treatments-xml/data/A8/2B/87/A82B87C50E07F2324EB9FE7BA640D6B9.xml

<document id="B73227126CFB3897165A87499266F619" ID-DOI="10.1080/00222930110062345" ID-ISSN="1464-5262" ID-Zenodo-Dep="5301013" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630001574096" checkinUser="felipe" docAuthor="Balke, M., Hendrich, L., Mazzoldi, P. &amp; Biström, O." docDate="2002" docId="A82B87C50E07F2324EB9FE7BA640D6B9" docLanguage="en" docName="JNatHist.36.8.963-978.pdf" docOrigin="Journal of Natural History 36 (8)" docSource="http://www.tandfonline.com/doi/abs/10.1080/00222930110062345" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Borneodessus zetteli subsp. zetteli Balke &amp; Hendrich &amp; Mazzoldi &amp; Biström 2002, sp. nov." docType="treatment" docVersion="4" lastPageNumber="967" masterDocId="5412FFBD0E01F2344F18FF9FA244D331" masterDocTitle="Borneodessus zetteli, new genus and new species, notes on biogeography and a checklist of Dytiscidae from Borneo (Insecta: Coleoptera: Dytiscidae)" masterLastPageNumber="978" masterPageNumber="963" pageNumber="967" updateTime="1699135344134" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E68D7CC00F0225F12029BE6C94DC0050">Borneodessus zetteli, new genus and new species, notes on biogeography and a checklist of Dytiscidae from Borneo (Insecta: Coleoptera: Dytiscidae)</mods:title>
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<mods:name id="0CABEE32FCFDAAECA7037F2CA1A8D055" type="personal">
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<mods:namePart id="BD484869ADFBBDD7E9D4C8EB56387C01">Balke, M.</mods:namePart>
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<mods:namePart id="4D663D8966CB5168508082317792B1A1">Hendrich, L.</mods:namePart>
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<mods:namePart id="3A471ABDB1C8913C5BDA4D8EA04F0A9C">Mazzoldi, P.</mods:namePart>
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<mods:namePart id="FE040D08DD13E635477883A334DD253F">Biström, O.</mods:namePart>
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<mods:title id="B179D8157CB5888E817C9CC06E700AE4">Journal of Natural History</mods:title>
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<mods:date id="DA71574B988BA7805DE955081778C04A">2002</mods:date>
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<mods:number id="A1524776EFBB87F14642344BD7B86728">2002-06-30</mods:number>
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<mods:number id="7300CA757F464BFE58B8B639C66B3324">36</mods:number>
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<mods:number id="301268EA7A494489590A651DAEA0401B">8</mods:number>
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<treatment id="A82B87C50E07F2324EB9FE7BA640D6B9" ID-DOI="http://doi.org/10.5281/zenodo.5308481" ID-Zenodo-Dep="5308481" LSID="urn:lsid:plazi:treatment:A82B87C50E07F2324EB9FE7BA640D6B9" httpUri="http://treatment.plazi.org/id/A82B87C50E07F2324EB9FE7BA640D6B9" lastPageNumber="967" pageId="6" pageNumber="967">
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<paragraph id="203D36D30E07F2324EB9FE7BA174D2CD" blockId="6.[417,816,484,541]" box="[417,816,484,508]" pageId="6" pageNumber="967">
<heading id="7B7581BF0E07F2324EB9FE7BA174D2CD" box="[417,816,484,508]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="967" reason="3">
<taxonomicName id="E7824D500E07F2324EB9FE7BA089D2CD" ID-CoL="5FZ8Y" authority="Balke &amp; Hendrich &amp; Mazzoldi &amp; Biström, 2002" authorityName="Balke &amp; Hendrich &amp; Mazzoldi &amp; Biström" authorityYear="2002" box="[417,717,484,508]" class="Insecta" family="Dytiscidae" genus="Borneodessus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="967" phylum="Arthropoda" rank="subSpecies" species="zetteli" status="sp. nov." subSpecies="zetteli">
<emphasis id="12F6EAC10E07F2324EB9FE7BA089D2CD" bold="true" box="[417,717,484,508]" italics="true" pageId="6" pageNumber="967">Borneodessus zetteli zetteli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09C557BA0E07F2324DCEFE7AA174D2CD" box="[726,816,485,508]" pageId="6" pageNumber="967" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="203D36D30E07F2324D1BFD9BA08BD12D" blockId="6.[417,816,484,541]" box="[515,719,516,541]" pageId="6" pageNumber="967">(®gures 1, 4, 6, 7)</paragraph>
</subSubSection>
<subSubSection id="689865580E07F2324FABFDABA640D6B9" pageId="6" pageNumber="967" type="materials_examined">
<paragraph id="203D36D30E07F2324FABFDABA3F9D1BA" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFDABA312D17A" box="[179,342,563,587]" italics="true" pageId="6" pageNumber="967">Type material.</emphasis>
<materialsCitation id="90EA3C8E0E07F2324E46FDABA088D15A" collectingDate="1997-02-12" collectionCode="NMW" collectorName="H. Zettel" country="Malaysia" location="Sapat Kalisan" municipality="Danum Valley" pageId="6" pageNumber="967" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="FF3988710E07F2324E46FDABA395D17A" box="[350,465,563,587]" pageId="6" pageNumber="967" type="holotype">HOLOTYPE</typeStatus>
male. 
<collectingCountry id="589576430E07F2324D38FDACA0C3D17A" box="[544,647,563,587]" name="Malaysia" pageId="6" pageNumber="967">Malaysia</collectingCountry>
, 
<collectingRegion id="E246F8310E07F2324D8FFDACA098D17A" box="[663,732,563,587]" country="Malaysia" name="Sabah" pageId="6" pageNumber="967">Sabah</collectingRegion>
, 
<collectingMunicipality id="C059ACA90E07F2324DF4FDABA1D4D17A" box="[748,912,563,587]" pageId="6" pageNumber="967">Danum Valley</collectingMunicipality>
, 
<location id="255D60080E07F2324C87FDACA67BD17A" LSID="urn:lsid:plazi:treatment:A82B87C50E07F2324EB9FE7BA640D6B9:255D60080E07F2324C87FDACA67BD17A" box="[927,1087,563,587]" country="Malaysia" municipality="Danum Valley" name="Sapat Kalisan" pageId="6" pageNumber="967" stateProvince="Sabah">Sapat Kalisan</location>
, locality 15, 
<date id="543C10130E07F2324E0DFDCCA3A5D15A" box="[277,481,595,619]" pageId="6" pageNumber="967" value="1997-02-12">
<collectingDate id="4478E9FB0E07F2324E0DFDCCA3A5D15A" box="[277,481,595,619]" pageId="6" pageNumber="967" value="1997-02-12">12 February 1997</collectingDate>
</date>
, 
<collectorName id="8D7753050E07F2324EEBFDCBA018D15A" box="[499,604,595,619]" pageId="6" pageNumber="967">H. Zettel</collectorName>
(
<collectionCode id="4693AE160E07F2324D6AFDCBA0F8D15A" box="[626,700,596,619]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="6" pageNumber="967">NMW</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="90EA3C8E0E07F2324DC3FDCBA3FCD1BA" collectingDate="1997-02-12" collectionCode="NMW" collectorName="H. Zettel" country="Malaysia" location="Sapat Kalisan" municipality="Danum Valley" pageId="6" pageNumber="967" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="FF3988710E07F2324DC3FDCBA109D15A" box="[731,845,595,619]" pageId="6" pageNumber="967" type="paratype">PARATYPE</typeStatus>
: 
<specimenCount id="3684FD5A0E07F2324C42FDCCA1F8D15A" box="[858,956,595,619]" pageId="6" pageNumber="967" type="female">1 female</specimenCount>
, same label data as holotype (
<collectionCode id="4693AE160E07F2324E46FDEBA3ECD1BA" box="[350,424,628,651]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="6" pageNumber="967">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="203D36D30E07F2324FABFD0CA30ED1FA" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFD0CA2A6D19A" box="[179,226,659,683]" italics="true" pageId="6" pageNumber="967">Size</emphasis>
. Length of beetle 
<quantity id="E77A9B360E07F2324EDCFD0CA051D19A" box="[452,533,659,683]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="967" unit="mm" value="2.9">2.9 mm</quantity>
(male), 
<quantity id="E77A9B360E07F2324D65FD0CA08BD19A" box="[637,719,659,683]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="967" unit="mm" value="2.8">2.8 mm</quantity>
(female); width 
<quantity id="E77A9B360E07F2324C80FD0CA1AED19A" box="[920,1002,659,683]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="967" unit="mm" value="1.6">1.6 mm</quantity>
(male), 
<quantity id="E77A9B360E07F2324F94FD2CA299D1FA" box="[140,221,691,715]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="967" unit="mm" value="1.5">1.5 mm</quantity>
(female).
</paragraph>
<paragraph id="203D36D30E07F2324FABFD4BA0B8D03A" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFD4BA3ABD1DA" box="[179,495,723,747]" italics="true" pageId="6" pageNumber="967">Description (male). Colour.</emphasis>
Beetle almost completely ferruginous. Elytron with dark brown, barely contrasting maculations (®gure 1).
</paragraph>
<paragraph id="203D36D30E07F2324FABFC8CA0A5D71B" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFC8CA35FD01A" box="[179,283,787,811]" italics="true" pageId="6" pageNumber="967">Sculpture</emphasis>
. Beetle dorsally and ventrally shiny. Head smooth medially, with small punctures laterally along eyes; with microreticulation and some minute punctures posterior of an imaginary line connecting hind margins of eyes. Pronotum with dense punctation, punctures larger than those on head, regularly spaced, separated by about twice their diameter; no microreticulation present; basal stria sinuate and faint, about one-third of pronotal length. Elytron with dense punctation as on pronotum, not microreticulate; no serial rows of larger punctures present. Metasternum and metacoxal plates microreticulate and with larger punctures; sternites microreticulate with few smaller punctures.
</paragraph>
<paragraph id="203D36D30E07F2324FABFBADA307D7F8" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFBADA30ED77B" box="[179,330,1074,1098]" italics="true" pageId="6" pageNumber="967">Median lobe.</emphasis>
Median lobe of aedeagus simple, symmetrical in ventral view. Rather straight and only slightly bent distally in lateral view; in ventral view broad, more or less parallel-sided over most of its length, narrowing apically, apex pointed. Paramere two-segmented, apical segment almost as long as basal segment; tip twisted (®gures 4, 6, 7).
</paragraph>
<paragraph id="203D36D30E07F2324FABFB4EA17FD638" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFB4EA345D7D8" box="[179,257,1233,1257]" italics="true" pageId="6" pageNumber="967">Female</emphasis>
. As in male except: beetle dull because of distinct microreticulation of small cells present all over the head, pronotum and elytron.
</paragraph>
<paragraph id="203D36D30E07F2324FABFA8EA055D679" blockId="6.[140,1095,563,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFA8EA36ED618" box="[179,298,1297,1321]" italics="true" pageId="6" pageNumber="967">Etymology</emphasis>
. Named for our colleague and friend, the collector of this species, Herbert Zettel (
<collectingRegion id="E246F8310E07F2324E5CFAAEA3D0D678" box="[324,404,1329,1353]" country="Austria" name="Wien" pageId="6" pageNumber="967">Vienna</collectingRegion>
Museum).
</paragraph>
<paragraph id="203D36D30E07F2324FABFACEA1F0D658" blockId="6.[140,1095,563,1417]" box="[179,948,1361,1385]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFACEA373D658" box="[179,311,1361,1385]" italics="true" pageId="6" pageNumber="967">Distribution</emphasis>
. East 
<collectingCountry id="589576430E07F2324E9BFACEA3AED658" box="[387,490,1361,1385]" name="Malaysia" pageId="6" pageNumber="967">Malaysia</collectingCountry>
, northern 
<collectingRegion id="E246F8310E07F2324D70FACEA0EAD658" box="[616,686,1361,1385]" country="Malaysia" name="Sabah" pageId="6" pageNumber="967">Sabah</collectingRegion>
(Borneo), see ®gure 8.
</paragraph>
<paragraph id="203D36D30E07F2324FABFAEEA640D6B9" blockId="6.[140,1095,563,1417]" box="[179,1028,1393,1417]" pageId="6" pageNumber="967">
<emphasis id="12F6EAC10E07F2324FABFAEEA329D6B8" box="[179,365,1393,1417]" italics="true" pageId="6" pageNumber="967">Collecting notes.</emphasis>
In shallow, small isolated puddles, at the edge of a river.
</paragraph>
</subSubSection>
</treatment>
</document>