235 lines
21 KiB
XML
235 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.701.13261" ID-GBIF-Dataset="a9b15909-ea61-40d7-ba04-325b8af91669" ID-PMC="PMC5674090" ID-Pensoft-Pub="1313-2970-701-1" ID-PubMed="29118596" ID-ZBK="23BDD7CE1C7E4D2092A8ED47267579FD" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-701-1" ModsDocOrigin="ZooKeys 701" ModsDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" checkinTime="1506095869158" checkinUser="pensoft" docAuthor="Fisher, J. Ray, Fisher, Danielle M., Skvarla, Michael J., Nelson, Whitney A. & Dowling, Ashley P. G." docDate="2017" docId="FDF673E9842E0784C17FA81DBE4FA3BF" docLanguage="en" docName="ZooKeys 701: 1-496" docOrigin="ZooKeys 701" docSource="http://dx.doi.org/10.3897/zookeys.701.13261" docTitle="Torrenticola olliei Fisher & Dowling, sp. n." docType="treatment" docUuid="EF825E5C-935A-4BDD-AEDA-A599A4839D7A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="3D440132FFABFFFC1505FFA0FFDDEA09" masterDocTitle="Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species" masterLastPageNumber="496" masterPageNumber="1" pageNumber="1" updateTime="1668164828938" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of torrent mites (Parasitengona, Torrenticolidae, Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, J. Ray</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Danielle M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skvarla, Michael J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nelson, Whitney A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dowling, Ashley P. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>701</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>496</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.701.13261</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.701.13261</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-701-1</mods:identifier>
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<mods:identifier type="ZBK">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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<mods:identifier type="ZooBank">23BDD7CE1C7E4D2092A8ED47267579FD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="134839613" LSID="urn:lsid:zoobank.org:act:EF825E5C-935A-4BDD-AEDA-A599A4839D7A" httpUri="http://treatment.plazi.org/id/FDF673E9842E0784C17FA81DBE4FA3BF" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/EF825E5C-935A-4BDD-AEDA-A599A4839D7A" authority="Fisher & Dowling" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei Fisher & Dowling</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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HOLOTYPE (♀): from USA, Oregon, Douglas County, Umpqua National Forest, Umpqua River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.29111">43°17'28"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.62">122°37'12"W</geoCoordinate>
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), 12 Aug 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0812-006.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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PARATYPES (9 ♀; 12 ♂): British Columbia, Canada: 1 ♀ and 1 ♂ from Vancouver Island, Malahat, Goldstream Provincial Park, Goldstream River, 26 Jun 1979, by IM Smith, IMS790028A
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<normalizedToken originalValue="•">*</normalizedToken>
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California, USA: 1 ♀ and 1 ♂ from Humboldt County, Honeydew, Mattole River, beside road to Bull Creek on east side of bridge, 8 Aug 1987, by IM Smith, IMS870135A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Trinity County, Trinity River, beside Route 299, 8.7 kilometers northwest of Del Loma, 9 Aug 1987, by IM Smith, IMS870137A
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Trinity County, Weaver Creek, beside Route 299, 4.3 kilometers north of Route 3 West, 9 Aug 1987, by IM Smith, IMS870138A & IMS870138B
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<normalizedToken originalValue="•">*</normalizedToken>
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Oregon, USA: 1 ♂ from Coos County, Siskiyou National Forest, Road 33 between Powers & Agness, Daphne Grove campground, Coquille River, 2 Jul 1983, by IM Smith, IMS830016
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<normalizedToken originalValue="•">*</normalizedToken>
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3 ♀ and 6 ♂ from Curry County, Port Orford, Butler Bar campground, Elk River, 25-26 Jun 1976, by IM Smith, IMS760163
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ from Curry County, Port Orford, small spring run beside road from Humbug Mountain State Park to McGribble campground, 25 Jun 1976, by IM Smith, IMS760161
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♂ (ALLOTYPE) from Douglas County, Umpqua National Forest, Umpqua River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.29111">43°17'28"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.62">122°37'12"W</geoCoordinate>
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), 12 Aug 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0812-006
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<normalizedToken originalValue="•">*</normalizedToken>
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1 ♀ and 1 ♂ from Douglas County, Umpqua National Forest, Umpqua River (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="43.29111">43°17'28"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-122.62">122°37'12"W</geoCoordinate>
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), 12 Aug 2013, by JC
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<normalizedToken originalValue="O’Neill">O'Neill</normalizedToken>
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, & WA Nelson, JNOW 13-0812-006.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type deposition">
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<paragraph pageId="0" pageNumber="1">Type deposition.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype (♀), allotype (♂), and most paratypes (5 ♀; 6 ♂) deposited in the CNC; other paratypes (4 ♀; 5 ♂) deposited in the ACUA.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei</taxonomicName>
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are similar to other members of the Tricolor Complex (
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<taxonomicName lsidName="T. bittikoferae" pageId="0" pageNumber="1" rank="species" species="bittikoferae">T. bittikoferae</taxonomicName>
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,
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<taxonomicName lsidName="T. hoosieri" pageId="0" pageNumber="1" rank="species" species="hoosieri">T. hoosieri</taxonomicName>
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,
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<taxonomicName lsidName="T. larvata" pageId="0" pageNumber="1" rank="species" species="larvata">T. larvata</taxonomicName>
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,
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<taxonomicName lsidName="T. pearsoni" pageId="0" pageNumber="1" rank="species" species="pearsoni">T. pearsoni</taxonomicName>
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,
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<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
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,
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<taxonomicName lsidName="T. tricolor" pageId="0" pageNumber="1" rank="species" species="tricolor">T. tricolor</taxonomicName>
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,
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<taxonomicName lsidName="T. trimaculata" pageId="0" pageNumber="1" rank="species" species="trimaculata">T. trimaculata</taxonomicName>
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,
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<taxonomicName lsidName="T. unimaculata" pageId="0" pageNumber="1" rank="species" species="unimaculata">T. unimaculata</taxonomicName>
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,
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<taxonomicName lsidName="T. cardia" pageId="0" pageNumber="1" rank="species" species="cardia">T. cardia</taxonomicName>
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,
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<taxonomicName lsidName="T. kringi" pageId="0" pageNumber="1" rank="species" species="kringi">T. kringi</taxonomicName>
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,
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<taxonomicName lsidName="T. dimorpha" pageId="0" pageNumber="1" rank="species" species="dimorpha">T. dimorpha</taxonomicName>
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, and
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<taxonomicName lsidName="T. mohawk" pageId="0" pageNumber="1" rank="species" species="mohawk">T. mohawk</taxonomicName>
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) in having a short, conical rostrum.
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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can be differentiated from most
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
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, including most other members of the Tricolor Complex, by having a very short rostrum (length/width = 1.56-1.81 in
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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, 1.80-3.13 in others), except
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<taxonomicName lsidName="T. bittikoferae" pageId="0" pageNumber="1" rank="species" species="bittikoferae">T. bittikoferae</taxonomicName>
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(1.81-1.9) and
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<taxonomicName lsidName="T. dimorpha" pageId="0" pageNumber="1" rank="species" species="dimorpha">T. dimorpha</taxonomicName>
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(1.3-1.6). Additionally, males have a shorter anterior venter than all other Tricolor Complex (♂ = 205-225 in
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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, 230-335 in others), except
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<taxonomicName lsidName="T. kringi" pageId="0" pageNumber="1" rank="species" species="kringi">T. kringi</taxonomicName>
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(220-255), and both males and females have a wider genital field than most other Tricolor Complex (♀ = 190-202.5 in
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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, 145-180 in others; ♂ = 130-137.5 in
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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, 92.5-120 in others) except for
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<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
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(♀ = 180-212.5; ♂ = 135-175) and female
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<taxonomicName lsidName="T. leviathan" pageId="0" pageNumber="1" rank="species" species="leviathan">T. leviathan</taxonomicName>
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(180-195). Finally,
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<taxonomicName lsidName="T. olliei" pageId="0" pageNumber="1" rank="species" species="olliei">T. olliei</taxonomicName>
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can be differentiated from most Tricolor Complex, except for
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<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
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, by being distributed in the west (all others are eastern).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Female (Figure 176) (n=5) (holotype measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (690-900 (690) long; 500-670 (500) wide) ovoid with orange coloration separated into posterior and anterior portions with faint orange medially. Anterio-medial platelets (130-172.5 (130) long; 60-80 (60) wide). Anterio-lateral platelets (190-235 (190) long; 70-92.5 (70) wide) free from dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 360-470 (360)). Dorsal plate proportions: dorsum length/width 1.34-1.42 (1.38); dorsal width/distance between Dgl-4 1.31-1.43 (1.39); anterio-medial platelet length/width 1.95-2.23 (2.17); anterio-lateral platelet length/width 2.34-2.71 (2.71); anterio-lateral/anterio-medial length 1.36-1.46 (1.46).</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (245-287.5 (245) long (ventral); 155-225 (155) long (dorsal); 137.5-152.5 (137.5) tall) colorless and tall. Rostrum (85-112.5 (85) long; 50-67.5 (50) wide) short and conical. Chelicerae (225-305 (225) long) with curved fangs (55-65 (55) long). Subcapitular proportions: ventral length/height 1.78-1.92 (1.78); rostrum length/width 1.62-1.76 (1.70). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (37.5-50 long); femur (87.5-122.5 (87.55) long); genu (58.75-82.5 (60) long); tibia (75-102.5 (75) long; 25-33.75 (25) wide); tarsus (25-30 (26.25) long). Palpomere proportions: femur/genu 1.46-1.51 (1.46); tibia/femur 0.84-0.90 (0.86); tibia length/width 3.00-3.08 (3.00).</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (790-1050 (810) long; 550-765 (550) wide) colorless. Gnathosomal bay (120-170 (150) long; 75-97.5 (75) wide). Cxgl-4 subapical. Medial suture (12.5-50 (12.5) long). Genital plates (202.5-230 (212.5) long; 190-202.5 (190) wide). Additional measurements: Cx-1 (260-375 (280) long (total); 130-195 (130) long (medial)); Cx-3 (350-465 (350) wide); anterior venter (162.5-255 (162.5) long). Ventral proportions: gnathosomal bay length/width 1.37-2.00 (2.00); anterior venter/genital field length 0.76-1.11 (0.76); anterior venter length/genital field width 0.85-1.28 (0.86); anterior venter/medial suture 5.10-13.00 (13.00).</paragraph>
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<paragraph pageId="0" pageNumber="1">Male (Figure 177) (n = 5) (allotypic measurements in parentheses when available) with characters of the genus with following specifications.</paragraph>
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<paragraph pageId="0" pageNumber="1">Dorsum - (560-640 (570) long; 410-460 (410) wide) ovoid with orange coloration separated into posterior and anterior portions with faint orange medially. Anterio-medial platelets (102.5-125 (107.5) long; 50-70 (56.25) wide). Anterio-lateral platelets (170-187.5 (170) long; 62.5-80 (62.5) wide) free from dorsal plate. Dgl-4 much closer to the edge of the dorsum than to the muscle scars (distance between Dgl-4 320-375 (320)). Dorsal plate proportions: dorsum length/width 1.33-1.44 (1.39); dorsal width/distance between Dgl-4 1.19-1.33 (1.28); anterio-medial platelet length/width 1.71-2.25 (1.91); anterio-lateral platelet length/width 2.34-2.72 (2.72); anterio-lateral/anterio-medial length 1.50-1.68 (1.58).</paragraph>
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<paragraph pageId="0" pageNumber="1">Gnathosoma - Subcapitulum (197.5-220 (212.5) long (ventral); 147.5-160 (159) long (dorsal); 115-125 (115) tall) colorless and tall. Rostrum (70-82.5 (72.5) long; 40-50 (40) wide) short and conical. Chelicerae (195-220 long) with curved fangs (47.5-52.5 long). Subcapitular proportions: ventral length/height 1.65-1.85 (1.85); rostrum length/width 1.56-1.81 (1.81). Pedipalps with tuberculate ventral extensions on femora and genua. Palpomeres: trochanter (32.5-35 (32.5) long); femur (75-77.5 (76.25) long); genu (52.5-55 (52.5) long); tibia (67.5-75 (67.5) long; 22.5-25 (22.5) wide); tarsus (20-25 (20) long). Palpomere proportions: femur/genu 1.41-1.48 (1.45); tibia/femur 0.89-0.97 (0.89); tibia length/width 2.70-3.00 (3.00).</paragraph>
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<paragraph pageId="0" pageNumber="1">Venter - (650-710 (650) long; 444-530 (444) wide) colorless. Gnathosomal bay (122.5-137.5 (127.5) long; 62.5-75 (62.5) wide). Cxgl-4 subapical. Medial suture (70-80 (75) long). Genital plates (165-172.5 (165) long; 130-137.5 (130) wide). Additional measurements: Cx-1 (234-270 (235) long (total); 102-135 (102) long (medial)); Cx-3 (311-355 (312) wide); anterior venter (200-225 (207.5) long). Ventral proportions: gnathosomal bay length/width 1.70-2.04 (2.04); anterior venter/genital field length 1.19-1.30 (1.26); anterior venter length/genital field width 1.54-1.67 (1.60); anterior venter/medial suture 2.67-2.93 (2.77).</paragraph>
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<paragraph pageId="0" pageNumber="1">Immatures unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Specific epithet (
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<taxonomicName class="Arachnida" family="Arrenuridae" genus="Lebertioidea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lebertioidea olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">olliei</taxonomicName>
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) named in honor of Ollie-pet Boston Terrier of JRF and DMF-whose brachycephaly resembles the short rostrum of this species, the shortest of all
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Torrenticola</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Pacific coastal mountains (Figure 175).</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 175.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei</taxonomicName>
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sp. n. distribution.
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</paragraph>
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</caption>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 176.
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<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei</taxonomicName>
|
||
sp. n. female: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Figure 177.
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei</taxonomicName>
|
||
sp. n. male: A dorsal plates B venter (legs removed) C subcapitulum D pedipalp (setae not accurately depicted). Scale = 100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName class="Arachnida" family="Torrenticolidae" genus="Torrenticola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Torrenticola olliei" order="Trombidiformes" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="olliei">Torrenticola olliei</taxonomicName>
|
||
groups with other members of the Tricolor Complex with high support in all analyses and group with
|
||
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
|
||
to form the western portion of this complex. Only one specimen was analyzed phylogenetically, so intraspecific variation remains unknown, but that specimen is greater than 6% different from all specimens of
|
||
<taxonomicName lsidName="T. sierrensis" pageId="0" pageNumber="1" rank="species" species="sierrensis">T. sierrensis</taxonomicName>
|
||
. This species hypothesis is supported by high divergence between species (3-15%), and by the morphological characters outlined in the diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |