396 lines
30 KiB
XML
396 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="772940C18E935467A73420AD0C3F704E" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria mcelroyorum Means & Hennen & Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="181004738" LSID="urn:lsid:zoobank.org:act:37EFC360-6A08-41DD-848F-2C5C97C60E70" httpUri="http://treatment.plazi.org/id/772940C18E935467A73420AD0C3F704E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="8RVT2" LSID="http://zoobank.org/37EFC360-6A08-41DD-848F-2C5C97C60E70" authority="Means & Hennen & Marek, 2021" authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria mcelroyorum" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mcelroyorum" status="sp. nov.">Nannaria mcelroyorum</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Nannaria mcelroyorum sp. nov. holotype ♂ (VTEC, MPE 03113) left gonopod A anterior view; red triangle indicates medial flange B medial view C posterior view; red arrow indicates prefemoral spine secondary hump. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure26" httpUri="https://binary.pensoft.net/fig/531129" pageId="0" pageNumber="1">Figs 26</figureCitation>
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Nannaria mcelroyorum sp. nov. non-type ♂ (VTEC, MPE 03125) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure27" httpUri="https://binary.pensoft.net/fig/531130" pageId="0" pageNumber="1">, 27 Vernacular name: "The McElroy Twisted-Claw Millipede"</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355337" accessionNumber="MPE03113" collectingDate="2017-09-23" collectorName="J. Means, D. Hennen" country="United States" county="Wayne County" elevation="194" latitude="38.3047" location="Beech Fork State Park" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" specimenCount="♂" stateProvince="West Virginia" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount>♂</specimenCount>
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;
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<collectingCounty>Wayne County</collectingCounty>
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,
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<collectingMunicipality>Barboursville</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:910F8FD5B2274483FD680874734C2A3B" country="United States" county="Wayne County" latitude="38.3047" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" name="Beech Fork State Park" stateProvince="West Virginia">Beech Fork State Park</location>
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,
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<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:987EA256E930AF559606EBB3EC22A324" country="United States" county="Wayne County" latitude="38.3047" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" name="Lost Trail" stateProvince="West Virginia">Lost Trail</location>
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near
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<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:16E6E292F1F468DE0CC6E7651B971314" country="United States" county="Wayne County" latitude="38.3047" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" name="Moxley Branch Campground" stateProvince="West Virginia">Moxley Branch Campground</location>
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;
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<geoCoordinate degrees="38.3047" direction="north" orientation="latitude" precision="5" value="38.3047">38.3047°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.3512" direction="west" orientation="longitude" precision="5" value="-82.3512">82.3512°W</geoCoordinate>
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; elev.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="1.94" unit="m" value="194.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="1.94" unit="m" value="194.0">194 m</elevation>
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</quantity>
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;
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<collectingDate value="2017-09-23">23 Sep. 2017</collectingDate>
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; hand collected;
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<collectorName>J. Means</collectorName>
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and
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<collectorName>D. Hennen</collectorName>
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leg.; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03113">MPE03113</accessionNumber>
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.
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355554" accessionNumber="MPE03116" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Wayne County" elevation="194" latitude="38.3047" location="West Virginia" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" specimenCount="3" specimenCount-male="3" stateProvince="West Virginia" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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</emphasis>
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">3 ♂♂</specimenCount>
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; same collection data as holotype; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03116">MPE03116</accessionNumber>
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</materialsCitation>
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, 3118, 3120 •
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<materialsCitation ID-GBIF-Occurrence="3071355421" accessionNumber="MPE03135" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Wayne County" elevation="194" latitude="38.3047" location="Beech Fork State Park" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" specimenCount="4" specimenCount-male="4" stateProvince="West Virginia" typeStatus="paratype">
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<specimenCount type="male">4 ♂♂</specimenCount>
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; same collection data as holotype; VMNH
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03135">MPE03135</accessionNumber>
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</materialsCitation>
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, 3151, 3154, 3156 •
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<materialsCitation ID-GBIF-Occurrence="3071355317" accessionNumber="MPE03117" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Wayne County" elevation="194" latitude="38.3047" location="Beech Fork State Park" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" specimenCount="3" specimenCount-female="3" stateProvince="West Virginia" typeStatus="paratype">
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<specimenCount type="female">3 ♀♀</specimenCount>
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; same collection data as holotype; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03117">MPE03117</accessionNumber>
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</materialsCitation>
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, 3119, 3121 •
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<materialsCitation ID-GBIF-Occurrence="3071355647" accessionNumber="MPE03122" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Wayne County" elevation="194" latitude="38.3047" location="Beech Fork State Park" longLatPrecision="6" longitude="-82.3512" municipality="Barboursville" specimenCount="4" specimenCount-female="4" stateProvince="West Virginia" typeStatus="paratype">
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<specimenCount type="female">4 ♀♀</specimenCount>
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; same collection data as holotype; VMNH
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03122">MPE03122</accessionNumber>
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</materialsCitation>
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, 3136, 3137, 3155.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355490" accessionNumber="MPE03125" collectingDate="2017-09-24" collectorName="J. Means, D. Hennen" country="United States" county="Boyd County" elevation="219" latitude="38.3475" location="Bramble Drive" longLatPrecision="6" longitude="-82.6866" municipality="Fannin Park" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky">
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">1 ♂</specimenCount>
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;
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<collectingCounty>Boyd County</collectingCounty>
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,
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<collectingMunicipality>Fannin Park</collectingMunicipality>
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, off
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<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:2A53483974BBFE98C3D5CB81C7787309" country="United States" county="Boyd County" latitude="38.3475" longLatPrecision="6" longitude="-82.6866" municipality="Fannin Park" name="Bramble Drive" stateProvince="Kentucky">Bramble Drive</location>
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, hillside across road from park;
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<geoCoordinate degrees="38.3475" direction="north" orientation="latitude" precision="5" value="38.3475">38.3475°N</geoCoordinate>
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, -
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<geoCoordinate degrees="82.6866" direction="west" orientation="longitude" precision="5" value="-82.6866">82.6866°W</geoCoordinate>
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; elev.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="2.19" unit="m" value="219.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="2.19" unit="m" value="219.0">219 m</elevation>
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</quantity>
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;
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<collectingDate value="2017-09-24">24 Sep. 2017</collectingDate>
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;
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<collectorName>J. Means</collectorName>
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,
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<collectorName>D. Hennen</collectorName>
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leg.; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03125">MPE03125</accessionNumber>
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</materialsCitation>
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; SCAU -
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<materialsCitation ID-GBIF-Occurrence="3071355472" accessionNumber="MPE03262" collectingDate="2017-09-23" collectorName="J. Means, D. Hennen" country="United States" county="Boone County" elevation="223" latitude="38.1802" location="Big Pinnacle Road" longLatPrecision="6" longitude="-81.8384" municipality="Julian" specimenCount="3" specimenCount-male="3" stateProvince="West Virginia">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">3 ♂♂</specimenCount>
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;
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<collectingCounty>Boone County</collectingCounty>
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,
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<collectingMunicipality>Julian</collectingMunicipality>
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, along
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<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:EDB7492E2E9D2F5EDDD678859BD0C8F1" country="United States" county="Boone County" latitude="38.1802" longLatPrecision="6" longitude="-81.8384" municipality="Julian" name="Big Pinnacle Road" stateProvince="West Virginia">Big Pinnacle Road</location>
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||
;
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<geoCoordinate degrees="38.1802" direction="north" orientation="latitude" precision="5" value="38.1802">38.1802°N</geoCoordinate>
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, -
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<geoCoordinate degrees="81.8384" direction="west" orientation="longitude" precision="5" value="-81.8384">81.8384°W</geoCoordinate>
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; elev.
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<quantity metricMagnitude="2" metricUnit="m" metricValue="2.23" unit="m" value="223.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="2.23" unit="m" value="223.0">223 m</elevation>
|
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</quantity>
|
||
;
|
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<collectingDate value="2017-09-23">23 Sep. 2017</collectingDate>
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; hand collected;
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||
<collectorName>J. Means</collectorName>
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||
,
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<collectorName>D. Hennen</collectorName>
|
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leg.; VTEC
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03262">MPE03262</accessionNumber>
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||
</materialsCitation>
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||
, 63, 3289 •
|
||
<materialsCitation ID-GBIF-Occurrence="3071355587" accessionNumber="MPE03290" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Boone County" elevation="223" latitude="38.1802" location="Big Pinnacle Road" longLatPrecision="6" longitude="-81.8384" municipality="Julian" specimenCount="2" specimenCount-female="2" stateProvince="West Virginia">
|
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<specimenCount type="female">2 ♀♀</specimenCount>
|
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; same collection data as preceding; VTEC
|
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE03290">MPE03290</accessionNumber>
|
||
</materialsCitation>
|
||
, 91 •
|
||
<materialsCitation ID-GBIF-Occurrence="3071355601" accessionNumber="MPE02240" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Boone County" elevation="223" latitude="38.1802" location="Big Pinnacle Road" longLatPrecision="6" longitude="-81.8384" municipality="Julian" specimenCount="7" specimenCount-male="7" stateProvince="West Virginia">
|
||
<specimenCount type="male">7 ♂♂</specimenCount>
|
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; same collection data as preceding; VTEC
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02240">MPE02240</accessionNumber>
|
||
</materialsCitation>
|
||
,
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||
<materialsCitation ID-GBIF-Occurrence="3071355603" accessionNumber="MPE02240" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Boone County" elevation="223" latitude="38.1802" location="Big Pinnacle Road" longLatPrecision="6" longitude="-81.8384" municipality="Julian" specimenCount="1" stateProvince="West Virginia">2257, 2258, 2260-62, 3727 • same collection data as preceding</materialsCitation>
|
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;
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<materialsCitation ID-GBIF-Occurrence="3071355357" accessionNumber="MPE02248" collectingDate="2016-11-18" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means & D. Hennen" country="United States" county="Boone County" elevation="223" latitude="38.1802" location="Big Pinnacle Road" longLatPrecision="6" longitude="-81.8384" municipality="Julian" specimenCount="5" specimenCount-female="5" stateProvince="West Virginia">
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<specimenCount type="female">5 ♀♀</specimenCount>
|
||
; same collection data as preceding;
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||
<collectingDate value="2016-11-18">18 Nov. 2016</collectingDate>
|
||
; VTEC
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02248">MPE02248</accessionNumber>
|
||
</materialsCitation>
|
||
-
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<materialsCitation ID-GBIF-Occurrence="3071355743" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Suppl." country="United States" location="For" specimenCount="1" stateProvince="West Virginia">
|
||
50, 3728, 3729.
|
||
<location LSID="urn:lsid:plazi:treatment:772940C18E935467A73420AD0C3F704E:73B36A83F9F8D0F43B1B3080EA07A7BB" country="United States" name="For" stateProvince="West Virginia">For</location>
|
||
detailed collection data see
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||
<collectorName>Suppl.</collectorName>
|
||
material 7
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||
</materialsCitation>
|
||
.
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||
</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Adult males of
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria mcelroyorum" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mcelroyorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria mcelroyorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are distinct from other
|
||
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
|
||
</taxonomicName>
|
||
and the nearby
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
, based on the following combination of characters:
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
|
||
.
|
||
</emphasis>
|
||
Gonopodal acropodite gently curving medially throughout, not straight as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
. Acropodite gradually tapering towards tip, with small lobed medial flange (Fig.
|
||
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Nannaria mcelroyorum sp. nov. holotype ♂ (VTEC, MPE 03113) left gonopod A anterior view; red triangle indicates medial flange B medial view C posterior view; red arrow indicates prefemoral spine secondary hump. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure26" httpUri="https://binary.pensoft.net/fig/531129" pageId="0" pageNumber="1">26A</figureCitation>
|
||
, red triangle), lacking swollen medial area as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Tip directed medially, rounded and simple, not with folds, grooves, flanges as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
. Distal zone greatly reduced, not enlarged and curving caudolaterally as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
. Prefemur with thin, sinuous, acicular prefemoral process, not curved, saber-like as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
. Prefemoral spine small, medially directed, with secondary hump proximal to acropodite base (Fig.
|
||
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Nannaria mcelroyorum sp. nov. holotype ♂ (VTEC, MPE 03113) left gonopod A anterior view; red triangle indicates medial flange B medial view C posterior view; red arrow indicates prefemoral spine secondary hump. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure26" httpUri="https://binary.pensoft.net/fig/531129" pageId="0" pageNumber="1">26C</figureCitation>
|
||
, red arrow), not enlarged, cephalically directed as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., or lacking as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
. Telopodite basal zone height ca. 1/2 length of acropodite, not> 1/2 length of acropodite as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. serpens" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="serpens">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. serpens</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. or ca. 1/5 length as in
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. shenandoa" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="shenandoa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. shenandoa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
|
||
.
|
||
</emphasis>
|
||
Tergites with light orange paranotal spots and faint orange stripes (Fig.
|
||
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Nannaria mcelroyorum sp. nov. non-type ♂ (VTEC, MPE 03125) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure27" httpUri="https://binary.pensoft.net/fig/531130" pageId="0" pageNumber="1">27</figureCitation>
|
||
). Light brown background. Dorsum of collum smooth with orange margin.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1030.62544.figure26" httpUri="https://binary.pensoft.net/fig/531129" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria mcelroyorum" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mcelroyorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria mcelroyorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype ♂ (VTEC, MPE03113) left gonopod
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
anterior view; red triangle indicates medial flange
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
medial view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
posterior view; red arrow indicates prefemoral spine secondary hump. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1030.62544.figure27" httpUri="https://binary.pensoft.net/fig/531130" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
|
||
<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria mcelroyorum" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mcelroyorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria mcelroyorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. non-type ♂ (VTEC, MPE03125) coloration. Scale bar: 4.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">♂ paratype (VTEC, MPE03116; holotype too damaged for measurement): BL = 33.9, CW = 4.7, IW = 2.3, ISW = 0.9, B11W = 5.4, B11H = 3.6; ♀ paratype (VTEC, MPE03117): BL = 33.2, CW = 4.4, IW = 2.5, ISW = 1.3, B11W = 5.9, B11H = 4.3.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="variation">
|
||
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Individuals of
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. mcelroyorum" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="mcelroyorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. mcelroyorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from the Boone County, West Virginia site have prefemoral processes ca. 2/3 the length of those of the holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Known from a linear area in western West Virginia and Eastern Kentucky (Kentucky: Boyd County; West Virginia: Boone and Wayne counties, Suppl. material 7; Fig.
|
||
<figureCitation captionStart="Figure 127" captionStartId="F127" captionText="Figure 127. Distribution map of Nannaria minor species group collection localities in the northern Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure127" httpUri="https://binary.pensoft.net/fig/531230" pageId="0" pageNumber="1">127</figureCitation>
|
||
). Distribution area: 105 km2; status: MRE.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Individuals of
|
||
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. mcelroyorum" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="mcelroyorum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N. mcelroyorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. were collected from mesic deciduous forests dominated by oak, maple, laurel, and some pine, often under 1-2 cm soil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This species is named for Clint, Justin, Travis, and Griffin McElroy from Huntington, West Virginia, who provided endless hours of emotional support to JCM and DAH during field collections through their podcasts, 'The Adventure
|
||
<normalizedToken originalValue="Zone’">Zone'</normalizedToken>
|
||
and 'My Brother, My Brother, and
|
||
<normalizedToken originalValue="Me’">Me'</normalizedToken>
|
||
. The specific name is a plural genitive derived as a patronym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type locality">
|
||
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
United States, West Virginia, Wayne County, Barboursville, Beech Fork State Park, Lost Trail near Moxley Branch Campground,
|
||
<geoCoordinate degrees="38.3047" direction="north" orientation="latitude" precision="5" value="38.3047">38.3047°N</geoCoordinate>
|
||
, -
|
||
<geoCoordinate degrees="82.3512" direction="west" orientation="longitude" precision="5" value="-82.3512">82.3512°W</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |