164 lines
18 KiB
XML
164 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4327827" ID-GBIF-Dataset="6e84f4d6-64d0-408b-889b-e270ab6be3a7" ID-GBIF-Taxon="164845022" ID-Zenodo-Dep="4327827" approvalRequired="9" approvalRequired_for_textStreams="8" approvalRequired_for_treatments="1" checkinTime="1593591561348" checkinUser="plazi" docAuthor="Kamura, Takahide" docDate="2020" docId="F224878C3B00FFA8FBDAFFFDFCA80FB5" docLanguage="en" docName="actaArcachnologica.69.1.1-9.pdf" docOrigin="Acta Arachnologica 69 (1)" docStyle="DocumentStyle{}" docTitle="Aponetius Kamura 2020, n. gen." docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="7" masterDocId="0E1DFFF43B05FFAEFFC7FA54FFAA0B53" masterDocTitle="A new genus of the subfamily Zelotinae (Araneae: Gnaphosidae), with descrip- tions of two new species from Amami-Ōshima Island, Southwest Japan" masterLastPageNumber="9" masterPageNumber="1" pageId="5" pageNumber="6" updateTime="1634701899359" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus of the subfamily Zelotinae (Araneae: Gnaphosidae), with descrip- tions of two new species from Amami-Ōshima Island, Southwest Japan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kamura, Takahide</mods:namePart>
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<mods:affiliation>1 – 1 – 527, Chiyoda-cho, Takatsuki-shi, Osaka 569 – 0087 Japan</mods:affiliation>
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<mods:nameIdentifier type="email">kamura@haruka.otemon.ac.jp</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Arachnologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-06-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>9</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3925102</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6e84f4d6-64d0-408b-889b-e270ab6be3a7</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3925102</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4327827" ID-GBIF-Taxon="164845022" ID-Zenodo-Dep="4327827" LSID="urn:lsid:plazi:treatment:F224878C3B00FFA8FBDAFFFDFCA80FB5" httpUri="http://treatment.plazi.org/id/F224878C3B00FFA8FBDAFFFDFCA80FB5" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection box="[1053,1242,1449,1473]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph blockId="5.[909,1386,1449,1505]" box="[1053,1242,1449,1473]" pageId="5" pageNumber="6">
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<heading bold="true" box="[1053,1242,1449,1473]" fontSize="10" level="3" pageId="5" pageNumber="6" reason="6">
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<taxonomicName authority="Kamura, 2020" authorityName="Kamura" authorityYear="2020" box="[1053,1163,1449,1473]" class="Arachnida" family="Gnaphosidae" genus="Aponetius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="n. gen.">
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<emphasis bold="true" box="[1053,1163,1449,1473]" italics="true" pageId="5" pageNumber="6">Aponetius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1170,1242,1452,1472]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[909,1386,1483,1505]" pageId="5" pageNumber="6" type="vernacular_names">
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<paragraph blockId="5.[909,1386,1449,1505]" box="[909,1386,1483,1505]" pageId="5" pageNumber="6">[Japanese name: Tsuzura-kemurigumo-zoku]</paragraph>
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</subSubSection>
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<subSubSection box="[839,1311,1545,1570]" pageId="5" pageNumber="6" type="type_taxon">
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<paragraph blockId="5.[813,1456,1545,1954]" box="[839,1311,1545,1570]" pageId="5" pageNumber="6">
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<emphasis bold="true" box="[839,986,1545,1570]" pageId="5" pageNumber="6">
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<typeStatus box="[839,894,1546,1570]" pageId="5" pageNumber="6">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName authority="Kamura 1999" authorityName="Kamura" authorityYear="1999" box="[992,1308,1545,1569]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gladius">
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<emphasis box="[992,1153,1545,1569]" italics="true" pageId="5" pageNumber="6">Zelotes gladius</emphasis>
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<bibRefCitation author="Kamura, T." box="[1160,1308,1547,1568]" journalOrPublisher="Acta Arachnol." pageId="5" pageNumber="6" pagination="79 - 91" part="48" refId="ref4446" refString="Kamura, T. 1999. Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan. Acta Arachnol., 48: 79 - 91." title="Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan" type="journal article" year="1999">Kamura 1999</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="diagnosis">
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<paragraph blockId="5.[813,1456,1545,1954]" pageId="5" pageNumber="6">
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<emphasis bold="true" box="[839,965,1577,1601]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
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The new genus is distinguished from the known zelotine genera except
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<taxonomicName authorityName="Gistel" authorityYear="1848" box="[1141,1217,1609,1633]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis box="[1141,1217,1609,1633]" italics="true" pageId="5" pageNumber="6">Zelotes</emphasis>
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</taxonomicName>
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by having an intercalary sclerite in the male palp, and from
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<taxonomicName authorityName="Gistel" authorityYear="1848" box="[1243,1319,1642,1666]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis box="[1243,1319,1642,1666]" italics="true" pageId="5" pageNumber="6">Zelotes</emphasis>
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</taxonomicName>
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by the male palp with an embolar radix reduced, a long embolus arising ventrally and a curved ridge on the retrolateral side of the cymbium, and the female genitalia with internal ducts complexly winding.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph blockId="5.[813,1456,1545,1954]" lastBlockId="6.[132,776,172,1255]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<emphasis bold="true" box="[839,977,1802,1826]" pageId="5" pageNumber="6">Description.</emphasis>
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Anterior eye row weakly recurved and posterior eye row straight or slightly procurved as seen from dorsal side,
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<collectionCode box="[942,996,1870,1890]" pageId="5" pageNumber="6">PME</collectionCode>
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separated from each other by less than the eye size,
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<collectionCode box="[910,970,1901,1921]" pageId="5" pageNumber="6">MOA</collectionCode>
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slightly longer than wide, with anterior width slightly narrower than or almost as wide as the posterior. Chelicera with three or four relatively large teeth on promargin of fang furrow, with three or four relatively small teeth on retromargin (
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<figureCitation box="[301,394,236,258]" captionStart="Fig" captionStartId="2.[132,165,1511,1533]" captionTargetBox="[183,1383,1038,1487]" captionTargetId="figure@2.[132,1455,1017,1512]" captionTargetPageId="2" captionText="Fig. 3. Aponetius gladius (Kamura 1999) n. comb. A, left chelicera, ventral view; B, left female spinnerets, posteroventral view; C, left male spinnerets, posteroventral view. Abbreviations: ac, aciniform gland spigot; ALS, anterior lateral spinneret; cy, cylindrical gland spigot; ma am, major ampullate gland spigot; mi am, minor ampullate gland spigot; pi, piriform gland spigot; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3925108" httpUri="https://zenodo.org/record/3925108/files/figure.png" pageId="6" pageNumber="7">Fig. 3 A</figureCitation>
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). Thoracic groove longitudinal, distinct. Trochanters of legs without ventral notch. Metatarsi III and IV with preening comb on ventrodistal end. Metatarsi and tarsi of legs without scopula. Leg formula 4-1-2-3. Male abdomen with dorsal scutum variable from two- to threefifths of abdomen in length. Spinnerets: female (
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<figureCitation box="[662,762,397,419]" captionStart="Fig" captionStartId="2.[132,165,1511,1533]" captionTargetBox="[183,1383,1038,1487]" captionTargetId="figure@2.[132,1455,1017,1512]" captionTargetPageId="2" captionText="Fig. 3. Aponetius gladius (Kamura 1999) n. comb. A, left chelicera, ventral view; B, left female spinnerets, posteroventral view; C, left male spinnerets, posteroventral view. Abbreviations: ac, aciniform gland spigot; ALS, anterior lateral spinneret; cy, cylindrical gland spigot; ma am, major ampullate gland spigot; mi am, minor ampullate gland spigot; pi, piriform gland spigot; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3925108" httpUri="https://zenodo.org/record/3925108/files/figure.png" pageId="6" pageNumber="7">Fig. 3 B</figureCitation>
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):
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<collectionCode box="[132,182,430,450]" pageId="6" pageNumber="7">ALS</collectionCode>
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with five or six piriform gland spigots and two major ampullate gland spigots;
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<collectionCode box="[402,455,462,482]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="6" pageNumber="7">PMS</collectionCode>
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swollen with four cylindrical gland spigots, one minor ampullate gland spigot and about ten aciniform gland spigots;
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<collectionCode box="[440,486,527,547]" pageId="6" pageNumber="7">PLS</collectionCode>
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with two cylindrical gland spigots, one minor ampullate gland spigot and about ten aciniform gland spigots; male (
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<figureCitation box="[469,568,590,612]" captionStart="Fig" captionStartId="2.[132,165,1511,1533]" captionTargetBox="[183,1383,1038,1487]" captionTargetId="figure@2.[132,1455,1017,1512]" captionTargetPageId="2" captionText="Fig. 3. Aponetius gladius (Kamura 1999) n. comb. A, left chelicera, ventral view; B, left female spinnerets, posteroventral view; C, left male spinnerets, posteroventral view. Abbreviations: ac, aciniform gland spigot; ALS, anterior lateral spinneret; cy, cylindrical gland spigot; ma am, major ampullate gland spigot; mi am, minor ampullate gland spigot; pi, piriform gland spigot; PLS, posterior lateral spinneret; PMS, posterior median spinneret. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3925108" httpUri="https://zenodo.org/record/3925108/files/figure.png" pageId="6" pageNumber="7">Fig. 3 C</figureCitation>
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):
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<collectionCode box="[589,639,591,611]" pageId="6" pageNumber="7">ALS</collectionCode>
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with five or six piriform gland spigots and one or two major ampullate gland spigots;
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<collectionCode box="[288,341,655,675]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="6" pageNumber="7">PMS</collectionCode>
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and
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<collectionCode box="[393,439,655,675]" pageId="6" pageNumber="7">PLS</collectionCode>
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each with one minor ampullate gland spigot and about ten aciniform gland spigots. Male palp (
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<figureCitation box="[192,263,718,740]" captionStart="Fig" captionStartId="1.[132,165,1758,1780]" captionTargetBox="[165,1431,205,1729]" captionTargetId="figure@1.[142,1451,164,1752]" captionTargetPageId="1" captionText="Fig. 1. A–D, Zelotes kimwha Paik 1986; E–I, Aponetius gladius (Kamura 1999) n. comb. A, E, left male palp, ventral view; B, F, bulb of left male palp, apical view; C, same, dorsoretrolateral view; D, I, female internal genitalia, dorsal view; G, bulb of left male palp, dorsoapical view; H, same, prolateral view. Abbreviations: C, conductor; E, embolus; EB, embolar base; EBP, embolar base projection; ER, embolar radix; IS, intercalary sclerite; MA, median apophysis; T, tegulum; TA, terminal apophysis; TB, tegular base. Scales = 0.2 mm. A, after Kamura (2000); D, after Kamura (2003); E, I, after Kamura (1999)." figureDoi="http://doi.org/10.5281/zenodo.3925104" httpUri="https://zenodo.org/record/3925104/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
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E–H): intercalary sclerite, terminal apophysis and median apophysis present; embolus long, arising ventrally, surrounding bulb dorsally and reappearing retrolaterally; embolar radix reduced, situated in central part of bulb; conductor reduced, situated from dorsally to prolaterally to embolar base; proximal part of embolus pushing aside intercalary sclerite, and often intercalary sclerite and tegulum extremely pressed retrolaterally; cymbium with distinct curved ridge on retrolateral side; retrolateral tibial apophysis present. Female internal genitalia with copulatory ducts complexly winding (
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<figureCitation box="[348,434,1039,1061]" captionStart="Fig" captionStartId="1.[132,165,1758,1780]" captionTargetBox="[165,1431,205,1729]" captionTargetId="figure@1.[142,1451,164,1752]" captionTargetPageId="1" captionText="Fig. 1. A–D, Zelotes kimwha Paik 1986; E–I, Aponetius gladius (Kamura 1999) n. comb. A, E, left male palp, ventral view; B, F, bulb of left male palp, apical view; C, same, dorsoretrolateral view; D, I, female internal genitalia, dorsal view; G, bulb of left male palp, dorsoapical view; H, same, prolateral view. Abbreviations: C, conductor; E, embolus; EB, embolar base; EBP, embolar base projection; ER, embolar radix; IS, intercalary sclerite; MA, median apophysis; T, tegulum; TA, terminal apophysis; TB, tegular base. Scales = 0.2 mm. A, after Kamura (2000); D, after Kamura (2003); E, I, after Kamura (1999)." figureDoi="http://doi.org/10.5281/zenodo.3925104" httpUri="https://zenodo.org/record/3925104/files/figure.png" pageId="6" pageNumber="7">Fig. 1 I</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="etymology">
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<paragraph blockId="6.[132,776,172,1255]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[159,286,1070,1094]" pageId="6" pageNumber="7">Etymology.</emphasis>
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Generic name is an arbitrary combination of letters and is masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="7" type="synonymic_list">
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<paragraph blockId="6.[132,776,172,1255]" pageId="6" pageNumber="7">
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<emphasis bold="true" box="[159,369,1134,1158]" pageId="6" pageNumber="7">Species included.</emphasis>
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Three species previously assigned to
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<taxonomicName authorityName="Gistel" authorityYear="1848" box="[164,246,1166,1190]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis box="[164,246,1166,1190]" italics="true" pageId="6" pageNumber="7">Zelotes</emphasis>
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</taxonomicName>
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:
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<taxonomicName authority="(Kamura 1999)" authorityName="Kamura" authorityYear="2020" baseAuthorityName="Kamura" baseAuthorityYear="1999" box="[264,654,1166,1190]" class="Arachnida" family="Gnaphosidae" genus="Aponetius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gladius" status="n. comb.">
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<emphasis box="[264,469,1166,1190]" italics="true" pageId="6" pageNumber="7">Aponetius gladius</emphasis>
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(
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<bibRefCitation author="Kamura, T." box="[484,649,1168,1189]" journalOrPublisher="Acta Arachnol." pageId="6" pageNumber="7" pagination="79 - 91" part="48" refId="ref4446" refString="Kamura, T. 1999. Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan. Acta Arachnol., 48: 79 - 91." title="Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan" type="journal article" year="1999">Kamura 1999</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[664,767,1167,1191]" pageId="6" pageNumber="7" rank="species">
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<emphasis bold="true" box="[664,761,1167,1191]" pageId="6" pageNumber="7">n. comb</emphasis>
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.
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</taxonomicNameLabel>
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,
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<taxonomicName authority="(Kamura 1999)" authorityName="Kamura" authorityYear="2020" baseAuthorityName="Kamura" baseAuthorityYear="1999" box="[132,472,1198,1222]" class="Arachnida" family="Gnaphosidae" genus="Aponetius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ryukyuensis" status="n. comb.">
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<emphasis box="[132,294,1198,1222]" italics="true" pageId="6" pageNumber="7">A. ryukyuensis</emphasis>
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(
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<bibRefCitation author="Kamura, T." box="[307,468,1200,1222]" journalOrPublisher="Acta Arachnol." pageId="6" pageNumber="7" pagination="79 - 91" part="48" refId="ref4446" refString="Kamura, T. 1999. Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan. Acta Arachnol., 48: 79 - 91." title="Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan" type="journal article" year="1999">Kamura 1999</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[482,583,1199,1223]" pageId="6" pageNumber="7" rank="species">
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<emphasis bold="true" box="[482,576,1199,1223]" pageId="6" pageNumber="7">n. comb</emphasis>
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.
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</taxonomicNameLabel>
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and
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<taxonomicName authority="(Kamura 1999)" authorityName="Kamura" authorityYear="2020" baseAuthorityName="Kamura" baseAuthorityYear="1999" class="Arachnida" family="Gnaphosidae" genus="Aponetius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flexuosus" status="n. comb.">
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<emphasis box="[641,775,1198,1222]" italics="true" pageId="6" pageNumber="7">A. flexuosus</emphasis>
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(
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<bibRefCitation author="Kamura, T." box="[136,290,1232,1254]" journalOrPublisher="Acta Arachnol." pageId="6" pageNumber="7" pagination="79 - 91" part="48" refId="ref4446" refString="Kamura, T. 1999. Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan. Acta Arachnol., 48: 79 - 91." title="Spiders of the genus Zelotes (Araneae: Gnaphosidae) from Ryukyu Islands, Southwest Japan" type="journal article" year="1999">Kamura 1999</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[301,397,1231,1255]" pageId="6" pageNumber="7" rank="species">
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<emphasis bold="true" box="[301,391,1231,1255]" pageId="6" pageNumber="7">n. comb</emphasis>
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.
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</taxonomicNameLabel>
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; two new species described below.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |