245 lines
31 KiB
XML
245 lines
31 KiB
XML
<document id="05A4364379FE15F3234B73469FB9A6D1" ID-DOI="10.26107/RBZ-2021-0005" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351678" ID-ZooBank="D63943BE-00B0-409E-BCE4-96942D33114E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630206537254" checkinUser="diego" docAuthor="Tran, A. D. & Zettel, H." docDate="2021" docId="610CF11BE005FFDBFEEE24D6FEEBFBB6" docLanguage="en" docName="RBZ-2021-0005.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Ranatra bendanilloi Tran & Zettel 2021, new species" docType="treatment" docUuid="BD4AF315-A356-4F14-B4CE-19812342AD3D" docUuidSource="ZooBank" docVersion="4" lastPageNumber="57" masterDocId="9D358963E00FFFD7FFE32768FFF2FFE3" masterDocTitle="Taxonomic review of the Ranatra gracilis group sensu Lansbury, 1972 (Nepomorpha: Nepidae), with descriptions of four new species" masterLastPageNumber="70" masterPageNumber="45" pageNumber="55" updateTime="1699185027225" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="44C8CFA8919FCB84F620E2823ED7D4A4">
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<mods:title id="27A1C2E402D8F1081D567182B952DC6E">Taxonomic review of the Ranatra gracilis group sensu Lansbury, 1972 (Nepomorpha: Nepidae), with descriptions of four new species</mods:title>
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<mods:name id="2E87A0F0509C2E1D3227B3E5B954CC74" type="personal">
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<mods:roleTerm id="73DA5C38B960F7D2E1AAF8AC88F0AB47">Author</mods:roleTerm>
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<mods:namePart id="B013B7690D4D52AAE24E8D0A5BE8F5AD">Tran, A. D.</mods:namePart>
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<mods:roleTerm id="ED0B6E21F2243AB3476385CDB3938DB6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3DBADE51DE26522EFC41C2D8D78D5979">Zettel, H.</mods:namePart>
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<mods:typeOfResource id="C434F5F4BAC4DA15F4EA31DE1897DB10">text</mods:typeOfResource>
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<mods:title id="66A75DFCD2990D5FD049D12D383ECC2E">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="67BBF948BAE6A2834C931D565E7E7839">
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<mods:date id="4495792581E4359A58B76A01DC8BB6BF">2021</mods:date>
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<mods:number id="F5374B1F26D72D4271233F1BAB0F3E7A">2021-02-25</mods:number>
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</mods:detail>
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<mods:detail id="D142F1192113DC0AAD82AC8670952F3C" type="volume">
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<mods:number id="E9F60BCC8CA50D1C9CA5D4A6AD95220A">69</mods:number>
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<mods:start id="2A085C34C995F3D3C6CB951CC050B53E">45</mods:start>
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<mods:classification id="D9C0BEE71B5888EBD875BCB57F08EC9E">journal article</mods:classification>
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<mods:identifier id="47260ED6D9A8E037E497397160F5AC67" type="DOI">10.26107/RBZ-2021-0005</mods:identifier>
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<mods:identifier id="82FC3DF0BC9152CB8B1446A10AB7CA72" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="8B319BBC93267EB7CBF9AE76D640EC2A" type="Zenodo-Dep">5351678</mods:identifier>
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<treatment id="610CF11BE005FFDBFEEE24D6FEEBFBB6" ID-DOI="http://doi.org/10.5281/zenodo.5449955" ID-Zenodo-Dep="5449955" LSID="urn:lsid:zoobank.org:act:BD4AF315-A356-4F14-B4CE-19812342AD3D" httpUri="http://treatment.plazi.org/id/610CF11BE005FFDBFEEE24D6FEEBFBB6" lastPageId="12" lastPageNumber="57" pageId="10" pageNumber="55">
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<subSubSection id="A1BF1386E005FFDDFEEE24D6FD70FC36" box="[269,642,957,981]" pageId="10" pageNumber="55" type="nomenclature">
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<paragraph id="E91A400DE005FFDDFEEE24D6FD70FC36" blockId="10.[269,642,957,1013]" box="[269,642,957,981]" pageId="10" pageNumber="55">
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<heading id="B252F761E005FFDDFEEE24D6FD70FC36" bold="true" box="[269,642,957,981]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="55" reason="2">
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<emphasis id="DBD19C1FE005FFDDFEEE24D6FD70FC36" bold="true" box="[269,642,957,981]" pageId="10" pageNumber="55">
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<taxonomicName id="2EA53B8EE005FFDDFEEE24D6FE1FFC36" authority="Tran & Zettel, 2021" authorityName="Tran & Zettel" authorityYear="2021" box="[269,493,957,981]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="55" phylum="Arthropoda" rank="species" species="bendanilloi" status="sp. nov.">
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<emphasis id="DBD19C1FE005FFDDFEEE24D6FE1FFC36" bold="true" box="[269,493,957,981]" italics="true" pageId="10" pageNumber="55">Ranatra bendanilloi</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="C0E22164E005FFDDFE1F24D5FD70FC36" box="[508,642,957,981]" pageId="10" pageNumber="55" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A1BF1386E005FFDDFE6124B5FDFFFC16" box="[386,525,989,1013]" pageId="10" pageNumber="55" type="description">
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<paragraph id="E91A400DE005FFDDFE6124B5FDFFFC16" blockId="10.[269,642,957,1013]" box="[386,525,989,1013]" pageId="10" pageNumber="55">
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(
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<figureCitation id="719E5C88E005FFDDFE6924B5FE1BFC16" box="[394,489,989,1013]" captionStart="Fig" captionStartId="5.[113,148,2000,2022]" captionTargetBox="[113,1451,253,1984]" captionTargetId="figure-10@5.[113,1451,253,1984]" captionTargetPageId="5" captionText="Fig. 1. Habitus of Ranatra spp., all males. A, R. stali Montandon, 1905; B, R. bisaya, new species; C, R. bendanilloi, new species; D, R. brevicauda, new species; E, R. pangantihoni, new species (all of same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351680" httpUri="https://zenodo.org/record/5351680/files/figure.png" pageId="10" pageNumber="55">Figs. 1C</figureCitation>
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,
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<figureCitation id="719E5C88E005FFDDFE1424B5FDF7FC16" box="[503,517,989,1013]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">4</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="A1BF1386E005FFDDFF6B2375FE2AFAF6" pageId="10" pageNumber="55" type="materials_examined">
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<paragraph id="E91A400DE005FFDDFF6B2375FE2AFAF6" blockId="10.[136,776,1053,1301]" pageId="10" pageNumber="55">
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<emphasis id="DBD19C1FE005FFDDFF6B2375FE97FBD6" bold="true" box="[136,357,1053,1077]" pageId="10" pageNumber="55">
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<collectingRegion id="2B618EEFE005FFDDFF6B2375FF19FBD6" box="[136,235,1053,1077]" pageId="10" pageNumber="55">Material</collectingRegion>
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examined.
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||
</emphasis>
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||
<materialsCitation id="59CD4A50E005FFDDFE892375FF21FB96" collectionCode="USC" collectorName="Fidel Bendanillo" country="Philippines" location="Banica River" pageId="10" pageNumber="55" specimenCount="1" stateProvince="Negros" typeStatus="holotype">
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<emphasis id="DBD19C1FE005FFDDFE892375FE20FBD6" bold="true" box="[362,466,1053,1077]" pageId="10" pageNumber="55">
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<typeStatus id="361EFEAFE005FFDDFE892375FE20FBD6" box="[362,466,1053,1077]" pageId="10" pageNumber="55" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
(male): “Philippines: Negros Or.\ Valencia, Apolong,\
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||
<location id="EC7A16D6E005FFDDFE6D2355FDE4FBB6" LSID="urn:lsid:plazi:treatment:610CF11BE005FFDBFEEE24D6FEEBFBB6:EC7A16D6E005FFDDFE6D2355FDE4FBB6" box="[398,534,1085,1109]" country="Philippines" name="Banica River" pageId="10" pageNumber="55" stateProvince="Negros">Banica River</location>
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||
\ leg.
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<collectorName id="445025DBE005FFDDFDAD2355FD0FFBB6" box="[590,765,1085,1109]" pageId="10" pageNumber="55">Fidel Bendanillo</collectorName>
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” (
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<collectionCode id="8FB4D8C8E005FFDDFF732335FF39FB96" box="[144,203,1117,1141]" pageId="10" pageNumber="55">USC</collectionCode>
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||
)
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||
</materialsCitation>
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.
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||
<materialsCitation id="59CD4A50E005FFDDFF012335FDE2FB56" collectionCode="NHMW, UPLB, USC, ZCW, ZRC" collectorName="Fidel Bendanillo" country="Philippines" location="Banica River" pageId="10" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Negros Oriental" typeStatus="paratype">
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<emphasis id="DBD19C1FE005FFDDFF012335FEA5FB96" bold="true" box="[226,343,1117,1141]" pageId="10" pageNumber="55">
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<typeStatus id="361EFEAFE005FFDDFF012335FEA5FB96" box="[226,343,1117,1141]" pageId="10" pageNumber="55" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(
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<collectionCode id="8FB4D8C8E005FFDDFE8A2335FE35FB96" box="[361,455,1117,1141]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="10" pageNumber="55" type="Museum">NHMW</collectionCode>
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,
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||
<collectionCode id="8FB4D8C8E005FFDDFE362336FDEFFB96" box="[469,541,1118,1141]" country="Philippines" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="10" pageNumber="55">UPLB</collectionCode>
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,
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||
<collectionCode id="8FB4D8C8E005FFDDFDCF2335FD96FB96" box="[556,612,1117,1141]" pageId="10" pageNumber="55">USC</collectionCode>
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,
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<collectionCode id="8FB4D8C8E005FFDDFD902335FD40FB96" box="[627,690,1117,1141]" pageId="10" pageNumber="55">ZCW</collectionCode>
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,
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||
<collectionCode id="8FB4D8C8E005FFDDFD222335FD09FB96" box="[705,763,1117,1141]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="55">ZRC</collectionCode>
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||
):
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||
<collectingCountry id="91B2009DE005FFDDFF6B2315FEDBFB76" box="[136,297,1149,1173]" name="Philippines" pageId="10" pageNumber="55">PHILIPPINES</collectingCountry>
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:
|
||
<emphasis id="DBD19C1FE005FFDDFED72315FE7FFB76" bold="true" box="[308,397,1149,1173]" pageId="10" pageNumber="55">Negros:</emphasis>
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||
<collectingRegion id="2B618EEFE005FFDDFE762316FD5EFB76" box="[405,684,1149,1174]" country="Philippines" name="Negros Oriental" pageId="10" pageNumber="55">Negros Oriental Province</collectingRegion>
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||
,
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||
<specimenCount id="FFA38B84E005FFDDFD542315FCF0FB76" box="[695,770,1149,1173]" pageId="10" pageNumber="55" type="male">1 male</specimenCount>
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||
, same collection data as holotype
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="59CD4A50E005FFDDFDFC23F5FF1CFB16" country="Philippines" county="Valencia" location="Banica River" municipality="Apolong" pageId="10" pageNumber="55" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Negros Oriental" typeStatus="paratype">
|
||
<specimenCount id="FFA38B84E005FFDDFDFC23F5FD8CFB56" box="[543,638,1181,1205]" pageId="10" pageNumber="55" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="FFA38B84E005FFDDFD6D23F5FCF0FB56" box="[654,770,1181,1205]" pageId="10" pageNumber="55" type="female">5 females</specimenCount>
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,
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||
<collectingRegion id="2B618EEFE005FFDDFF6A23D6FE68FB36" box="[137,410,1213,1238]" country="Philippines" name="Negros Oriental" pageId="10" pageNumber="55">Negros Oriental Province</collectingRegion>
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,
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<collectingCounty id="007B3881E005FFDDFE4123D5FDF1FB36" box="[418,515,1213,1237]" pageId="10" pageNumber="55">Valencia</collectingCounty>
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,
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||
<collectingMunicipality id="097EDA77E005FFDDFDEF23D5FD9EFB36" box="[524,620,1213,1237]" pageId="10" pageNumber="55">Apolong</collectingMunicipality>
|
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,
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||
<location id="EC7A16D6E005FFDDFD9523D5FCF6FB36" LSID="urn:lsid:plazi:treatment:610CF11BE005FFDBFEEE24D6FEEBFBB6:EC7A16D6E005FFDDFD9523D5FCF6FB36" box="[630,772,1213,1237]" country="Philippines" county="Valencia" municipality="Apolong" name="Banica River" pageId="10" pageNumber="55" stateProvince="Negros Oriental">Banica River</location>
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.
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||
<collectingRegion id="2B618EEFE005FFDDFF6B23B5FF1CFB16" box="[136,238,1245,1269]" country="Philippines" name="Masbate" pageId="10" pageNumber="55">
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<emphasis id="DBD19C1FE005FFDDFF6B23B5FF1CFB16" bold="true" box="[136,238,1245,1269]" pageId="10" pageNumber="55">Masbate</emphasis>
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</collectingRegion>
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</materialsCitation>
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<emphasis id="DBD19C1FE005FFDDFF0D23B5FF01FB16" bold="true" box="[238,243,1245,1269]" pageId="10" pageNumber="55">:</emphasis>
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<materialsCitation id="59CD4A50E005FFDDFF1F23B5FE26FAF6" country="Philippines" county="Island" location="Tugbo River" municipality="Tugbo" pageId="10" pageNumber="55" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Masbate" typeStatus="paratype">
|
||
<specimenCount id="FFA38B84E005FFDDFF1F23B5FEA6FB16" box="[252,340,1245,1269]" pageId="10" pageNumber="55" type="male">6 males</specimenCount>
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,
|
||
<specimenCount id="FFA38B84E005FFDDFE8123B5FE3DFB16" box="[354,463,1245,1269]" pageId="10" pageNumber="55" type="female">4 females</specimenCount>
|
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,
|
||
<collectingRegion id="2B618EEFE005FFDDFE3F23B5FDC4FB16" box="[476,566,1245,1269]" country="Philippines" name="Masbate" pageId="10" pageNumber="55">Masbate</collectingRegion>
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<collectingCounty id="007B3881E005FFDDFDA323B5FD76FB16" box="[576,644,1245,1269]" pageId="10" pageNumber="55">Island</collectingCounty>
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||
,
|
||
<locationDeviation id="F8A8592FE005FFDDFD7223B5FF17FAF6" location="Masbate" pageId="10" pageNumber="55">
|
||
<quantity id="2E5DEDE8E005FFDDFD7223B5FD2CFB16" box="[657,734,1245,1269]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="55" unit="km" value="3.5">3.5 km</quantity>
|
||
SE
|
||
<collectingRegion id="2B618EEFE005FFDDFF6B2395FF17FAF6" box="[136,229,1277,1301]" country="Philippines" name="Masbate" pageId="10" pageNumber="55">Masbate</collectingRegion>
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="097EDA77E005FFDDFF112395FEC8FAF6" box="[242,314,1277,1301]" pageId="10" pageNumber="55">Tugbo</collectingMunicipality>
|
||
,
|
||
<location id="EC7A16D6E005FFDDFEA42395FE26FAF6" LSID="urn:lsid:plazi:treatment:610CF11BE005FFDBFEEE24D6FEEBFBB6:EC7A16D6E005FFDDFEA42395FE26FAF6" box="[327,468,1277,1301]" country="Philippines" county="Island" municipality="Tugbo" name="Tugbo River" pageId="10" pageNumber="55" stateProvince="Masbate">Tugbo River</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BF1386E005FFDBFF6B2255FE93FCF6" lastPageId="12" lastPageNumber="57" pageId="10" pageNumber="55" type="description">
|
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<paragraph id="E91A400DE005FFDDFF6B2255FEC1F9F6" blockId="10.[136,776,1341,1557]" pageId="10" pageNumber="55">
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||
<emphasis id="DBD19C1FE005FFDDFF6B2255FEE4FAB6" bold="true" box="[136,278,1341,1365]" pageId="10" pageNumber="55">Description.</emphasis>
|
||
General colouration: mostly light brown to brown; eyes dark brown; all coxae brown; all femora and tibiae usually distinctly annulated brown and yellow; fore tarsus mostly yellowish with dark brown apex; apices of middle and hind tibiae, middle and hind tarsi dark brown. Hemelytra mostly light coloured, usually with mottled greyish corium.
|
||
</paragraph>
|
||
<paragraph id="E91A400DE005FFDDFF6B2156FE89F896" blockId="10.[136,776,1597,1909]" pageId="10" pageNumber="55">
|
||
Measurements. Males: body length 31–32 (
|
||
<typeStatus id="361EFEAFE005FFDDFD892155FD3CF9B6" box="[618,718,1597,1621]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 32); length of siphon 8.7–9.3 (
|
||
<typeStatus id="361EFEAFE005FFDDFE572135FDE9F996" box="[436,539,1629,1653]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 9.0); width of head 2.65–2.73 (
|
||
<typeStatus id="361EFEAFE005FFDDFEE92115FE83F976" box="[266,369,1661,1685]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 2.71); interocular width 1.09–1.13 (
|
||
<typeStatus id="361EFEAFE005FFDDFF7321F5FF01F956" box="[144,243,1693,1717]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 1.12); width of eye 0.76–0.81 (
|
||
<typeStatus id="361EFEAFE005FFDDFDBA21F5FD4EF956" box="[601,700,1693,1717]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 0.79); pronotal length 7.8, anterior pronotal length 5.9; posterior pronotal length 3.3; anterior width of pronotum 2.30; posterior width of pronotum 2.98; lengths of leg segments: fore leg: coxa 4.8, femur 8.23, tibia 3.58, tarsus 0.75; middle leg: femur 11.2, tibia 10.0, tarsus 1.42; hind leg: femur 11.5, tibia 12.7, tarsus 1.84.
|
||
</paragraph>
|
||
<paragraph id="E91A400DE005FFDDFF6B20F5FAD7FEF6" blockId="10.[136,776,1949,2101]" lastBlockId="10.[835,1475,189,277]" pageId="10" pageNumber="55">Females: body length 32–38; length of siphon 8.4–9.3; width of head 2.68–3.04; interocular width 1.10–1.22; width of eye 0.79–0.92; pronotal length 8.06, anterior pronotal length 6.1; posterior pronotal length 3.5; anterior width of pronotum 2.48; posterior width of pronotum 3.2; lengths of leg segments: fore leg: coxa 5.25, femur 8.61, tibia 3.67, tarsus 0.76; middle leg: femur 11.7, tibia 10.25, tarsus 1.81; hind leg: femur 12.0, tibia 13.3, tarsus 2.02.</paragraph>
|
||
<paragraph id="E91A400DE005FFDDFCA02655FB5EFDB5" blockId="10.[835,1475,317,598]" pageId="10" pageNumber="55">
|
||
Head (
|
||
<figureCitation id="719E5C88E005FFDDFC712655FBE4FEB6" box="[914,1046,317,341]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">Fig. 4B, C</figureCitation>
|
||
): Vertex above eyes with an obtuse tubercle, narrow in dorsal view; width of eye clearly less than interocular width; clypeus smooth and convex, in lateral view about as high as or slightly higher than lora and slightly surpassing lora anteriorly; lora swollen, with a small, but distinct dorsal nodule, usually bearing few sparse, long, pale setae along dorsal side. Antenna (
|
||
<figureCitation id="719E5C88E005FFDDFB042695FACDFDF6" box="[1255,1343,509,533]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">Fig. 4A</figureCitation>
|
||
): finger-like projection of second antennal segment very short, about 0.2× the length of third segment.
|
||
</paragraph>
|
||
<paragraph id="E91A400DE005FFDDFCA02515FA31FAD6" blockId="10.[835,1475,637,1334]" pageId="10" pageNumber="55">
|
||
Thorax: Prothorax in lateral view distinctly longer than fore coxa (about 1.5–1.6× the length of fore coxa) and subequal to or slightly shorter than fore femur (0.92–0.99× the length of fore femur); anterior lobe about 1.7–1.9× as long as posterior lobe; anterior collar of pronotum distinctly raised when viewed laterally; ratio of posterior width / anterior width 1.25–1.36; posterior lobe with humeri broadly rounded, on each sublateral side with two swellings separated by a longitudinal groove (
|
||
<figureCitation id="719E5C88E005FFDDFBD12415FB42FC76" box="[1074,1200,893,917]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">Fig. 4B, C</figureCitation>
|
||
). Scutellum with length ca. 1.8–2.1× width, usually swollen at midpoint followed by a transverse depression before posterior third, apex pointed. Prosternum with paired broad, shallow longitudinal depressions separated by low and blunt median carina; median carina rather indistinct in posterior half. Mesosternum with slightly raised anterolateral margin, posterior projection between middle coxae truncate, weakly grooved along midline. Metasternum with anterior part grooved along midline, posterior part distinctly raised along midline and grooved sublaterally, posterior margin angularly emarginated (
|
||
<figureCitation id="719E5C88E005FFDDFCA823B5FC53FB16" box="[843,929,1245,1269]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">Fig. 4F</figureCitation>
|
||
). Space between middle coxae about same as or slightly greater than that between hind coxae. Hemelytra: membrane only reaching mid-length of abdominal tergum VI.
|
||
</paragraph>
|
||
<paragraph id="E91A400DE005FFDBFCA02236FCFAFE56" blockId="10.[835,1475,1373,2069]" lastBlockId="12.[136,776,189,437]" lastPageId="12" lastPageNumber="57" pageId="10" pageNumber="55">
|
||
Legs: Fore femur (
|
||
<figureCitation id="719E5C88E005FFDDFBFD2235FB6CFA96" box="[1054,1182,1373,1397]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="10" pageNumber="55">Fig. 4D, E</figureCitation>
|
||
): in both sexes relatively thick (ratio of length / maximum width of femur: 9.8–11.0,
|
||
<typeStatus id="361EFEAFE005FFDDFCA022F5FC52FA56" box="[835,928,1437,1461]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
10.4), widest at basal part, distal part nearly as wide as basal part, ratio in males: 1.13–1.17 (
|
||
<typeStatus id="361EFEAFE005FFDDFB1D22D5FAAFFA36" box="[1278,1373,1469,1493]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
1.17), in females: 1.13–1.19; flexor side with a median carina bearing dense, short setae and a tooth on mesal (anterior) surface situated distally to median carina; distal part with a pair of small teeth on lateral (posterior) surface flexor side, proximal to sinuous pre-apical ventral margin; distal teeth slightly longer than surrounding setae on flexor side of femur (more prominent in female); ratio of width of femur across median tooth (excluding tuft of setae) / width of femur at basal part: 1.07–1.14 (
|
||
<typeStatus id="361EFEAFE005FFDDFC2321B5FBEDF916" box="[960,1055,1757,1781]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
1.07); ratio of width of femur across median carina (excluding setae, on proximal side of median tooth) / width of femur at basal part: 0.97–1.02 (
|
||
<typeStatus id="361EFEAFE005FFDDFABF2075FA4DF8D6" box="[1372,1471,1821,1845]" pageId="10" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
: 1.00). Middle femur slightly shorter than hind femur; hind femur, when folded back parallel to body only reaching to anterior third of abdominal sternum VI (in males) or slightly surpassing anterior margin of sternum VI (in females). Middle tibia shorter than middle femur; hind tibia longer than hind femur; middle and hind tibiae both bearing dense fringes of long hairs on posterior margins along their distal two-thirds. Abdomen: Operculum of male slightly shorter than connexivum, medially keeled, apex pointed. Sternum VI of female with median keel distinctly raised on posterior half. Operculum of female clearly longer than connexivum, surpassing the apex of connexivum of about one third the length of operculum. Respiratory siphon clearly shorter (0.70–0.82×) than the length of sterna III–V combined, with sparse long, thin hairs along its length, more on apical part.
|
||
</paragraph>
|
||
<caption id="BDDA1085E004FFDCFF9220A9FE31F7F2" ID-DOI="http://doi.org/10.5281/zenodo.5351686" ID-Zenodo-Dep="5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="11" pageNumber="56" startId="11.[113,148,1985,2007]" targetBox="[123,1431,252,1963]" targetPageId="11">
|
||
<paragraph id="E91A400DE004FFDCFF9220A9FE31F7F2" blockId="11.[113,1452,1985,2065]" pageId="11" pageNumber="56">
|
||
Fig. 4.
|
||
<taxonomicName id="2EA53B8EE004FFDCFF5A20AAFE8EF835" authority="Tran & Zettel, 2021" authorityName="Tran & Zettel" authorityYear="2021" box="[185,380,1985,2007]" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="56" phylum="Arthropoda" rank="species" species="bendanilloi" status="sp. nov.">
|
||
<emphasis id="DBD19C1FE004FFDCFF5A20AAFE8EF835" box="[185,380,1985,2007]" italics="true" pageId="11" pageNumber="56">Ranatra bendanilloi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C0E22164E004FFDCFE6920AAFDF0F834" box="[394,514,1985,2007]" pageId="11" pageNumber="56" rank="species">new species</taxonomicNameLabel>
|
||
, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E91A400DE003FFDBFF6B26B5FE93FCF6" blockId="12.[136,776,477,789]" pageId="12" pageNumber="57">
|
||
Male genitalia: Paramere (
|
||
<figureCitation id="719E5C88E003FFDBFE5E26B5FDC7FE16" box="[445,565,477,501]" captionStart="Fig" captionStartId="11.[113,148,1985,2007]" captionTargetBox="[123,1431,252,1963]" captionTargetId="figure-11@11.[114,1452,239,1970]" captionTargetPageId="11" captionText="Fig. 4. Ranatra bendanilloi, new species, male. A, antenna; B, C, head and prothorax, lateral and dorsal views; D, E, fore femur, lateral and mesal views; F, metasternum; G, paramere, lateral view; H, I, apical part of paramere, sub-caudal and inner views. (B, C same scale; D, E same scale; G–I same scale)." figureDoi="http://doi.org/10.5281/zenodo.5351686" httpUri="https://zenodo.org/record/5351686/files/figure.png" pageId="12" pageNumber="57">Fig. 4G–I</figureCitation>
|
||
): thickest at basal third, constricted at distal quarter; ventral margin gradually tapering from middle part towards the constricted part, then followed by a large sub-triangular process bearing tuft of setae; on inner face, pre-apical process bearing a pointed projection slightly shorter than tuft of setae; apical hook long, slender with narrow, rounded tip; dorsal margin of paramere almost straight, slightly concave at same section with ventral, pre-apical process (ca. at distal fifth). Phallotheca strongly sclerotised.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BF1386E003FFDBFF6B2455FD2FFC76" pageId="12" pageNumber="57" type="etymology">
|
||
<paragraph id="E91A400DE003FFDBFF6B2455FD2FFC76" blockId="12.[136,775,829,917]" pageId="12" pageNumber="57">
|
||
<emphasis id="DBD19C1FE003FFDBFF6B2455FEFAFCB6" bold="true" box="[136,264,829,853]" pageId="12" pageNumber="57">Etymology.</emphasis>
|
||
This new species is dedicated to the entomologist and curator Mr. Fidel E. Bendanillo MSc from the University of San Carlos (
|
||
<collectingRegion id="2B618EEFE003FFDBFECE2415FE95FC76" box="[301,359,893,917]" country="Philippines" name="Cebu" pageId="12" pageNumber="57">Cebu</collectingRegion>
|
||
City) who collected the
|
||
<typeStatus id="361EFEAFE003FFDBFD9B2415FD2BFC76" box="[632,729,893,917]" pageId="12" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BF1386E003FFDBFF6B24D5FE5DFC16" pageId="12" pageNumber="57" type="discussion">
|
||
<paragraph id="E91A400DE003FFDBFF6B24D5FE5DFC16" blockId="12.[136,775,957,1013]" pageId="12" pageNumber="57">
|
||
<emphasis id="DBD19C1FE003FFDBFF6B24D5FF0BFC36" bold="true" box="[136,249,957,981]" pageId="12" pageNumber="57">Remarks.</emphasis>
|
||
For comparative notes, see Remarks under
|
||
<taxonomicName id="2EA53B8EE003FFDBFD1324D6FEE4FC16" authority="Tran & Zettel, 2021" authorityName="Tran & Zettel" authorityYear="2021" class="Insecta" family="Nepidae" genus="Ranatra" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="57" phylum="Arthropoda" rank="species" species="pangantihoni" status="sp. nov.">
|
||
<emphasis id="DBD19C1FE003FFDBFD1324D6FCF2FC36" box="[752,768,958,981]" italics="true" pageId="12" pageNumber="57">R</emphasis>
|
||
.
|
||
<emphasis id="DBD19C1FE003FFDBFF6B24B5FEE4FC16" box="[136,278,989,1013]" italics="true" pageId="12" pageNumber="57">pangantihoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="C0E22164E003FFDBFEC524B6FE59FC16" box="[294,427,989,1013]" pageId="12" pageNumber="57" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A1BF1386E003FFDBFF6B2375FEEBFBB6" pageId="12" pageNumber="57" type="distribution">
|
||
<paragraph id="E91A400DE003FFDBFF6B2375FEEBFBB6" blockId="12.[136,775,1053,1109]" pageId="12" pageNumber="57">
|
||
<emphasis id="DBD19C1FE003FFDBFF6B2375FEE8FBD6" bold="true" box="[136,282,1053,1077]" pageId="12" pageNumber="57">Distribution.</emphasis>
|
||
<collectingCountry id="91B2009DE003FFDBFEC72375FE52FBD6" box="[292,416,1053,1077]" name="Philippines" pageId="12" pageNumber="57">Philippines</collectingCountry>
|
||
: Greater Visayas region: Negros and
|
||
<collectingRegion id="2B618EEFE003FFDBFF542355FEE7FBB6" box="[183,277,1085,1109]" country="Philippines" name="Masbate" pageId="12" pageNumber="57">Masbate</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |