals from the Red Sea (not found in the MNHN collections). At least one of those specimens belongs to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
because of the length of the spine of the accessory penial gland mentioned by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
as 6 to 7 mm. The specimens from Sumatra and the Seychelles were fully dissected by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
(the Sumatra individual is basically empty): the male parts are missing, and it is not possible to determine the type of intestinal loops. However, given their huge size, they most likely belong to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. The presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in the Red Sea is possible but, at this stage, questionable: the size mentioned by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
for those specimens (17/12 mm) strongly suggests that he did not examine
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
from the Red Sea. Those specimens from the Red Sea identified as
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
could not be located at the MNHN (there are no specimens collected by Clot-Bey in the collections, and there are too many jars of specimens collected by Jousseaume to determine which jar corresponds to the species description in
<normalizedToken originalValue="Labbés">Labbe's</normalizedToken>
monograph).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Labbe, A" journalOrPublisher="Annales de l'Institut Oceanographique" pageId="0" pageNumber="1" pagination="173 - 246" refId="B108" refString="Labbe, A, 1934a. Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues. Annales de l'Institut Oceanographique 14: 173 - 246" title="Les Silicodermes Labbe du Museum d'Histoire Naturelle de Paris. Premiere partie: classification, formes nouvelles ou peu connues." volume="14" year="1934 a">
<normalizedToken originalValue="Labbés">Labbe's</normalizedToken>
(1934a
</bibRefCitation>
: 191-192, figs 4-7) re-description of
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
was based on one individual from Djibouti (Obock), one individual (85/60 mm) from the Seychelles (MNHN-IM-2012-25148), two individuals from New Ireland, and one individual from Tonga which happens to be part of the type series by Quoy and Gaimard (MNHN-IM-2000-22937) even though
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
does not mention it. The specimen from the Seychelles was re-examined for the present study and, given its huge size, it is confirmed that it belongs to
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
: its intestinal loops are of type I, with a transitional loop at 3
<normalizedToken originalValue="oclock">o'clock</normalizedToken>
; the male parts are missing. There are two specimens (60/50 mm) from New Ireland at the MNHN which could potentially be the two specimens mentioned by
<normalizedToken originalValue="Labbé">Labbe</normalizedToken>
, but the collecting dates do not match. At any rate, it does not matter much since our fresh specimens demonstrate that
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is present in New Ireland (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). The specimen from Obock could not be traced back at the MNHN; there is a specimen (80/60 mm) which could possibly correspond to it but it is a problematic specimen as it could also be a type specimen for
<taxonomicName lsidName="P. gaimardi" pageId="0" pageNumber="1" rank="species" species="gaimardi">
<emphasis italics="true" pageId="0" pageNumber="1">P. gaimardi</emphasis>
</taxonomicName>
, and is now an empty notum (see below, remarks on the type material of
<taxonomicName lsidName="P. gaimardi" pageId="0" pageNumber="1" rank="species" species="gaimardi">
<emphasis italics="true" pageId="0" pageNumber="1">P. gaimardi</emphasis>
</taxonomicName>
in
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
). Thus, the presence of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in Djibouti is not accepted here and would need to be supported by positive evidence.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Risbec, J" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="387 - 417" refId="B152" refString="Risbec, J, 1935. Biologie et ponte de Mollusques Gasteropodes Neo-Caledoniens. Bulletin de la Societe Zoologique de France 60: 387 - 417" title="Biologie et ponte de Mollusques Gasteropodes Neo-Caledoniens." volume="60" year="1935">Risbec (1935</bibRefCitation>
: 415) illustrated the eggs of an onchidiid individual from New Caledonia which he called &quot;
<taxonomicName class="Gastropoda" family="Onchidiidae" genus="Oncidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncidium tonga" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonga">
<emphasis italics="true" pageId="0" pageNumber="1">Oncidium tonga</emphasis>
</taxonomicName>
Q et G,&quot; clearly a spelling mistake for
<taxonomicName authorityName="Quoy &amp; Gaimard" authorityYear="1832" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium tonganum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonganum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium tonganum</emphasis>
</taxonomicName>
Quoy &amp; Gaimard, 1832. It is not possible to know what species Risbec was referring to because there are three
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species in New Caledonia (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a064590" author="White, KM" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="241 - 253" refId="B195" refString="White, KM, 1951. On a collection of Molluscs, mainly Nudibranchs from the Red Sea. Proceedings of the Malacological Society of London 28 (6): 241 - 253, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a064590" title="On a collection of Molluscs, mainly Nudibranchs from the Red Sea." url="https://doi.org/10.1093/oxfordjournals.mollus.a064590" volume="28" year="1951">White (1951</bibRefCitation>
: 241) reported a single specimen (53/38 mm) of
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from the Persian Gulf. The radular formula (88
<normalizedToken originalValue="×">x</normalizedToken>
88-1-88) is not enough to identify a
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species.
<normalizedToken originalValue="Whites">White's</normalizedToken>
record referred either to
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #4) or
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
In Japan,
<bibRefCitation DOI="https://doi.org/10.2307/2520262" author="Baba, K" journalOrPublisher="Zoological Memoirs, University of Bombay" pageId="0" pageNumber="1" refId="B12" refString="Baba, K, 1958. Seashore life of Japan [in Japanese]. Hokuryukan, Tokyo, 148 pp. [5 pls] https://doi.org/10.2307/2520262" title="Seashore life of Japan [in Japanese]. Hokuryukan, Tokyo, 148 pp. [5 pls]" url="https://doi.org/10.2307/2520262" year="1958">Baba (1958</bibRefCitation>
: 144) indicated that some specimens of
<taxonomicName authorityName="Cuvier" authorityYear="1830" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium verruculatum" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium verruculatum</emphasis>
</taxonomicName>
from Tokara Islands, south of Kyushu (ca. 30°N) are very large (up to 120 mm long), suggesting that
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
is found there, which would be its most northern record.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Macnae, W" journalOrPublisher="Mitochondrial DNA" pageId="0" pageNumber="1" refId="B118" refString="Macnae, W, Kalk, M [Eds], 1958. A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." title="A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." year="1958">Macnae and Kalk (1958</bibRefCitation>
: 34, 44, 128) mentioned
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Inhaca Island, Mozambique (ca. 26°S). Given that no information is provided for species identification, this record is not taken for granted.
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
was likely confused with
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5), which our material indicates is present in Inhaca, or even
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, known from South Africa to western India (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). The fact that the slugs were found on sand (
<bibRefCitation author="Macnae, W" journalOrPublisher="Mitochondrial DNA" pageId="0" pageNumber="1" refId="B118" refString="Macnae, W, Kalk, M [Eds], 1958. A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." title="A Natural History of Inhaca Island, Mocambique. Wiwatersrand University Press, Johannesburg, vi + 163 pp." year="1958">Macnae and Kalk 1958</bibRefCitation>
: 128) could suggest that they saw
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #5).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2888" author="Solem, A" journalOrPublisher="Fieldiana Zoology" pageId="0" pageNumber="1" pagination="1 - 359" refId="B169" refString="Solem, A, 1959. Systematics and zoogeography of the land and fresh-water Mollusca of the New Hebrides. Fieldiana Zoology 4: 1 - 359, DOI: https://doi.org/10.5962/bhl.title.2888" title="Systematics and zoogeography of the land and fresh-water Mollusca of the New Hebrides." url="https://doi.org/10.5962/bhl.title.2888" volume="4" year="1959">Solem (1959</bibRefCitation>
: 39) did not report any new material or localities for
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
. The references that he mentioned (e.g.,
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.12.1919.888" author="Bretnall, W" journalOrPublisher="Records of the Australian Museum" pageId="0" pageNumber="1" pagination="303 - 328" refId="B26" refString="Bretnall, W, 1919. Onchidiidae from Australia and the South-Western Pacific Islands. Records of the Australian Museum 12: 303 - 328, DOI: https://doi.org/10.3853/j.0067-1975.12.1919.888" title="Onchidiidae from Australia and the South-Western Pacific Islands." url="https://doi.org/10.3853/j.0067-1975.12.1919.888" volume="12" year="1919">Bretnall 1919</bibRefCitation>
;
<bibRefCitation author="Hoffmann, H" journalOrPublisher="Zoologische Jahrbuecher" pageId="0" pageNumber="1" pagination="29 - 118" refId="B92" refString="Hoffmann, H, 1928. Zur Kenntnis der Oncidiiden. Zoologische Jahrbuecher 55: 29 - 118" title="Zur Kenntnis der Oncidiiden." volume="55" year="1928">Hoffmann 1928</bibRefCitation>
) are already commented on above. His proposed distribution (&quot;from the Red Sea and Mauritius to New Caledonia, Samoa, and the Marshall Islands&quot;) is not fully accurate because it is based on the assumption that people never made any mistakes when identifying
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
, which is unfortunately not true.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.2888" author="Solem, A" journalOrPublisher="Fieldiana Zoology" pageId="0" pageNumber="1" pagination="1 - 359" refId="B169" refString="Solem, A, 1959. Systematics and zoogeography of the land and fresh-water Mollusca of the New Hebrides. Fieldiana Zoology 4: 1 - 359, DOI: https://doi.org/10.5962/bhl.title.2888" title="Systematics and zoogeography of the land and fresh-water Mollusca of the New Hebrides." url="https://doi.org/10.5962/bhl.title.2888" volume="4" year="1959">Solem (1959</bibRefCitation>
: 38) mentioned what he thought were the three &quot;most obvious&quot; of the &quot;numerous differences&quot; between
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. verruculatum" pageId="0" pageNumber="1" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. verruculatum</emphasis>
</taxonomicName>
: distribution of branchial plumes (dorsal gills) on the notum, relative position of the pneumostome and the anus, and relative width of the hyponotum and pedal sole. But those features vary among individuals and should not be used for species identification.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Marcus, Er" journalOrPublisher="Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse" pageId="0" pageNumber="1" pagination="873 - 934" refId="B122" refString="Marcus, Er, Marcus, Ev, 1960. Opisthobranchia aus dem Roten Meer und von den Malediven. Akademie der Wissenschaften und der Literature, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse 12: 873 - 934" title="Opisthobranchia aus dem Roten Meer und von den Malediven." volume="12" year="1960">Marcus and Marcus (1960</bibRefCitation>
: 877) described
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
from the Maldives based on eight specimens. Given that they report a maximum animal length of 155 mm, a long (4.5 mm) spine of the accessory penial gland, as well as a retractor muscle inserting near the heart, there is little doubt that they did examine
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Later,
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus (1970</bibRefCitation>
: 213) added that they observed a retractor muscle inserting near the nerve ring in another of their specimens from the Maldives, which also is compatible with our present observations: a vestigial retractor muscle was even observed here in
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of traits that can help identify Peronia species, based exclusively on specimens examined for the present revision. Species are arranged in a phylogenetic order (Figs 2 - 4). Traits are described in detail in species descriptions." httpUri="http://table.plazi.org/id/8BDBED7FCCE9A8E1C67A716B088DDD7D" pageId="0" pageNumber="1" tableUuid="8BDBED7FCCE9A8E1C67A716B088DDD7D">4</tableCitation>
). Some of the material examined from historical museum collections for the present work also came from the Maldives (ANSP 304860).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Webb, M" journalOrPublisher="South African Journal of Science" pageId="0" pageNumber="1" pagination="107 - 118" refId="B193" refString="Webb, M, Moodley, LG, Thandar, AS, 1969. The copulatory mechanism in Onchidium peronii. South African Journal of Science 65: 107 - 118" title="The copulatory mechanism in Onchidium peronii." volume="65" year="1969">Webb et al. (1969</bibRefCitation>
: 107-112) described copulatory mechanisms in specimens they identified as
<taxonomicName lsidName="O. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">O. peronii</emphasis>
</taxonomicName>
. It is unclear from where those specimens were, possibly South Africa. At any rate, given that they illustrate a spine of the accessory penial gland which is only 1.4 mm long (
<bibRefCitation author="Webb, M" journalOrPublisher="South African Journal of Science" pageId="0" pageNumber="1" pagination="107 - 118" refId="B193" refString="Webb, M, Moodley, LG, Thandar, AS, 1969. The copulatory mechanism in Onchidium peronii. South African Journal of Science 65: 107 - 118" title="The copulatory mechanism in Onchidium peronii." volume="65" year="1969">Webb et al. 1969</bibRefCitation>
: 110, fig. 3), they did not examine individuals of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
It is not possible to determine whether
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus (1970</bibRefCitation>
: 213) examined an individual of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
from Madagascar because they do not provide the key features that characterize it. They could have seen a large individual of
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
instead.
<bibRefCitation DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065863" author="Britton, KM" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="179 - 191" refId="B27" refString="Britton, KM, 1984. The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera. Journal of Molluscan Studies 50: 179 - 191, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a065863" title="The Onchidiacea (Gastropoda, Pulmonata) of Hong Kong with a worldwide review of the genera." url="https://doi.org/10.1093/oxfordjournals.mollus.a065863" volume="50" year="1984">Britton (1984</bibRefCitation>
: 183) merely mentioned the fact that
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus (1970)</bibRefCitation>
accepted only two valid species names (
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
), which is not strictly exact because
<bibRefCitation author="Marcus, Ev" journalOrPublisher="Malacologia" pageId="0" pageNumber="1" pagination="181 - 223" publicationUrl="https://www.biodiversitylibrary.org/page/13147904" refId="B123" refString="Marcus, Ev, Marcus, Er, 1970. Some gastropods from Madagascar and west Mexico. Malacologia 10: 181 - 223, https://www.biodiversitylibrary.org/page/13147904" title="Some gastropods from Madagascar and west Mexico." url="https://www.biodiversitylibrary.org/page/13147904" volume="10" year="1970">Marcus and Marcus (1970)</bibRefCitation>
did not address the nomenclatural status of
<taxonomicName lsidName="P. tongana" pageId="0" pageNumber="1" rank="species" species="tongana">
<emphasis italics="true" pageId="0" pageNumber="1">P. tongana</emphasis>
</taxonomicName>
and did say that
<taxonomicName lsidName="P. branchifera" pageId="0" pageNumber="1" rank="species" species="branchifera">
<emphasis italics="true" pageId="0" pageNumber="1">P. branchifera</emphasis>
</taxonomicName>
was close to
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
but not that it was its synonym.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Patil, P" journalOrPublisher="International Journal of Zoology and Research" pageId="0" pageNumber="1" pagination="23 - 30" publicationUrl="http://www.tjprc.org/view_paper.php?id=2556" refId="B143" refString="Patil, P, Kulkarni, BG, 2013. The Onchidium (Gastropoda: Pulmonata: Onchidiidae: Genus: Onchidium) of the Uran, West Coast of India. International Journal of Zoology and Research 3 (4): 23 - 30, http://www.tjprc.org/view_paper.php?id=2556" title="The Onchidium (Gastropoda: Pulmonata: Onchidiidae: Genus: Onchidium) of the Uran, West Coast of India." url="http://www.tjprc.org/view_paper.php?id=2556" volume="3" year="2013">Patil and Kulkarni (2013)</bibRefCitation>
reported
<taxonomicName authorityName="Cuvier" authorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Onchidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onchidium peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Onchidium peronii</emphasis>
</taxonomicName>
from Uran City, near Mumbai, India, but it is impossible to determine what species they saw (most likely it was
<taxonomicName lsidName="P. madgascariensis" pageId="0" pageNumber="1" rank="species" species="madgascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madgascariensis</emphasis>
</taxonomicName>
or the unit #4 of
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
, or both).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Many chemical studies have mentioned
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in the past few decades. However, the name
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
was used arbitrarily. The individuals used for the extraction of natural products may not have been properly identified.
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)98720-3" author="Biskupiak, JE" journalOrPublisher="Tetrahedon Letters" pageId="0" pageNumber="1" pagination="4307 - 4310" refId="B19" refString="Biskupiak, JE, Ireland, CM, 1985. Cytotoxic metabolites from the mollusc Peronia peronii. Tetrahedon Letters 26: 4307 - 4310, DOI: https://doi.org/10.1016/S0040-4039(00)98720-3" title="Cytotoxic metabolites from the mollusc Peronia peronii." url="https://doi.org/10.1016/S0040-4039(00)98720-3" volume="26" year="1985">Biskupiak and Ireland (1985)</bibRefCitation>
extracted peroniatriols from specimens identified as
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
from Guam.
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
is undeniably present in Guam. However, it is possible that
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #1) could be present there as well.
<bibRefCitation author="Pietra, F" journalOrPublisher="International Journal of Zoology and Research" pageId="0" pageNumber="1" refId="B144" refString="Pietra, F, 1990. A Secret World. Natural Products of Marine Life. Birkhaeuser Verlag, Basel, Boston, Berlin, viii + 280 pp." title="A Secret World. Natural Products of Marine Life. Birkhaeuser Verlag, Basel, Boston, Berlin, viii + 280 pp." year="1990">Pietra (1990</bibRefCitation>
: 145) mentioned peroniatriols in
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
from Micronesia where more than one species may be found.
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)73859-7" author="Arimoto, H" journalOrPublisher="Tetrahedron Letters" pageId="0" pageNumber="1" pagination="5781 - 5784" refId="B8" refString="Arimoto, H, Cheng, JF, Nishiyama, S, Yamamura, S, 1993. Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II. Tetrahedron Letters 34 (36): 5781 - 5784, DOI: https://doi.org/10.1016/S0040-4039(00)73859-7" title="Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II." url="https://doi.org/10.1016/S0040-4039(00)73859-7" volume="34" year="1993">Arimoto et al. (1993)</bibRefCitation>
did not indicate where specimens of
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. verruculatum" pageId="0" pageNumber="1" rank="species" species="verruculatum">
<emphasis italics="true" pageId="0" pageNumber="1">O. verruculatum</emphasis>
</taxonomicName>
were collected. In Japan, where the individuals used by
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)73859-7" author="Arimoto, H" journalOrPublisher="Tetrahedron Letters" pageId="0" pageNumber="1" pagination="5781 - 5784" refId="B8" refString="Arimoto, H, Cheng, JF, Nishiyama, S, Yamamura, S, 1993. Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II. Tetrahedron Letters 34 (36): 5781 - 5784, DOI: https://doi.org/10.1016/S0040-4039(00)73859-7" title="Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II." url="https://doi.org/10.1016/S0040-4039(00)73859-7" volume="34" year="1993">Arimoto et al. (1993)</bibRefCitation>
possibly came from, there are four
<taxonomicName authorityName="Fleming" authorityYear="1822" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia</emphasis>
</taxonomicName>
species which are all cryptic externally.
<bibRefCitation DOI="https://doi.org/10.1016/S1460-1567(02)80014-9" author="Pietra, F" journalOrPublisher="Pergamon Elsevier Science, Amsterdam" pageId="0" pageNumber="1" refId="B145" refString="Pietra, F, 2002. Biodiversity and Natural Product Diversity. Pergamon Elsevier Science, Amsterdam, DOI: https://doi.org/10.1016/S1460-1567(02)80014-9" title="Biodiversity and Natural Product Diversity." url="https://doi.org/10.1016/S1460-1567(02)80014-9" year="2002">Pietra (2002</bibRefCitation>
: 290) briefly cited peroniatriols in
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
based on the work by
<bibRefCitation DOI="https://doi.org/10.1016/S0040-4039(00)73859-7" author="Arimoto, H" journalOrPublisher="Tetrahedron Letters" pageId="0" pageNumber="1" pagination="5781 - 5784" refId="B8" refString="Arimoto, H, Cheng, JF, Nishiyama, S, Yamamura, S, 1993. Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II. Tetrahedron Letters 34 (36): 5781 - 5784, DOI: https://doi.org/10.1016/S0040-4039(00)73859-7" title="Synthetic studies on fully substituted γ-pyrone-containing natural products: The absolute configurations of Ilikonapyrone and peroniatriols I and II." url="https://doi.org/10.1016/S0040-4039(00)73859-7" volume="34" year="1993">Arimoto et al. (1993)</bibRefCitation>
. Finally, the antibacterial peptide extracted from individuals of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
from the Persian Gulf (
<bibRefCitation DOI="https://doi.org/10.13005/bbra/1870" author="Bitaab, MA" journalOrPublisher="Biosciences Biotechnology Research Asia" pageId="0" pageNumber="1" pagination="2023 - 2035" refId="B20" refString="Bitaab, MA, Ranaei Siadat, SO, Pazooki, J, Sefidbakht, Y, 2015. Antibacterial and molecular dynamics study of the Dolabellanin B2 isolated from sea slug, Peronia peronii. Biosciences Biotechnology Research Asia 12 (3): 2023 - 2035, DOI: https://doi.org/10.13005/bbra/1870" title="Antibacterial and molecular dynamics study of the Dolabellanin B 2 isolated from sea slug, Peronia peronii." url="https://doi.org/10.13005/bbra/1870" volume="12" year="2015">Bitaab et al. 2015</bibRefCitation>
) was most likely extracted from individuals of either
<taxonomicName lsidName="P. verruculata" pageId="0" pageNumber="1" rank="species" species="verruculata">
<emphasis italics="true" pageId="0" pageNumber="1">P. verruculata</emphasis>
</taxonomicName>
(unit #4), or
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, or both (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F8" captionText="Figure 6. Geographical distribution of the Peronia species A distribution of all Peronia species except for P. peronii B distribution of P. peronii. The colors are the same as those used in Figs 2 - 5. Colored areas correspond to hypothetical geographical ranges based on confirmed records only. Distinct colors are used for each unit of P. verruculata. The distribution of P. verruculata in the Indo-West Pacific is actually continuous. However, because it is unclear which units are present in regions from where we have no fresh material of P. verruculata (red areas), no unit of P. verruculata is shown there. For P. peronii, black dots correspond to material identified based on anatomical characters and blue dots correspond to material with DNA sequences. Details on species distribution can be found in each species description." figureDoi="10.3897/zookeys.972.52853.figure6" httpUri="https://binary.pensoft.net/fig/458306" pageId="0" pageNumber="1">6</figureCitation>
). The same general remark applies to ecological studies:
<bibRefCitation DOI="https://doi.org/10.1201/EBK1439821169-2" author="Morrisey, D" editor="Gibson, RN" journalOrPublisher="(Vol. 48). CRC Press, Boca Raton" pageId="0" pageNumber="1" pagination="43 - 160" refId="B132" refString="Morrisey, D, Swales, A, Ditttmann, S, Morrison, MA, Lovelock, CE, Beard, CM, 2010. The ecology and management of temperate mangroves. In: Gibson, RN, Atkinson, RJA, Gordon, JDM, Eds., Oceanography and Marine Biology: an Annual Review. (Vol. 48). CRC Press, Boca Raton: 43 - 160, DOI: https://doi.org/10.1201/EBK1439821169-2" title="The ecology and management of temperate mangroves." url="https://doi.org/10.1201/EBK1439821169-2" volumeTitle="Oceanography and Marine Biology: an Annual Review." year="2010">Morrisey et al. (2010</bibRefCitation>
: 72) listed (with no justification for species identification) the presence of
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
in mangroves of the estuary of the Mtata River (31°57'S), South Africa; most likely,
<bibRefCitation DOI="https://doi.org/10.1201/EBK1439821169-2" author="Morrisey, D" editor="Gibson, RN" journalOrPublisher="(Vol. 48). CRC Press, Boca Raton" pageId="0" pageNumber="1" pagination="43 - 160" refId="B132" refString="Morrisey, D, Swales, A, Ditttmann, S, Morrison, MA, Lovelock, CE, Beard, CM, 2010. The ecology and management of temperate mangroves. In: Gibson, RN, Atkinson, RJA, Gordon, JDM, Eds., Oceanography and Marine Biology: an Annual Review. (Vol. 48). CRC Press, Boca Raton: 43 - 160, DOI: https://doi.org/10.1201/EBK1439821169-2" title="The ecology and management of temperate mangroves." url="https://doi.org/10.1201/EBK1439821169-2" volumeTitle="Oceanography and Marine Biology: an Annual Review." year="2010">Morrisey et al. (2010</bibRefCitation>
: 72) encountered
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
instead.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Finally, a few last words on
<taxonomicName lsidName="P. peronii" pageId="0" pageNumber="1" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">P. peronii</emphasis>
</taxonomicName>
in phylogenetic studies.
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. (2011</bibRefCitation>
: 428) and
<bibRefCitation DOI="https://doi.org/10.1186/1471-2148-11-295" author="White, TR" journalOrPublisher="BMC Evolutionary Biology" pageId="0" pageNumber="1" pagination="1 - 295" refId="B196" refString="White, TR, Conrad, M, Tseng, R, Golding, R, Gomes, S, de Frias Martins, AM, Dayrat, B, 2011. Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications. BMC Evolutionary Biology 11: 1 - 295, DOI: https://doi.org/10.1186/1471-2148-11-295" title="Ten new complete mitochondrial genomes of pulmonate gastropods (Mollusca): Systematic and macro-evolutionary implications." url="https://doi.org/10.1186/1471-2148-11-295" volume="11" year="2011">White et al. (2011</bibRefCitation>
: 4) identified a specimen from Guam (CASIZ 180486) as
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia peronii</emphasis>
</taxonomicName>
, which is specimen [443] in the present study (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
). The specimen tentatively identified as
<taxonomicName baseAuthorityName="Cuvier" baseAuthorityYear="1804" class="Gastropoda" family="Onchidiidae" genus="Peronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Peronia peronii" order="Systellommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peronii">
<emphasis italics="true" pageId="0" pageNumber="1">Peronia cf. peronii</emphasis>
</taxonomicName>
from Mozambique (NHMUK 20060414) by
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2011.02.014" author="Dayrat, B" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="1" pagination="425 - 437" refId="B50" refString="Dayrat, B, Conrad, M, Balayan, S, White, TR, Albrecht, C, Golding, R, Gomes, S, Harasewych, MG, Frias Martins, AM de, 2011. Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling. Molecular Phylogenetics and Evolution 59: 425 - 437, DOI: https://doi.org/10.1016/j.ympev.2011.02.014" title="Phylogenetic relationships and evolution of pulmonate gastropods (Mollusca): new insights from increased taxon sampling." url="https://doi.org/10.1016/j.ympev.2011.02.014" volume="59" year="2011">Dayrat et al. (2011</bibRefCitation>
: 428) belongs to
<taxonomicName lsidName="P. madagascariensis" pageId="0" pageNumber="1" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">P. madagascariensis</emphasis>
</taxonomicName>
, which is specimen [735] in the present study (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phylogenetic relationships between Peronia species based on concatenated mitochondrial COI and 16 S DNA sequences. Numbers by the branches are the bootstrap values (maximum likelihood analysis) and the posterior probabilities (Bayesian analysis). Only the values&gt; 50 % (ML) and&gt; 0.9 (Bayesian) are indicated. Numbers for each individual correspond to unique identifiers for DNA extraction. Information on specimens can be found in the lists of material examined and in Table 2. The color used for each species or mitochondrial unit is the same as the color used in Figs 3 - 6." figureDoi="10.3897/zookeys.972.52853.figure2" httpUri="https://binary.pensoft.net/fig/458300" pageId="0" pageNumber="1">2</figureCitation>
). The DNA sequences of the specimen from Guam were used again in several studies (e.g.,
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0067299" author="Gaitan-Espitia, JD" journalOrPublisher="Annual Reviews of Ecology, Evolution and Systematics" pageId="0" pageNumber="1" refId="B76" refString="Gaitan-Espitia, JD, Nespolo, RF, Opazo, JC, 2013. The Complete Mitochondrial Genome of the Land Snail Cornuaspersum (Helicidae: Mollusca): Intra-Specific Divergence of Protein-Coding Genes and Phylogenetic Considerations within Euthyneura. PLoS ONE 8: e67299. https://doi.org/10.1371/journal.pone.0067299" title="The Complete Mitochondrial Genome of the Land Snail Cornuaspersum (Helicidae: Mollusca): Intra-Specific Divergence of Protein-Coding Genes and Phylogenetic Considerations within Euthyneura. PLoS ONE 8: e 67299." url="https://doi.org/10.1371/journal.pone.0067299" year="2013">
<normalizedToken originalValue="Gaitán-Espitia">Gaitan-Espitia</normalizedToken>
et al. 2013
</bibRefCitation>
;
<bibRefCitation author="Harasewych, MG" journalOrPublisher="The Nautilus" pageId="0" pageNumber="1" pagination="156 - 162" publicationUrl="https://www.biodiversitylibrary.org/page/59108238" refId="B89" refString="Harasewych, MG, Windsor, AM, Lopez-Vera, E, Thompson, FG, 2015. On the phylogenetic relationships of the genus Mexistrophia and of the family Cerionidae (Gastropoda: Eupulmonata). The Nautilus 129: 156 - 162, https://www.biodiversitylibrary.org/page/59108238" title="On the phylogenetic relationships of the genus Mexistrophia and of the family Cerionidae (Gastropoda: Eupulmonata)." url="https://www.biodiversitylibrary.org/page/59108238" volume="129" year="2015">Harasewych et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>