631 lines
66 KiB
XML
631 lines
66 KiB
XML
<document id="A37CEE9F590D133B67964BE9D8092983" ID-CLB-Dataset="46698" ID-DOI="10.11646/zootaxa.3753.6.3" ID-GBIF-Dataset="715c7010-c4b2-4910-b707-62a84ba9a6d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="224299" ID-ZooBank="8DF3EFAA-F8CC-4D4C-A830-D07D76C2D1C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460366938684" checkinUser="plazi" docAuthor="Fragoso, Carlos & Rojas, Patricia" docDate="2014" docId="FD1F87899A43FF9D1DECAB4FFE6AF8F5" docLanguage="en" docName="zt03753p572.pdf" docOrigin="Zootaxa 3753 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ramiellona tojolabala Fragoso & Rojas, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="553" masterDocId="0126FFF19A42FF991D7BAE78FFDBFFB9" masterDocTitle="New species and records of the earthworm genus Ramiellona (Annelida, Oligochaeta, Acanthodrilidae) from southern Mexico and Guatemala" masterLastPageNumber="572" masterPageNumber="549" pageNumber="550" updateTime="1698295306276" updateUser="ExternalLinkService">
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<mods:mods id="29DD8B3D58F371DD0E47E2DCD7305A5A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6C9117D98D33E81BB4AAD1F1924110CF">
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<mods:title id="8EE68FA5336D43F3E08ACEB76D7B685C">New species and records of the earthworm genus Ramiellona (Annelida, Oligochaeta, Acanthodrilidae) from southern Mexico and Guatemala</mods:title>
|
||
</mods:titleInfo>
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<mods:name id="02A0C73A8EB975328B4DAF789451F259" type="personal">
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<mods:role id="020B059088FDCCDD82E4B55FB324274E">
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<mods:roleTerm id="14E105722106E5B2820631B772534F6D">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="1C302DC62A520968A79EA7B05C280DEC">Fragoso, Carlos</mods:namePart>
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</mods:name>
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||
<mods:name id="D303872DD6B1F88A3AB48876BE6FB14A" type="personal">
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<mods:role id="0A8E75D49D301512554BEB69A7B6E8BC">
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<mods:roleTerm id="2A00869054EA122325DE30C2578F853F">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="A356AE87C5B6758DF48A510A58547B1E">Rojas, Patricia</mods:namePart>
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</mods:name>
|
||
<mods:typeOfResource id="1FC7350183D669F2FE17B820003B6EE3">text</mods:typeOfResource>
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<mods:titleInfo id="AAB85E21D0E114FE287B0152B2B7E173">
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||
<mods:title id="77B82EBC00A873C7C1CF88E7A4807BF8">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="A1F7151CE804EE41673C73946744115D">
|
||
<mods:date id="9EAC9B54CCC03B4750ED32A943210FB0">2014</mods:date>
|
||
<mods:detail id="A38D853388FF9DED8B44BD6541A88EE4" type="volume">
|
||
<mods:number id="1A4A7A18E58CD9E7324FED2B7534BF07">3753</mods:number>
|
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</mods:detail>
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<mods:detail id="0D653E8D5D84409C11E813CB5AD81AEA" type="issue">
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<mods:number id="B458775C55B2822013FC50D55DFB5A91">6</mods:number>
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<mods:start id="423110BF1948650B498C8830C4C77E65">549</mods:start>
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<mods:end id="D27E00E04D6EA9996952C6855F10B0EC">572</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="BFACC20D5448A1A878F225D8C10BD6AA">journal article</mods:classification>
|
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<mods:identifier id="754A55B286E8620A4B622A6BCC1DECFA" type="CLB-Dataset">46698</mods:identifier>
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<mods:identifier id="3A5C5DD7436CCF8448DB9E76D428D1BC" type="DOI">10.11646/zootaxa.3753.6.3</mods:identifier>
|
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<mods:identifier id="D52A9AF2B31A2F5AB29DA03D24D66C3E" type="GBIF-Dataset">715c7010-c4b2-4910-b707-62a84ba9a6d0</mods:identifier>
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<mods:identifier id="4A250AD76C88C5CA9D996FBA45E870FC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A6F7ACCAD97AE1C94CDEEF1376BC73A4" type="Zenodo-Dep">224299</mods:identifier>
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<mods:identifier id="141F9F19D3597FB231A83D57D8AC2943" type="ZooBank">8DF3EFAA-F8CC-4D4C-A830-D07D76C2D1C0</mods:identifier>
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</mods:mods>
|
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<treatment id="FD1F87899A43FF9D1DECAB4FFE6AF8F5" ID-DOI="http://doi.org/10.5281/zenodo.5616737" ID-GBIF-Taxon="119411010" ID-Zenodo-Dep="5616737" LSID="urn:lsid:plazi:treatment:FD1F87899A43FF9D1DECAB4FFE6AF8F5" httpUri="http://treatment.plazi.org/id/FD1F87899A43FF9D1DECAB4FFE6AF8F5" lastPageId="4" lastPageNumber="553" pageId="1" pageNumber="550">
|
||
<subSubSection id="3DAC65149A43FF981DECAB4FFEC6FACA" pageId="1" pageNumber="550" type="nomenclature">
|
||
<paragraph id="7509369F9A43FF981DECAB4FFDD3FAEB" blockId="1.[151,520,1335,1395]" box="[151,520,1335,1362]" pageId="1" pageNumber="550">
|
||
<heading id="2E4181F39A43FF981DECAB4FFDD3FAEB" bold="true" box="[151,520,1335,1362]" fontSize="11" level="1" pageId="1" pageNumber="550" reason="1">
|
||
<emphasis id="47C2EA8D9A43FF981DECAB4FFDD3FAEB" bold="true" box="[151,520,1335,1362]" pageId="1" pageNumber="550">
|
||
<taxonomicName id="B2B64D1C9A43FF981DECAB4FFE79FAE8" box="[151,418,1335,1361]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="550" phylum="Annelida" rank="species" species="tojolabala" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A43FF981DECAB4FFE79FAE8" bold="true" box="[151,418,1335,1361]" italics="true" pageId="1" pageNumber="550">Ramiellona tojolabala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A43FF981CD1AB40FDD3FAEB" box="[426,520,1336,1362]" pageId="1" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7509369F9A43FF981DECAB22FEC6FACA" blockId="1.[151,520,1335,1395]" box="[151,285,1370,1395]" pageId="1" pageNumber="550">
|
||
<figureCitation id="ED8D2A1A9A43FF981DECAB22FEDBFACA" box="[151,256,1370,1395]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="1" pageNumber="550">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="ED8D2A1A9A43FF981C75AB23FEC6FACA" box="[270,285,1371,1395]" captionStart="FIGURE 2" captionStartId="3.[151,255,1234,1257]" captionTargetBox="[263,1323,333,1193]" captionTargetId="figure@3.[263,1323,333,1193]" captionTargetPageId="3" captionText="FIGURE 2. Ramiellona tojolabala sp. nov. A. Penial seta a of segment 19 (scale 200 µm). SEM photographs. B. Apex of penial seta 19 a (scale 50 µm). C. Spermathecal seta a of segment 7 (scale 100 µm). D. Apex of spermathecal seta 7 a (scale 20 µm). Paratype IEOL 2149." httpUri="https://zenodo.org/record/224301/files/figure.png" pageId="1" pageNumber="550">2</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DAC65149A43FF981DECABDAFCFAF80A" pageId="1" pageNumber="550" type="materials_examined">
|
||
<paragraph id="7509369F9A43FF981DECABDAFF33F947" blockId="1.[151,1437,1442,1971]" pageId="1" pageNumber="550">
|
||
<emphasis id="47C2EA8D9A43FF981DECABDAFE68FA02" bold="true" box="[151,435,1442,1467]" pageId="1" pageNumber="550">Localities and material.</emphasis>
|
||
<collectingCountry id="0DA1760F9A43FF981CC0ABDAFDC8FA02" box="[443,531,1442,1467]" name="Mexico" pageId="1" pageNumber="550">Mexico</collectingCountry>
|
||
, Chiapas, National Park Lagunas de Montebello:
|
||
<emphasis id="47C2EA8D9A43FF98193DABDAFBBDFA02" bold="true" box="[1094,1126,1442,1467]" pageId="1" pageNumber="550">(1)</emphasis>
|
||
<typeStatus id="AA0D883D9A43FF981914ABDBFB7CFA02" box="[1135,1191,1443,1467]" pageId="1" pageNumber="550">Type</typeStatus>
|
||
locality: 3.5 km after the entrance to the park, towards Tziscao. In the forests at right hand of the entrance to “Lago de Montebello”,
|
||
<quantity id="B24E9B7A9A43FF981809ABBFFF76F9BB" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="550" unit="m" value="100.0">100 m</quantity>
|
||
away from the lake shore. Slightly disturbed pine-oak forest over a N-faced slope; within soil at
|
||
<quantity id="B24E9B7A9A43FF981995AB93FA94F9BB" box="[1262,1359,1515,1539]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="1" pageNumber="550" unit="cm" value="10.0" valueMax="20.0" valueMin="0.0">0–20 cm</quantity>
|
||
depth.
|
||
<geoCoordinate id="108250589A43FF981DECA876FEF8F99E" box="[151,291,1550,1575]" direction="north" orientation="latitude" pageId="1" pageNumber="550" precision="15" value="16.104445">16°06’16”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="108250589A43FF981C55A877FE60F99E" box="[302,443,1551,1575]" direction="west" orientation="longitude" pageId="1" pageNumber="550" precision="15" value="-91.703606">91º42’13”W</geoCoordinate>
|
||
,
|
||
<quantity id="B24E9B7A9A43FF981CBDA877FDC0F99F" box="[454,539,1551,1575]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="1" pageNumber="550" unit="m" value="1490.0">1490 m</quantity>
|
||
a.s.l.. 19J-23A-15CA (with 2, 9, and 4 posterior amputees, respectively),
|
||
<date id="0108105F9A43FF98182EA877FF09F9F2" pageId="1" pageNumber="550" value="1981-11-19">11/19/ 1981</date>
|
||
, C. Fragoso and P. Lavelle. 6J-4A-3CA,
|
||
<date id="0108105F9A43FF981FDBA84BFCFBF9F2" box="[672,800,1587,1611]" pageId="1" pageNumber="550" value="1997-11-27">11/27/1997</date>
|
||
, C. Fragoso and J. Bueno.
|
||
<emphasis id="47C2EA8D9A43FF98192AA84AFBA9F9F2" bold="true" box="[1105,1138,1586,1611]" pageId="1" pageNumber="550">(2)</emphasis>
|
||
<quantity id="B24E9B7A9A43FF981901A84BFB68F9F2" box="[1146,1203,1586,1611]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="550" unit="km" value="2.0">2 km</quantity>
|
||
after the entrance to the Park, towards Tziscao
|
||
<quantity id="B24E9B7A9A43FF981CBBA82FFDD3F9D7" box="[448,520,1623,1647]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="550" unit="m" value="300.0">300 m</quantity>
|
||
by a rural road. Ecotone cloud forest - pine oak forest with abundant bromeliads, within soil at
|
||
<quantity id="B24E9B7A9A43FF981C54A803FE4BF92B" box="[303,400,1659,1683]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="1" pageNumber="550" unit="cm" value="10.0" valueMax="20.0" valueMin="0.0">0–20 cm</quantity>
|
||
depth.
|
||
<geoCoordinate id="108250589A43FF981C98A802FDBAF92A" box="[483,609,1658,1683]" direction="north" orientation="latitude" pageId="1" pageNumber="550" precision="15" value="16.077223">16°4’38”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="108250589A43FF981F17A803FD27F92A" box="[620,764,1659,1683]" direction="west" orientation="longitude" pageId="1" pageNumber="550" precision="15" value="-91.719444">91°43’10”W</geoCoordinate>
|
||
,
|
||
<quantity id="B24E9B7A9A43FF981E7CA802FC87F92B" box="[775,860,1658,1683]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="1" pageNumber="550" unit="m" value="1540.0">1540 m</quantity>
|
||
a.s.l., 8J-1A,
|
||
<date id="0108105F9A43FF981E8DA803FBACF92A" box="[1014,1143,1659,1683]" pageId="1" pageNumber="550" value="1997-11-27">11/27/1997</date>
|
||
, C. Fragoso and J. Bueno.
|
||
<emphasis id="47C2EA8D9A43FF981DECA8E6FF63F90E" bold="true" box="[151,184,1694,1719]" pageId="1" pageNumber="550">(3)</emphasis>
|
||
km 6 on route to Tziscao,
|
||
<quantity id="B24E9B7A9A43FF981C91A8E7FDFFF90E" box="[490,548,1694,1719]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="550" unit="km" value="4.0">4 km</quantity>
|
||
after “Cinco lagos”. Well preserved cloud forest and adjacent recently cleared forest, within soil at
|
||
<quantity id="B24E9B7A9A43FF981C04A8BBFE3BF963" box="[383,480,1731,1755]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="1" pageNumber="550" unit="cm" value="10.0" valueMax="20.0" valueMin="0.0">0–20 cm</quantity>
|
||
depth.
|
||
<geoCoordinate id="108250589A43FF981F4FA8BAFD7FF962" box="[564,676,1730,1755]" direction="north" orientation="latitude" pageId="1" pageNumber="550" precision="15" value="16.117222">16°7’2”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="108250589A43FF981FCBA8BBFCE9F962" box="[688,818,1731,1755]" direction="west" orientation="longitude" pageId="1" pageNumber="550" precision="15" value="-91.668884">91°40’8”W</geoCoordinate>
|
||
,
|
||
<quantity id="B24E9B7A9A43FF981E46A8BAFC48F963" box="[829,915,1730,1755]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="550" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l., 2A-4CA,
|
||
<date id="0108105F9A43FF981930A8BBFB17F962" box="[1099,1228,1731,1755]" pageId="1" pageNumber="550" value="1997-11-28">11/28/1997</date>
|
||
, C. Fragoso and J. Bueno.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A43FF981DBCA972FBE7F89A" blockId="1.[151,1437,1442,1971]" box="[199,1084,1802,1827]" pageId="1" pageNumber="550">
|
||
<emphasis id="47C2EA8D9A43FF981DBCA972FEE0F89A" bold="true" box="[199,315,1802,1827]" pageId="1" pageNumber="550">
|
||
<typeStatus id="AA0D883D9A43FF981DBCA972FEEDF89A" box="[199,310,1802,1827]" pageId="1" pageNumber="550" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
One clitellate adult collected
|
||
<date id="0108105F9A43FF981FF1A973FCD3F89A" box="[650,776,1803,1827]" pageId="1" pageNumber="550" value="1997-11-27">
|
||
11/27/
|
||
<quantity id="B24E9B7A9A43FF981FABA973FCD3F89A" box="[720,776,1803,1827]" metricMagnitude="1" metricUnit="m" metricValue="5.07238" pageId="1" pageNumber="550" unit="in" value="1997.0">1997</quantity>
|
||
</date>
|
||
in locality
|
||
<emphasis id="47C2EA8D9A43FF981EFCA972FC7CF89A" bold="true" box="[903,935,1802,1827]" pageId="1" pageNumber="550">(1)</emphasis>
|
||
,
|
||
<collectionCode id="13A7AE5A9A43FF981ECDA973FC2DF89A" box="[950,1014,1803,1827]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
4238.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A43FF981DBCA956FCFAF80A" blockId="1.[151,1437,1442,1971]" pageId="1" pageNumber="550">
|
||
<emphasis id="47C2EA8D9A43FF981DBCA956FE9DF8FE" bold="true" box="[199,326,1838,1863]" pageId="1" pageNumber="550">
|
||
<typeStatus id="AA0D883D9A43FF981DBCA956FE9AF8FE" box="[199,321,1838,1863]" pageId="1" pageNumber="550" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Five clitellate adults collected in locality
|
||
<emphasis id="47C2EA8D9A43FF981E51A956FC91F8FE" bold="true" box="[810,842,1838,1863]" pageId="1" pageNumber="550">(1)</emphasis>
|
||
,
|
||
<date id="0108105F9A43FF981E20A957FC06F8FE" box="[859,989,1839,1863]" pageId="1" pageNumber="550" value="1981-11-19">11/19/1981</date>
|
||
:
|
||
<collectionCode id="13A7AE5A9A43FF981E91A957FBF1F8FE" box="[1002,1066,1839,1863]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
2149, 2150, 2151, 2155, and in locality
|
||
<emphasis id="47C2EA8D9A43FF981D89A92AFEC9F8D2" bold="true" box="[242,274,1874,1899]" pageId="1" pageNumber="550">(3)</emphasis>
|
||
,
|
||
<collectionCode id="13A7AE5A9A43FF981C5BA92BFEBBF8D2" box="[288,352,1875,1899]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
3336, dissected. Further 20 individuals, characterized only externally: localities
|
||
<emphasis id="47C2EA8D9A43FF98199CA92AFADCF8D2" bold="true" box="[1255,1287,1874,1899]" pageId="1" pageNumber="550">(1)</emphasis>
|
||
,
|
||
<date id="0108105F9A43FF98186EA92BFA43F8D2" box="[1301,1432,1875,1899]" pageId="1" pageNumber="550" value="1981-11-19">11/19/1981</date>
|
||
:
|
||
<collectionCode id="13A7AE5A9A43FF981DECA90FFF0CF836" box="[151,215,1911,1935]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
2154, 3360, 4229, 4230, 4231, 4232, 4233, 4234, 4235, 4236, 4237;
|
||
<date id="0108105F9A43FF981E8DA90FFBA3F836" box="[1014,1144,1911,1935]" pageId="1" pageNumber="550" value="1997-11-27">11/27/1997</date>
|
||
:
|
||
<collectionCode id="13A7AE5A9A43FF9819FEA90FFB1EF836" box="[1157,1221,1911,1935]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
4219, 4220, 4239, 4243;
|
||
<emphasis id="47C2EA8D9A43FF981DA6A9E2FF25F80A" bold="true" box="[221,254,1946,1971]" pageId="1" pageNumber="550">(2)</emphasis>
|
||
,
|
||
<collectionCode id="13A7AE5A9A43FF981C77A9E3FE97F80A" box="[268,332,1947,1971]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
3320;
|
||
<emphasis id="47C2EA8D9A43FF981CE1A9E2FE60F80A" bold="true" box="[410,443,1946,1971]" pageId="1" pageNumber="550">(3)</emphasis>
|
||
,
|
||
<collectionCode id="13A7AE5A9A43FF981CB3A9E3FDD3F80A" box="[456,520,1947,1971]" pageId="1" pageNumber="550">IEOL</collectionCode>
|
||
4240, 4241, 4242, 4243.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="21C966179A40FF9B1DECABDBFC39FA42" httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="2" pageNumber="551" targetBox="[403,1182,200,1415]" targetPageId="2">
|
||
<paragraph id="7509369F9A40FF9B1DECABDBFC39FA42" blockId="2.[151,1436,1443,1531]" pageId="2" pageNumber="551">
|
||
<emphasis id="47C2EA8D9A40FF9B1DECABDBFDB6FA03" bold="true" box="[151,621,1443,1467]" pageId="2" pageNumber="551">
|
||
FIGURE 1.
|
||
<taxonomicName id="B2B64D1C9A40FF9B1C59ABDBFDCCFA02" box="[290,535,1443,1467]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="2" pageNumber="551" phylum="Annelida" rank="species" species="tojolabala" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A40FF9B1C59ABDBFDCCFA02" bold="true" box="[290,535,1443,1467]" italics="true" pageId="2" pageNumber="551">Ramiellona tojolabala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A40FF9B1F65ABDBFDB6FA03" box="[542,621,1443,1466]" pageId="2" pageNumber="551" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="47C2EA8D9A40FF9B1F00ABDBFD4EFA03" bold="true" box="[635,661,1443,1466]" pageId="2" pageNumber="551">A.</emphasis>
|
||
External ventral view of anterior region.
|
||
<emphasis id="47C2EA8D9A40FF9B192DABDBFBB5FA03" bold="true" box="[1110,1134,1443,1466]" pageId="2" pageNumber="551">B.</emphasis>
|
||
Spermatheca from segment
|
||
<emphasis id="47C2EA8D9A40FF9B1DECABBCFF7FFA62" bold="true" box="[151,164,1476,1499]" pageId="2" pageNumber="551">9</emphasis>
|
||
, dorsal view.
|
||
<emphasis id="47C2EA8D9A40FF9B1C41ABBCFE88FA62" bold="true" box="[314,339,1476,1499]" pageId="2" pageNumber="551">C.</emphasis>
|
||
Transverse section (hand made dissection) from segment
|
||
<emphasis id="47C2EA8D9A40FF9B1EBAABBCFC15FA62" bold="true" box="[961,974,1476,1499]" pageId="2" pageNumber="551">9</emphasis>
|
||
, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3DAC65149A40FF9D1DBCA85CFD2DFA24" lastPageId="4" lastPageNumber="553" pageId="2" pageNumber="551" type="description">
|
||
<paragraph id="7509369F9A40FF9A1DBCA85CFA47FF41" blockId="2.[151,1436,1572,2029]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="552" pageId="2" pageNumber="551">
|
||
<emphasis id="47C2EA8D9A40FF9B1DBCA85CFE0BF984" bold="true" box="[199,464,1572,1597]" pageId="2" pageNumber="551">Description. External.</emphasis>
|
||
Length
|
||
<quantity id="B24E9B7A9A40FF9B1F49A85CFD61F985" box="[562,698,1572,1597]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="2" pageNumber="551" unit="mm" value="85.0" valueMax="120.0" valueMin="50.0">50–120 mm</quantity>
|
||
(mean=78.9 mm, n=19;
|
||
<typeStatus id="AA0D883D9A40FF9B1EA2A85CFBE0F984" box="[985,1083,1572,1597]" pageId="2" pageNumber="551" type="holotype">holotype</typeStatus>
|
||
<quantity id="B24E9B7A9A40FF9B193FA85CFB7CF985" box="[1092,1191,1572,1596]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="2" pageNumber="551" unit="mm" value="103.0">103 mm</quantity>
|
||
); width (postclitellar) 1.6–2.5 mm (mean=2.11, n=23;
|
||
<typeStatus id="AA0D883D9A40FF9B1F7FA830FDBDF9D8" box="[516,614,1608,1633]" pageId="2" pageNumber="551" type="holotype">holotype</typeStatus>
|
||
1.8 mm). Segments 128–239 (mean=189, n=14;
|
||
<typeStatus id="AA0D883D9A40FF9B19E8A830FB2EF9D8" box="[1171,1269,1608,1633]" pageId="2" pageNumber="551" type="holotype">holotype</typeStatus>
|
||
213). Pigment absent. Prostomium prolobous, invaginated in the majority of individuals. Peristomium with several longitudinal annulations; in several individuals these grooves also present in the second segment. Secondary furrows: one postsetal in
|
||
<emphasis id="47C2EA8D9A40FF9B1C58A8CCFEEAF974" bold="true" box="[291,305,1716,1741]" pageId="2" pageNumber="551">7</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A40FF9B1C15A8CCFEA7F974" bold="true" box="[366,380,1716,1741]" pageId="2" pageNumber="551">8</emphasis>
|
||
; in
|
||
<emphasis id="47C2EA8D9A40FF9B1CD4A8CCFE3CF974" bold="true" box="[431,487,1716,1741]" pageId="2" pageNumber="551">9–13</emphasis>
|
||
one presetal and one postsetal; behind clitellum generally absent. Setae eight per segment, visible from segment
|
||
<emphasis id="47C2EA8D9A40FF9B1C8EA8A0FDD8F948" bold="true" box="[501,515,1752,1777]" pageId="2" pageNumber="551">2</emphasis>
|
||
, closely paired in the anterior region and quincuaxial in the posterior region. Setal formula in anterior region (average, n=5) (
|
||
<emphasis id="47C2EA8D9A40FF9B1FE9A884FC9BF8AC" box="[658,832,1788,1813]" italics="true" pageId="2" pageNumber="551">aa:ab:bc:cd:dd</emphasis>
|
||
):
|
||
<emphasis id="47C2EA8D9A40FF9B1E26A884FCA2F8AC" bold="true" box="[861,889,1788,1813]" pageId="2" pageNumber="551">10</emphasis>
|
||
: 2.9:1:7.3:0.6:34.9;
|
||
<emphasis id="47C2EA8D9A40FF9B1913A884FB5FF8AC" bold="true" box="[1128,1156,1788,1813]" pageId="2" pageNumber="551">30</emphasis>
|
||
: 4:1:4.2:1.2:39.1. In all individuals the first seta that shifts is seta
|
||
<emphasis id="47C2EA8D9A40FF9B1F03A959FD5FF881" box="[632,644,1825,1848]" italics="true" pageId="2" pageNumber="551">c</emphasis>
|
||
, then seta
|
||
<emphasis id="47C2EA8D9A40FF9B1E79A958FCCBF880" box="[770,784,1824,1849]" italics="true" pageId="2" pageNumber="551">d</emphasis>
|
||
and finally seta
|
||
<emphasis id="47C2EA8D9A40FF9B1EB4A958FC06F880" box="[975,989,1824,1849]" italics="true" pageId="2" pageNumber="551">b</emphasis>
|
||
; setae
|
||
<emphasis id="47C2EA8D9A40FF9B1955A959FBE7F881" box="[1070,1084,1825,1848]" italics="true" pageId="2" pageNumber="551">a</emphasis>
|
||
never move and thus row
|
||
<emphasis id="47C2EA8D9A40FF9B1814A959FA5BF881" box="[1391,1408,1825,1848]" italics="true" pageId="2" pageNumber="551">A</emphasis>
|
||
is recognized throughout. The quincuaxial arrangement occurs always in the second half of the body, after segment 100 (average beginning after 65% of segments, range 55–80%, n=12). Spermathecal setae present in some individuals (
|
||
<emphasis id="47C2EA8D9A40FF9B1C5CA9F5FEE8F81D" box="[295,307,1933,1956]" italics="true" pageId="2" pageNumber="551">c</emphasis>
|
||
. 50%), on one or both sides of
|
||
<emphasis id="47C2EA8D9A40FF9B1FD4A9F4FD66F81C" bold="true" box="[687,701,1932,1957]" pageId="2" pageNumber="551">6</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A40FF9B1F80A9F4FCD2F81C" bold="true" box="[763,777,1932,1957]" pageId="2" pageNumber="551">7</emphasis>
|
||
(one ind.),
|
||
<emphasis id="47C2EA8D9A40FF9B1EEEA9F4FC78F81C" bold="true" box="[917,931,1932,1957]" pageId="2" pageNumber="551">7</emphasis>
|
||
(14 ind.) or
|
||
<emphasis id="47C2EA8D9A40FF9B1945A9F4FB97F81C" bold="true" box="[1086,1100,1932,1957]" pageId="2" pageNumber="551">7</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A40FF9B19F1A9F4FB43F81C" bold="true" box="[1162,1176,1932,1957]" pageId="2" pageNumber="551">8</emphasis>
|
||
(one ind.), sometimes surrounded by swellings (
|
||
<figureCitation id="ED8D2A1A9A40FF9B1CC4A9C8FDDAF871" box="[447,513,1968,1993]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="2" pageNumber="551">Fig. 1</figureCitation>
|
||
A). Spermathecal setae slightly curved, without ornamentation, measuring 0.73– 0.93 mm, apex slightly arrow-shaped (
|
||
<figureCitation id="ED8D2A1A9A40FF9B1F3DA9ACFD52F855" box="[582,649,2004,2029]" captionStart="FIGURE 2" captionStartId="3.[151,255,1234,1257]" captionTargetBox="[263,1323,333,1193]" captionTargetId="figure@3.[263,1323,333,1193]" captionTargetPageId="3" captionText="FIGURE 2. Ramiellona tojolabala sp. nov. A. Penial seta a of segment 19 (scale 200 µm). SEM photographs. B. Apex of penial seta 19 a (scale 50 µm). C. Spermathecal seta a of segment 7 (scale 100 µm). D. Apex of spermathecal seta 7 a (scale 20 µm). Paratype IEOL 2149." httpUri="https://zenodo.org/record/224301/files/figure.png" pageId="2" pageNumber="551">Fig. 2</figureCitation>
|
||
C,D). Penial setae very conspicuous, robust, paired (
|
||
<emphasis id="47C2EA8D9A40FF9B19A9A9ADFB3BF855" box="[1234,1248,2005,2028]" italics="true" pageId="2" pageNumber="551">a</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A40FF9B186DA9ACFAFFF854" box="[1302,1316,2004,2029]" italics="true" pageId="2" pageNumber="551">b</emphasis>
|
||
) in
|
||
<emphasis id="47C2EA8D9A40FF9B182AA9ACFAB6F854" bold="true" box="[1361,1389,2004,2029]" pageId="2" pageNumber="551">17</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A41FF9A1DECAEEFFF68FF09" bold="true" box="[151,179,151,176]" pageId="3" pageNumber="552">19</emphasis>
|
||
, semicircular (
|
||
<figureCitation id="ED8D2A1A9A41FF9A1C27AEEFFE7BFF09" box="[348,416,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,255,1234,1257]" captionTargetBox="[263,1323,333,1193]" captionTargetId="figure@3.[263,1323,333,1193]" captionTargetPageId="3" captionText="FIGURE 2. Ramiellona tojolabala sp. nov. A. Penial seta a of segment 19 (scale 200 µm). SEM photographs. B. Apex of penial seta 19 a (scale 50 µm). C. Spermathecal seta a of segment 7 (scale 100 µm). D. Apex of spermathecal seta 7 a (scale 20 µm). Paratype IEOL 2149." httpUri="https://zenodo.org/record/224301/files/figure.png" pageId="3" pageNumber="552">Fig. 2</figureCitation>
|
||
A); those of
|
||
<emphasis id="47C2EA8D9A41FF9A1F55AEEFFD91FF09" bold="true" box="[558,586,151,176]" pageId="3" pageNumber="552">17</emphasis>
|
||
always slightly smaller (length 0.93–1.12 mm, width 70 µm) than those of
|
||
<emphasis id="47C2EA8D9A41FF9A1DECAEC4FF68FF6C" bold="true" box="[151,179,188,213]" pageId="3" pageNumber="552">19</emphasis>
|
||
(1.02–1.31 mm, 100 µm). Apex with a slight undulation, ending in an acute tip; ornamentation limited to undulation, consisting in several irregular rows of small, apex oriented thorns (
|
||
<figureCitation id="ED8D2A1A9A41FF9A197FAEA7FB93FF41" box="[1028,1096,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,255,1234,1257]" captionTargetBox="[263,1323,333,1193]" captionTargetId="figure@3.[263,1323,333,1193]" captionTargetPageId="3" captionText="FIGURE 2. Ramiellona tojolabala sp. nov. A. Penial seta a of segment 19 (scale 200 µm). SEM photographs. B. Apex of penial seta 19 a (scale 50 µm). C. Spermathecal seta a of segment 7 (scale 100 µm). D. Apex of spermathecal seta 7 a (scale 20 µm). Paratype IEOL 2149." httpUri="https://zenodo.org/record/224301/files/figure.png" pageId="3" pageNumber="552">Fig. 2</figureCitation>
|
||
B). Setae
|
||
<emphasis id="47C2EA8D9A41FF9A19C9AE99FB1BFF41" box="[1202,1216,225,248]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A41FF9A198EAEA7FAD8FF41" box="[1269,1283,223,248]" italics="true" pageId="3" pageNumber="552">b</emphasis>
|
||
of
|
||
<emphasis id="47C2EA8D9A41FF9A1853AEA7FA9FFF41" bold="true" box="[1320,1348,223,248]" pageId="3" pageNumber="552">18</emphasis>
|
||
visible.
|
||
</paragraph>
|
||
<caption id="21C966179A41FF9A1DECAAAAFDC0FA90" httpUri="https://zenodo.org/record/224301/files/figure.png" pageId="3" pageNumber="552" targetBox="[263,1323,333,1193]" targetPageId="3">
|
||
<paragraph id="7509369F9A41FF9A1DECAAAAFDC0FA90" blockId="3.[151,1436,1233,1321]" pageId="3" pageNumber="552">
|
||
<emphasis id="47C2EA8D9A41FF9A1DECAAAAFDB1FB50" bold="true" box="[151,618,1233,1257]" pageId="3" pageNumber="552">
|
||
FIGURE 2.
|
||
<taxonomicName id="B2B64D1C9A41FF9A1C5BAAA9FDCEFB50" box="[288,533,1233,1257]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="552" phylum="Annelida" rank="species" species="tojolabala" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A41FF9A1C5BAAA9FDCEFB50" bold="true" box="[288,533,1233,1257]" italics="true" pageId="3" pageNumber="552">Ramiellona tojolabala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A41FF9A1F60AAAAFDB1FB50" box="[539,618,1234,1257]" pageId="3" pageNumber="552" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="47C2EA8D9A41FF9A1F0CAAAAFD4AFB50" bold="true" box="[631,657,1234,1257]" pageId="3" pageNumber="552">A.</emphasis>
|
||
Penial seta
|
||
<emphasis id="47C2EA8D9A41FF9A1E69AAAAFCC4FB50" box="[786,799,1234,1257]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
of segment
|
||
<emphasis id="47C2EA8D9A41FF9A1ED8AAAAFC66FB50" bold="true" box="[931,957,1234,1257]" pageId="3" pageNumber="552">19</emphasis>
|
||
(scale 200 µm). SEM photographs.
|
||
<emphasis id="47C2EA8D9A41FF9A1838AAAAFA81FB50" bold="true" box="[1347,1370,1234,1257]" pageId="3" pageNumber="552">B.</emphasis>
|
||
Apex of penial seta
|
||
<emphasis id="47C2EA8D9A41FF9A1C56AA8AFE9CFAB0" bold="true" box="[301,327,1266,1289]" pageId="3" pageNumber="552">19</emphasis>
|
||
<emphasis id="47C2EA8D9A41FF9A1C33AA8AFE8EFAB0" box="[328,341,1266,1289]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
(scale 50 µm).
|
||
<emphasis id="47C2EA8D9A41FF9A1C85AA8AFDC3FAB0" bold="true" box="[510,536,1266,1289]" pageId="3" pageNumber="552">C.</emphasis>
|
||
Spermathecal seta
|
||
<emphasis id="47C2EA8D9A41FF9A1F93AA8AFD2EFAB0" box="[744,757,1266,1289]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
of segment
|
||
<emphasis id="47C2EA8D9A41FF9A1E01AA8AFC5CFAB0" bold="true" box="[890,903,1266,1289]" pageId="3" pageNumber="552">7</emphasis>
|
||
(scale
|
||
<emphasis id="47C2EA8D9A41FF9A1EAFAA8AFC27FAB0" bold="true" box="[980,1020,1266,1289]" pageId="3" pageNumber="552">100</emphasis>
|
||
µm).
|
||
<emphasis id="47C2EA8D9A41FF9A1946AA8AFB8CFAB0" bold="true" box="[1085,1111,1266,1289]" pageId="3" pageNumber="552">D.</emphasis>
|
||
Apex of spermathecal seta
|
||
<emphasis id="47C2EA8D9A41FF9A18FAAA8AFA55FAB0" bold="true" box="[1409,1422,1266,1289]" pageId="3" pageNumber="552">7</emphasis>
|
||
<emphasis id="47C2EA8D9A41FF9A18F5AA8AFA40FAB0" box="[1422,1435,1266,1289]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
(scale 20 µm). Paratype IEOL 2149.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7509369F9A41FF9A1DBCAB2AFEE2F812" blockId="3.[151,1436,1362,2035]" pageId="3" pageNumber="552">
|
||
Clitellum dark to light orange, dorsally in
|
||
<emphasis id="47C2EA8D9A41FF9A1FD5AB2AFD2FFAD2" bold="true" box="[686,756,1362,1387]" pageId="3" pageNumber="552">13–19</emphasis>
|
||
(one ind.),
|
||
<date id="0108105F9A41FF9A1E07AB2AFC1EFAD2" box="[892,965,1362,1387]" pageId="3" pageNumber="552" value="2013-02-01">
|
||
1/2
|
||
<emphasis id="47C2EA8D9A41FF9A1ED2AB2AFC1EFAD2" bold="true" box="[937,965,1362,1387]" pageId="3" pageNumber="552">13</emphasis>
|
||
</date>
|
||
–19 (8 ind.),
|
||
<emphasis id="47C2EA8D9A41FF9A1927AB2AFB79FAD2" bold="true" box="[1116,1186,1362,1387]" pageId="3" pageNumber="552">14–19</emphasis>
|
||
(6 ind.) or
|
||
<date id="0108105F9A41FF9A1852AB2AFA47FAD2" box="[1321,1436,1362,1387]" pageId="3" pageNumber="552" value="1920-02-14" valueMax="1920-02-01" valueMin="1920-02-14">
|
||
<emphasis id="47C2EA8D9A41FF9A1852AB2AFA9EFAD2" bold="true" box="[1321,1349,1362,1387]" pageId="3" pageNumber="552">14</emphasis>
|
||
–1/2
|
||
<emphasis id="47C2EA8D9A41FF9A18FBAB2AFA47FAD2" bold="true" box="[1408,1436,1362,1387]" pageId="3" pageNumber="552">20</emphasis>
|
||
</date>
|
||
(one ind.); ventrally in
|
||
<emphasis id="47C2EA8D9A41FF9A1CDEAB0EFE30FA36" bold="true" box="[421,491,1398,1423]" pageId="3" pageNumber="552">13–19</emphasis>
|
||
(one ind.),
|
||
<date id="0108105F9A41FF9A1F0EAB0FFD64FA36" box="[629,703,1398,1423]" pageId="3" pageNumber="552" value="2013-02-01">
|
||
1/2
|
||
<emphasis id="47C2EA8D9A41FF9A1FD8AB0EFD64FA36" bold="true" box="[675,703,1398,1423]" pageId="3" pageNumber="552">13</emphasis>
|
||
</date>
|
||
–19 (7 ind.),
|
||
<emphasis id="47C2EA8D9A41FF9A1E23AB0EFC45FA36" bold="true" box="[856,926,1398,1423]" pageId="3" pageNumber="552">14–19</emphasis>
|
||
(7 ind.) or
|
||
<date id="0108105F9A41FF9A1953AB0FFBA9FA36" box="[1064,1138,1398,1423]" pageId="3" pageNumber="552" value="2013-04-01">
|
||
1/4
|
||
<emphasis id="47C2EA8D9A41FF9A192DAB0EFBA9FA36" bold="true" box="[1110,1138,1398,1423]" pageId="3" pageNumber="552">13</emphasis>
|
||
</date>
|
||
–1/4
|
||
<emphasis id="47C2EA8D9A41FF9A19D6AB0EFB12FA36" bold="true" box="[1197,1225,1398,1423]" pageId="3" pageNumber="552">20</emphasis>
|
||
(one ind.); saddle shaped in the region of the genital zone (
|
||
<emphasis id="47C2EA8D9A41FF9A1F0DABE2FD67FA0A" bold="true" box="[630,700,1434,1459]" pageId="3" pageNumber="552">17–19</emphasis>
|
||
) and (in some individuals) in
|
||
<emphasis id="47C2EA8D9A41FF9A1964ABE2FBBEFA0A" bold="true" box="[1055,1125,1434,1459]" pageId="3" pageNumber="552">15–16</emphasis>
|
||
, reaching setae
|
||
<emphasis id="47C2EA8D9A41FF9A1859ABE2FAEBFA0A" box="[1314,1328,1434,1459]" italics="true" pageId="3" pageNumber="552">b</emphasis>
|
||
; in some individuals fused with swellings of segments
|
||
<emphasis id="47C2EA8D9A41FF9A1FD7ABC6FD2AFA6E" bold="true" box="[684,753,1470,1495]" pageId="3" pageNumber="552">14–16</emphasis>
|
||
. Intersegmental furrows of clitellum recognizable in some individuals (
|
||
<figureCitation id="ED8D2A1A9A41FF9A1C58AB9AFEB9FA43" box="[291,354,1506,1531]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
|
||
A). Large dorsal pores present all along the body, first pore in 12/13 (n=20). Spermathecal pores paired in 7/8 and 8/9, very large, centered in
|
||
<emphasis id="47C2EA8D9A41FF9A1FF7A87FFD74F9A7" box="[652,687,1543,1566]" italics="true" pageId="3" pageNumber="552">AB</emphasis>
|
||
(
|
||
<figureCitation id="ED8D2A1A9A41FF9A1FC5A87EFD27F9A6" box="[702,764,1542,1567]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
|
||
A), the posterior edge of each pore with iridescence. Due to their size, in some individuals the pores appear to be in segments
|
||
<emphasis id="47C2EA8D9A41FF9A1E0BA852FCA5F9FA" bold="true" box="[880,894,1578,1603]" pageId="3" pageNumber="552">8</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A41FF9A1ECFA852FC19F9FA" bold="true" box="[948,962,1578,1603]" pageId="3" pageNumber="552">9</emphasis>
|
||
, but when looking inside the pore the duct is clearly in the respective intersegment. Female pores in
|
||
<emphasis id="47C2EA8D9A41FF9A1E57A836FC93F9DE" bold="true" box="[812,840,1614,1639]" pageId="3" pageNumber="552">14</emphasis>
|
||
, presetal and slightly median to
|
||
<emphasis id="47C2EA8D9A41FF9A19BAA837FB14F9DF" box="[1217,1231,1615,1638]" italics="true" pageId="3" pageNumber="552">a</emphasis>
|
||
, within an ovoidtriangular papilla extending in
|
||
<emphasis id="47C2EA8D9A41FF9A1C8CA80BFDC1F933" box="[503,538,1651,1674]" italics="true" pageId="3" pageNumber="552">BB</emphasis>
|
||
(
|
||
<figureCitation id="ED8D2A1A9A41FF9A1F50A80AFDB7F933" box="[555,620,1650,1675]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
|
||
A). Two pairs of prostatic pores in
|
||
<emphasis id="47C2EA8D9A41FF9A1978A80AFBC4F932" bold="true" box="[1027,1055,1650,1675]" pageId="3" pageNumber="552">17</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A41FF9A1922A80AFBAEF932" bold="true" box="[1113,1141,1650,1675]" pageId="3" pageNumber="552">19</emphasis>
|
||
just at the base of seta
|
||
<emphasis id="47C2EA8D9A41FF9A18FCA80AFA4EF932" box="[1415,1429,1650,1675]" italics="true" pageId="3" pageNumber="552">b</emphasis>
|
||
, joined by square bracket-shaped seminal grooves, which run outside
|
||
<emphasis id="47C2EA8D9A41FF9A1ED8A8EFFC6FF917" box="[931,948,1687,1710]" italics="true" pageId="3" pageNumber="552">B</emphasis>
|
||
(
|
||
<figureCitation id="ED8D2A1A9A41FF9A1EBEA8EEFBDEF916" box="[965,1029,1686,1711]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
|
||
A). In some individuals the seminal groove is more bracket-like, whereas in others it curves slightly in
|
||
<emphasis id="47C2EA8D9A41FF9A1EF6A8C2FC72F96A" bold="true" box="[909,937,1722,1747]" pageId="3" pageNumber="552">18</emphasis>
|
||
towards seta
|
||
<emphasis id="47C2EA8D9A41FF9A193EA8C2FB88F96A" box="[1093,1107,1722,1747]" italics="true" pageId="3" pageNumber="552">b</emphasis>
|
||
. Male pores in 17/18, within the seminal groove, outside
|
||
<emphasis id="47C2EA8D9A41FF9A1CAAA8A7FE39F94F" box="[465,482,1759,1782]" italics="true" pageId="3" pageNumber="552">B</emphasis>
|
||
; in fully clitellate individuals pores difficult to see. Genital marks unpaired (
|
||
<figureCitation id="ED8D2A1A9A41FF9A1847A8A6FAA1F94E" box="[1340,1402,1758,1783]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="3" pageNumber="552">Fig. 1</figureCitation>
|
||
A) as mid-ventral, intersegmental papillae located in some intersegments of region 16/17–23/24; shape ovoidrectangular, extending into
|
||
<emphasis id="47C2EA8D9A41FF9A1CB6A95FFE34F887" box="[461,495,1831,1854]" italics="true" pageId="3" pageNumber="552">AA</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A41FF9A1C85A95FFDFAF887" box="[510,545,1831,1854]" italics="true" pageId="3" pageNumber="552">BB</emphasis>
|
||
or (more commonly) from outside
|
||
<emphasis id="47C2EA8D9A41FF9A1EC8A95FFC1FF887" box="[947,964,1831,1854]" italics="true" pageId="3" pageNumber="552">B</emphasis>
|
||
to outside
|
||
<emphasis id="47C2EA8D9A41FF9A1938A95FFB8FF887" box="[1091,1108,1831,1854]" italics="true" pageId="3" pageNumber="552">B</emphasis>
|
||
. Papillae in 20/21 and 21/22 mostly present (16 ind.).
|
||
<typeStatus id="AA0D883D9A41FF9A1CCCA932FDC4F8DA" box="[439,543,1866,1891]" pageId="3" pageNumber="552" type="holotype">Holotype</typeStatus>
|
||
with papillae in 19/20–22/23. Papillae largest in 2nd or 3rd position, smallest generally in the last position. In some clitellate adults papillae of 17/18, 18/19 difficult to recognize, seeming more like swellings.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A41FF9D1DBCA9CEFC45FE15" blockId="3.[151,1436,1362,2035]" lastBlockId="4.[151,1436,151,1869]" lastPageId="4" lastPageNumber="553" pageId="3" pageNumber="552">
|
||
<emphasis id="47C2EA8D9A41FF9A1DBCA9CEFEEBF876" bold="true" box="[199,304,1974,1999]" pageId="3" pageNumber="552">Internal.</emphasis>
|
||
<collectingCountry id="0DA1760F9A41FF9A1C4CA9CFFEAEF876" box="[311,373,1975,1999]" name="Spain" pageId="3" pageNumber="552">Septa</collectingCountry>
|
||
<date id="0108105F9A41FF9A1C06A9CEFDC3F876" box="[381,536,1974,1999]" pageId="3" pageNumber="552" value="2013-06-05" valueMax="2013-12-05" valueMin="2013-06-05">5/6 and 12/13</date>
|
||
very thin and membranous; 11/12 slightly muscular;
|
||
<date id="0108105F9A41FF9A190FA9CEFB3FF876" box="[1140,1252,1974,1999]" pageId="3" pageNumber="552" value="2011-07-06" valueMax="2011-10-06" valueMin="2011-07-06">6/7–10/11</date>
|
||
muscular; septa
|
||
<date id="0108105F9A41FF9A1DECA9A2FEDCF84A" box="[151,263,2010,2035]" pageId="3" pageNumber="552" value="2011-07-06" valueMax="2011-10-06" valueMin="2011-07-06">6/7–10/11</date>
|
||
funnel-shaped and imbricated, those of
|
||
<date id="0108105F9A41FF9A1FA4A9A2FC8BF84B" box="[735,848,2010,2034]" pageId="3" pageNumber="552" value="2011-09-08" valueMax="2011-10-08" valueMin="2011-09-08">8/9–10/11</date>
|
||
joined by 4–8 dorsal and lateral connective tissue fibers. One large gizzard in
|
||
<emphasis id="47C2EA8D9A46FF9D1CAAAEEFFE3DFF09" bold="true" box="[465,486,151,176]" pageId="4" pageNumber="553">5,</emphasis>
|
||
extending up to
|
||
<emphasis id="47C2EA8D9A46FF9D1FDFAEEFFD16FF09" bold="true" box="[676,717,151,176]" pageId="4" pageNumber="553">7–8</emphasis>
|
||
. Extramural calciferous glands present in
|
||
<emphasis id="47C2EA8D9A46FF9D19D2AEEFFB3AFF09" bold="true" box="[1193,1249,151,176]" pageId="4" pageNumber="553">7–12</emphasis>
|
||
(n=6), as dorsolateral sacs that open into the esophagus through a medium-sized conduct (
|
||
<figureCitation id="ED8D2A1A9A46FF9D1E9BAEC4FBC4FF6D" box="[992,1055,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="4" pageNumber="553">Fig. 1</figureCitation>
|
||
C); internally each sac with large, numerous lamellae with free margins. Size of sacs:
|
||
<emphasis id="47C2EA8D9A46FF9D1F9FAEA7FD29FF41" bold="true" box="[740,754,223,248]" pageId="4" pageNumber="553">7</emphasis>
|
||
≤
|
||
<emphasis id="47C2EA8D9A46FF9D1E68AEA7FC94FF41" bold="true" box="[787,847,223,248]" pageId="4" pageNumber="553">8 ≥ 9</emphasis>
|
||
>
|
||
<emphasis id="47C2EA8D9A46FF9D1E0BAEA7FC57FF41" bold="true" box="[880,908,223,248]" pageId="4" pageNumber="553">10</emphasis>
|
||
>
|
||
<emphasis id="47C2EA8D9A46FF9D1ED6AEA7FC13FF41" bold="true" box="[941,968,223,248]" pageId="4" pageNumber="553">11</emphasis>
|
||
>
|
||
<emphasis id="47C2EA8D9A46FF9D1E93AEA7FBD0FF41" bold="true" box="[1000,1035,223,248]" pageId="4" pageNumber="553">12.</emphasis>
|
||
Intestine begins in 14/15 or 15/16. Intestinal typhlosole starting in
|
||
<emphasis id="47C2EA8D9A46FF9D1C83AF7CFDCFFEA4" bold="true" box="[504,532,260,285]" pageId="4" pageNumber="553">15</emphasis>
|
||
or
|
||
<emphasis id="47C2EA8D9A46FF9D1F43AF7CFD8FFEA4" bold="true" box="[568,596,260,285]" pageId="4" pageNumber="553">16</emphasis>
|
||
, as a small fold increasing in size in
|
||
<emphasis id="47C2EA8D9A46FF9D1E95AF7CFBD1FEA4" bold="true" box="[1006,1034,260,285]" pageId="4" pageNumber="553">21</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D196CAF7CFBE8FEA4" bold="true" box="[1047,1075,260,285]" pageId="4" pageNumber="553">22</emphasis>
|
||
, lamina-shaped in
|
||
<emphasis id="47C2EA8D9A46FF9D187CAF7CFAF8FEA4" bold="true" box="[1287,1315,260,285]" pageId="4" pageNumber="553">23</emphasis>
|
||
or
|
||
<emphasis id="47C2EA8D9A46FF9D1833AF7CFABFFEA4" bold="true" box="[1352,1380,260,285]" pageId="4" pageNumber="553">24</emphasis>
|
||
with free edge divided into three deep ridges (i.e. typhlosole trifid), continuing with same shape until abrupt end in
|
||
<emphasis id="47C2EA8D9A46FF9D1810AF5FFA4EFEF9" bold="true" box="[1387,1429,295,320]" pageId="4" pageNumber="553">123</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1DECAF34FF1AFEDC" bold="true" box="[151,193,332,357]" pageId="4" pageNumber="553">124</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1DAAAF34FF20FEDC" bold="true" box="[209,251,332,357]" pageId="4" pageNumber="553">126</emphasis>
|
||
. In
|
||
<emphasis id="47C2EA8D9A46FF9D1C57AF34FEA8FEDC" bold="true" box="[300,371,332,357]" pageId="4" pageNumber="553">24–28</emphasis>
|
||
with up to 13 lateral folds, straight or slightly oblique in posterior direction. Smaller dorsolateral typhlosoles covering 6–7 segments in the region of
|
||
<emphasis id="47C2EA8D9A46FF9D1E5CAF17FCB6FE31" bold="true" box="[807,877,367,392]" pageId="4" pageNumber="553">21–28</emphasis>
|
||
, at both sides of main typhlosole. Intestinal caeca present as one or two paired dorsal pouches of the intestine in
|
||
<emphasis id="47C2EA8D9A46FF9D1E2AAFECFC4CFE14" bold="true" box="[849,919,404,429]" pageId="4" pageNumber="553">21–24</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A46FF9D1DBCAFCFFEAAFD49" blockId="4.[151,1436,151,1869]" pageId="4" pageNumber="553">
|
||
Single dorsal vessel visible throughout. Supra-esophageal vessel visible in
|
||
<emphasis id="47C2EA8D9A46FF9D1961AFCFFB89FE69" bold="true" box="[1050,1106,439,464]" pageId="4" pageNumber="553">8–12</emphasis>
|
||
. Lateral hearts in
|
||
<emphasis id="47C2EA8D9A46FF9D1858AFCFFAEAFE69" bold="true" box="[1315,1329,439,464]" pageId="4" pageNumber="553">7</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D183BAFCFFA95FE69" bold="true" box="[1344,1358,439,464]" pageId="4" pageNumber="553">8</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1826AFCFFAB0FE69" bold="true" box="[1373,1387,439,464]" pageId="4" pageNumber="553">9</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1DECAFA4FF68FE4C" bold="true" box="[151,179,476,501]" pageId="4" pageNumber="553">10</emphasis>
|
||
; latero-esophageal hearts in
|
||
<emphasis id="47C2EA8D9A46FF9D1C82AFA4FDCFFE4C" bold="true" box="[505,532,476,501]" pageId="4" pageNumber="553">11</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1F37AFA4FDB3FE4C" bold="true" box="[588,616,476,501]" pageId="4" pageNumber="553">12</emphasis>
|
||
. Ventral vessel present. Extra parietal ventral vessels in each side of
|
||
<emphasis id="47C2EA8D9A46FF9D1802AFA4FA4EFE4C" bold="true" box="[1401,1429,476,501]" pageId="4" pageNumber="553">13</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1DECAF87FF68FDA1" bold="true" box="[151,179,511,536]" pageId="4" pageNumber="553">14</emphasis>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1DBAAF87FF06FDA1" bold="true" box="[193,221,511,536]" pageId="4" pageNumber="553">15</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1C6FAF87FEEBFDA1" bold="true" box="[276,304,511,536]" pageId="4" pageNumber="553">16</emphasis>
|
||
, running outside male gonoduct joined in
|
||
<emphasis id="47C2EA8D9A46FF9D1E70AF87FCFCFDA1" bold="true" box="[779,807,511,536]" pageId="4" pageNumber="553">13</emphasis>
|
||
to paired infra-esophageal vessels, which run anteriorly until
|
||
<emphasis id="47C2EA8D9A46FF9D1DAAAC5CFF04FD84" bold="true" box="[209,223,548,573]" pageId="4" pageNumber="553">7</emphasis>
|
||
; vessels in
|
||
<emphasis id="47C2EA8D9A46FF9D1C19AC5CFEABFD84" bold="true" box="[354,368,548,573]" pageId="4" pageNumber="553">7</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1CDCAC5CFE6EFD84" bold="true" box="[423,437,548,573]" pageId="4" pageNumber="553">8</emphasis>
|
||
joined to lateral commissures before anastomosing with anterior septum and fusing with the ventral vessel. Paired clusters of tufted micronephridia along a longitudinal row in
|
||
<emphasis id="47C2EA8D9A46FF9D190FAC3FFB45FDD9" bold="true" box="[1140,1182,583,608]" pageId="4" pageNumber="553">2–4</emphasis>
|
||
; septal meronephridia observed in the anterior face of septa
|
||
<date id="0108105F9A46FF9D1F3DAC14FD63FD3D" box="[582,696,620,644]" pageId="4" pageNumber="553" value="2013-08-07" valueMax="2013-12-07" valueMin="2013-08-07">7/8–12/13</date>
|
||
(four, two on each side). Parietal, closed meronephridia from
|
||
<emphasis id="47C2EA8D9A46FF9D18FBAC14FA47FD3C" bold="true" box="[1408,1436,620,645]" pageId="4" pageNumber="553">14</emphasis>
|
||
backwards; in some individuals four on each side (eight per segment) from
|
||
<emphasis id="47C2EA8D9A46FF9D1E98ACF7FBF2FD11" bold="true" box="[995,1065,655,680]" pageId="4" pageNumber="553">28–39</emphasis>
|
||
; from
|
||
<emphasis id="47C2EA8D9A46FF9D190EACF7FB4AFD11" bold="true" box="[1141,1169,655,680]" pageId="4" pageNumber="553">40</emphasis>
|
||
on only three nephridia observed on each side. Median ventral nephridium of last segments larger and with a small nephrostome into the preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A46FF9D1DBCAC84FC5FFB26" blockId="4.[151,1436,151,1869]" pageId="4" pageNumber="553">
|
||
Holandric. Testes of
|
||
<emphasis id="47C2EA8D9A46FF9D1CCDAC84FE09FCAC" bold="true" box="[438,466,764,789]" pageId="4" pageNumber="553">10</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1F76AC84FDFDFCAC" bold="true" box="[525,550,764,789]" pageId="4" pageNumber="553">11</emphasis>
|
||
bushy and very large, joined to the male funnels by abundant coagulum and located mid-ventrally on the anterior septum. Male funnels iridescent and plicated, placed at both sides of the midventral line of the posterior septum; funnels of
|
||
<emphasis id="47C2EA8D9A46FF9D1FC9AD3CFD10FCE4" bold="true" box="[690,715,836,861]" pageId="4" pageNumber="553">11</emphasis>
|
||
larger than those of
|
||
<emphasis id="47C2EA8D9A46FF9D1EC1AD3CFC0DFCE4" bold="true" box="[954,982,836,861]" pageId="4" pageNumber="553">10</emphasis>
|
||
. Male gonoducts double, running along body wall in
|
||
<emphasis id="47C2EA8D9A46FF9D1C57AD10FE8BFCC6" box="[300,336,872,895]" italics="true" pageId="4" pageNumber="553">BC</emphasis>
|
||
of
|
||
<emphasis id="47C2EA8D9A46FF9D1C0DAD1FFE67FC39" bold="true" box="[374,444,871,896]" pageId="4" pageNumber="553">13–17</emphasis>
|
||
, muscular in region
|
||
<emphasis id="47C2EA8D9A46FF9D1FDEAD1FFD2AFC39" bold="true" box="[677,753,871,896]" pageId="4" pageNumber="553">15–16,</emphasis>
|
||
entering body wall in 17/18 just below the muscular duct of the prostate, towards mid-ventral line. Two pairs of seminal vesicles in
|
||
<emphasis id="47C2EA8D9A46FF9D1EBDADF4FC0FFC1C" bold="true" box="[966,980,908,933]" pageId="4" pageNumber="553">9</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1976ADF4FBF2FC1C" bold="true" box="[1037,1065,908,933]" pageId="4" pageNumber="553">12</emphasis>
|
||
, respectively fixed to septa
|
||
<date id="0108105F9A46FF9D1811ADF4FED3FC71" pageId="4" pageNumber="553" value="2012-10-09" valueMax="2012-11-09" valueMin="2012-10-09">9/10 and 11/12</date>
|
||
; the anterior pair slightly smaller and in some individuals flat; the posterior pair larger and acinous. Two pairs of tubular prostates of similar size in
|
||
<emphasis id="47C2EA8D9A46FF9D1F0FADACFD4BFC54" bold="true" box="[628,656,980,1005]" pageId="4" pageNumber="553">17</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1FBDADACFD39FC54" bold="true" box="[710,738,980,1005]" pageId="4" pageNumber="553">19</emphasis>
|
||
, strongly coiled and fixed to intestine and septa by connective tissue, extending 1–4 segments backwards; muscular duct narrower and shorter (1/4) than glandular part. Penial setae
|
||
<emphasis id="47C2EA8D9A46FF9D1DAEAA65FF38FB8D" box="[213,227,1053,1076]" italics="true" pageId="4" pageNumber="553">a</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1C62AA64FEFCFB8C" box="[281,295,1052,1077]" italics="true" pageId="4" pageNumber="553">b</emphasis>
|
||
of both
|
||
<emphasis id="47C2EA8D9A46FF9D1CFFAA64FE7BFB8C" bold="true" box="[388,416,1052,1077]" pageId="4" pageNumber="553">17</emphasis>
|
||
and
|
||
<emphasis id="47C2EA8D9A46FF9D1CADAA64FE29FB8C" bold="true" box="[470,498,1052,1077]" pageId="4" pageNumber="553">19</emphasis>
|
||
, in separate follicles that join to form an ovoidal muscle pouch; these pouches fixed by muscular stripes to lateral walls (one fiber) and to the floor (one or two fibers per pouch). Further retractor muscles in 16/17 and 18/19, extending from mid-ventral line to dorso-lateral walls; mid-ventral floor in segments
|
||
<emphasis id="47C2EA8D9A46FF9D1DECAAFFFF06FB19" bold="true" box="[151,221,1159,1184]" pageId="4" pageNumber="553">16–18</emphasis>
|
||
thicker. In each follicle, one undeveloped extra seta present.
|
||
</paragraph>
|
||
<paragraph id="7509369F9A46FF9D1DBCAAD4FD2DFA24" blockId="4.[151,1436,151,1869]" pageId="4" pageNumber="553">
|
||
Ovaries one pair, large, on the floor of
|
||
<emphasis id="47C2EA8D9A46FF9D1FEAAAD4FD76FB7C" bold="true" box="[657,685,1196,1221]" pageId="4" pageNumber="553">13</emphasis>
|
||
, with bushy coral shape and numerous eggs not in rows; female funnels one pair in
|
||
<emphasis id="47C2EA8D9A46FF9D1C15AAB7FE51FB51" bold="true" box="[366,394,1231,1256]" pageId="4" pageNumber="553">13</emphasis>
|
||
, at both sides of mid-ventral line. Two pairs of similar-sized spermathecae discharging in 7/8 and 8/9, ampulla and duct in the respective posterior segment, diverticulum in the anterior one; diverticulum sessile, discoidal, tightened to the parietes and with the overall appearance of a cabbage; duct wider and shorter than the elongated ampulla, which narrows in the middle part (
|
||
<figureCitation id="ED8D2A1A9A46FF9D1EDFAB44FC37FAED" box="[932,1004,1340,1365]" captionStart="FIGURE 1" captionStartId="2.[151,255,1443,1466]" captionTargetBox="[403,1182,200,1415]" captionTargetId="figure@2.[403,1184,195,1418]" captionTargetPageId="2" captionText="FIGURE 1. Ramiellona tojolabala sp. nov. A. External ventral view of anterior region. B. Spermatheca from segment 9, dorsal view. C. Transverse section (hand made dissection) from segment 9, showing the esophagus and dorsal stalked calciferous glands. D = dorsal, V = Ventral. Holotype IEOL 4238. Scale 1 mm." httpUri="https://zenodo.org/record/224300/files/figure.png" pageId="4" pageNumber="553">Fig. 1</figureCitation>
|
||
B). Length of spermathecae
|
||
<quantity id="B24E9B7A9A46FF9D1837AB44FA43FAED" box="[1356,1432,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="553" unit="mm" value="2.0">2 mm</quantity>
|
||
. Spermathecal setae embedded in ovoidal pouches of muscular tissue fixed to lateral parietes by muscular stripes. Two follicles within pouches, no extra setae observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DAC65149A46FF9D1DBCABDFFC7FFA5C" pageId="4" pageNumber="553" type="etymology">
|
||
<paragraph id="7509369F9A46FF9D1DBCABDFFC7FFA5C" blockId="4.[151,1436,151,1869]" pageId="4" pageNumber="553">
|
||
<emphasis id="47C2EA8D9A46FF9D1DBCABDFFE95FA79" bold="true" box="[199,334,1447,1472]" pageId="4" pageNumber="553">Etymology.</emphasis>
|
||
The name of the species is dedicated to the Tojolabal people who inhabit the region of Montebello lakes and other nearby zones, in the highlands of the state of Chiapas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3DAC65149A46FF9D1DBCAB97FE6AF8F5" pageId="4" pageNumber="553" type="discussion">
|
||
<paragraph id="7509369F9A46FF9D1DBCAB97FE6AF8F5" blockId="4.[151,1436,151,1869]" pageId="4" pageNumber="553">
|
||
<emphasis id="47C2EA8D9A46FF9D1DBCAB97FEE0F9B1" bold="true" box="[199,315,1519,1544]" pageId="4" pageNumber="553">Remarks.</emphasis>
|
||
<taxonomicName id="B2B64D1C9A46FF9D1C3FAB89FE02F9B1" box="[324,473,1519,1544]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="tojolabala">
|
||
<emphasis id="47C2EA8D9A46FF9D1C3FAB89FE02F9B1" box="[324,473,1519,1544]" italics="true" pageId="4" pageNumber="553">R. tojolabala</emphasis>
|
||
</taxonomicName>
|
||
belongs to the group of species with last hearts in
|
||
<emphasis id="47C2EA8D9A46FF9D1957AB97FB93F9B1" bold="true" box="[1068,1096,1519,1544]" pageId="4" pageNumber="553">12</emphasis>
|
||
and quincuaxial setae. Being holandric it is separated from the metandric
|
||
<emphasis id="47C2EA8D9A46FF9D1FDDA86DFF31F9E9" italics="true" pageId="4" pageNumber="553">
|
||
<taxonomicName id="B2B64D1C9A46FF9D1FDDA86DFC84F994" box="[678,863,1556,1581]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="guatemalana">R. guatemalana</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2B64D1C9A46FF9D1E15A86DFBDBF994" box="[878,1024,1556,1581]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="balantina">R. balantina</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2B64D1C9A46FF9D196BA86DFB1DF995" box="[1040,1222,1557,1580]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="tecumumami">R. tecumumami</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2B64D1C9A46FF9D19AEA86DFAAFF995" box="[1237,1396,1557,1580]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="americana">R. americana</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2B64D1C9A46FF9D18FFA86DFF31F9E9" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="lavellei">R. lavellei</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="B2B64D1C9A46FF9D1C5BA841FE77F9E9" box="[288,428,1591,1616]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="vulcanica">
|
||
<emphasis id="47C2EA8D9A46FF9D1C5BA841FE77F9E9" box="[288,428,1591,1616]" italics="true" pageId="4" pageNumber="553">R. vulcanica</emphasis>
|
||
</taxonomicName>
|
||
; by having two pairs of seminal vesicles it is separated from
|
||
<taxonomicName id="B2B64D1C9A46FF9D1926A841FB6CF9E9" box="[1117,1207,1593,1616]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="irpex">
|
||
<emphasis id="47C2EA8D9A46FF9D1926A841FB6CF9E9" box="[1117,1207,1593,1616]" italics="true" pageId="4" pageNumber="553">R. irpex</emphasis>
|
||
</taxonomicName>
|
||
(one pair), although both species share the dorsal calciferous glands in segments
|
||
<emphasis id="47C2EA8D9A46FF9D1E3AA824FCADF9CC" bold="true" box="[833,886,1628,1653]" pageId="4" pageNumber="553">7–11</emphasis>
|
||
. The quincuaxial arrangement of setae separates it from the other holandric species of the genus like
|
||
<taxonomicName id="B2B64D1C9A46FF9D1F8AA8F9FC49F921" box="[753,914,1663,1688]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="sauerlandti">
|
||
<emphasis id="47C2EA8D9A46FF9D1F8AA8F9FC49F921" box="[753,914,1663,1688]" italics="true" pageId="4" pageNumber="553">R. sauerlandti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="B2B64D1C9A46FF9D1ED9A8F9FB97F921" box="[930,1100,1663,1688]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="stadelmanni">
|
||
<emphasis id="47C2EA8D9A46FF9D1ED9A8F9FB97F921" box="[930,1100,1663,1688]" italics="true" pageId="4" pageNumber="553">R. stadelmanni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="47C2EA8D9A46FF9D1927A8F9FA92F921" box="[1116,1353,1665,1688]" italics="true" pageId="4" pageNumber="553">
|
||
<taxonomicName id="B2B64D1C9A46FF9D1927A8F9FB01F921" box="[1116,1242,1665,1688]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="strigosa">R. strigosa</taxonomicName>
|
||
, R,
|
||
<taxonomicName id="B2B64D1C9A46FF9D1873A8F9FA92F921" box="[1288,1353,1665,1688]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="eiseni">eiseni</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="B2B64D1C9A46FF9D18FFA8F9FE9AF904" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="microscolecina">
|
||
<emphasis id="47C2EA8D9A46FF9D18FFA8F9FE9AF904" italics="true" pageId="4" pageNumber="553">R. microscolecina</emphasis>
|
||
</taxonomicName>
|
||
, sharing with the last three species the shape of spermathecae. From the quincuaxial
|
||
<taxonomicName id="B2B64D1C9A46FF9D1987A8DDFAB7F904" box="[1276,1388,1700,1725]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="species" species="lasiura">
|
||
<emphasis id="47C2EA8D9A46FF9D1987A8DDFAB7F904" box="[1276,1388,1700,1725]" italics="true" pageId="4" pageNumber="553">R. lasiura</emphasis>
|
||
</taxonomicName>
|
||
it is also separated because, after the revision of one
|
||
<typeStatus id="AA0D883D9A46FF9D1FA4A8B0FCE4F959" box="[735,831,1736,1760]" pageId="4" pageNumber="553" type="paratype">paratype</typeStatus>
|
||
(see further on), this last species turned out to be metandric as was assumed by
|
||
<bibRefCitation id="11274B6E9A46FF9D1C8AA894FD5CF8BC" author="Gates" box="[497,647,1772,1797]" pageId="4" pageNumber="553" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article" year="1962">Gates (1962)</bibRefCitation>
|
||
. This species has previously been referred to as "
|
||
<taxonomicName id="B2B64D1C9A46FF9D19C7A894FAE6F8BC" box="[1212,1341,1772,1797]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="genus" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A46FF9D19C7A894FAE6F8BC" box="[1212,1341,1772,1797]" italics="true" pageId="4" pageNumber="553">Ramiellona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A46FF9D183EA895FA47F8BD" box="[1349,1436,1773,1796]" pageId="4" pageNumber="553" rank="species">
|
||
sp.
|
||
<date id="0108105F9A46FF9D1816A895FA47F8BD" box="[1389,1436,1773,1796]" pageId="4" pageNumber="553" value="1928-11">nov.</date>
|
||
</taxonomicNameLabel>
|
||
28" (
|
||
<bibRefCitation id="11274B6E9A46FF9D1DA9A968FEAEF891" author="Fragoso" box="[210,373,1808,1832]" pageId="4" pageNumber="553" refString="Fragoso, C. (1993) Les Peuplements de Vers de Terre dans L'Est et Sud-Est du Mexique. These Universite Paris 6, Paris, 228 pp." type="book" year="1993">Fragoso 1993</bibRefCitation>
|
||
), "
|
||
<taxonomicName id="B2B64D1C9A46FF9D1CE1A977FDC0F891" box="[410,539,1807,1832]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="genus" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A46FF9D1CE1A977FDC0F891" box="[410,539,1807,1832]" italics="true" pageId="4" pageNumber="553">Ramiellona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A46FF9D1F5CA968FD59F89E" box="[551,642,1808,1831]" pageId="4" pageNumber="553" rank="species">
|
||
sp.
|
||
<date id="0108105F9A46FF9D1F28A968FD59F89E" box="[595,642,1808,1831]" pageId="4" pageNumber="553" value="2013-11">nov.</date>
|
||
</taxonomicNameLabel>
|
||
13" (
|
||
<bibRefCitation id="11274B6E9A46FF9D1FB2A968FCB6F891" author="Fragoso" box="[713,877,1808,1832]" pageId="4" pageNumber="553" refString="Fragoso, C. (2001) Las lombrices de Tierra de Mexico (Annelida, Oligochaeta): Diversidad, Ecologia y Manejo. Acta Zoologica Mexicana (n. s.), Numero especial 1, 131 - 171." type="journal article" year="2001">Fragoso 2001</bibRefCitation>
|
||
), and "
|
||
<taxonomicName id="B2B64D1C9A46FF9D1EBEA977FB9DF891" box="[965,1094,1807,1832]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="553" phylum="Annelida" rank="genus" status="sp. nov.">
|
||
<emphasis id="47C2EA8D9A46FF9D1EBEA977FB9DF891" box="[965,1094,1807,1832]" italics="true" pageId="4" pageNumber="553">Ramiellona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5CF157F69A46FF9D1929A968FB75F89E" box="[1106,1198,1808,1831]" pageId="4" pageNumber="553" rank="species">
|
||
sp.
|
||
<date id="0108105F9A46FF9D1905A968FB75F89E" box="[1150,1198,1808,1831]" pageId="4" pageNumber="553" value="1914-11">nov.</date>
|
||
</taxonomicNameLabel>
|
||
14" (Fragoso 2007;
|
||
<bibRefCitation id="11274B6E9A46FF9D1DECA94DFE7EF8F5" author="Fragoso" box="[151,421,1844,1869]" pageId="4" pageNumber="553" refString="Fragoso, C. & Brown, G. (2007) Ecologia y Taxonomia de las lombrices de tierra en Latinoamerica: El primer Encuentro Latino-Americano de Ecologia y Taxonomia de Oligoquetos (ELAETAO 1). In: Brown, G. G. & Fragoso, C. (Eds.), Minhocas na America Latina: Biodiversidade e Ecologia. EMBRAPA Soja. Londrina, pp. 33 - 75." type="book chapter" year="2007">Fragoso & Brown 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |