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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.746.22020" ID-GBIF-Dataset="398e247a-455e-44af-b2ac-abb8b9a4942b" ID-PMC="PMC5904419" ID-Pensoft-Pub="1313-2970-746-51" ID-PubMed="29674896" ID-ZBK="DF12B2D041554B10B3A3A0B1E755DFF6" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-746-51" ModsDocOrigin="ZooKeys 746" ModsDocTitle="Deep intraspecific DNA barcode splits and hybridisation in the Udeaalpinalis group (Insecta, Lepidoptera, Crambidae) an integrative revision" checkinTime="1522144730391" checkinUser="pensoft" docAuthor="Mally, Richard, Huemer, Peter &amp; Nuss, Matthias" docDate="2018" docId="304D406A9DDE2627AC80B8EC88FB43A2" docLanguage="en" docName="ZooKeys 746: 51-90" docOrigin="ZooKeys 746" docSource="http://dx.doi.org/10.3897/zookeys.746.22020" docTitle="Udea altaica Zerny 1914, stat. n." docType="treatment" docVersion="4" lastPageNumber="70" masterDocId="5C28FFE7FF8D5848D4017C1B78380C2D" masterDocTitle="Deep intraspecific DNA barcode splits and hybridisation in the Udeaalpinalis group (Insecta, Lepidoptera, Crambidae) - an integrative revision" masterLastPageNumber="90" masterPageNumber="51" pageNumber="68" updateTime="1668165631775" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Deep intraspecific DNA barcode splits and hybridisation in the Udeaalpinalis group (Insecta, Lepidoptera, Crambidae) - an integrative revision</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mally, Richard</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huemer, Peter</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nuss, Matthias</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>746</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>90</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.746.22020</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.746.22020</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-746-51</mods:identifier>
<mods:identifier type="ZBK">DF12B2D041554B10B3A3A0B1E755DFF6</mods:identifier>
<mods:identifier type="ZooBank">DF12B2D041554B10B3A3A0B1E755DFF6</mods:identifier>
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<treatment ID-GBIF-Taxon="142656901" LSID="urn:lsid:plazi:treatment:304D406A9DDE2627AC80B8EC88FB43A2" httpUri="http://treatment.plazi.org/id/304D406A9DDE2627AC80B8EC88FB43A2" lastPageId="19" lastPageNumber="70" pageId="17" pageNumber="68">
<subSubSection pageId="17" pageNumber="68" type="nomenclature">
<paragraph pageId="17" pageNumber="68">
<taxonomicName authority="Zerny, 1914" authorityName="Zerny" authorityYear="1914" class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea altaica" order="Lepidoptera" pageId="17" pageNumber="68" phylum="Arthropoda" rank="species" species="altaica">Udea altaica (Zerny, 1914)</taxonomicName>
<taxonomicNameLabel pageId="17" pageNumber="68">stat. n.</taxonomicNameLabel>
Figs 6, 15-18, 36-37, 47
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="68" type="reference_group">
<paragraph pageId="17" pageNumber="68">
<taxonomicName class="Insecta" family="Crambidae" genus="Pyrausta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrausta austriacalis" order="Lepidoptera" pageId="17" pageNumber="68" phylum="Arthropoda" rank="species" species="austriacalis">Pyrausta austriacalis</taxonomicName>
v.
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea altaica" order="Lepidoptera" pageId="17" pageNumber="68" phylum="Arthropoda" rank="species" species="altaica">altaica</taxonomicName>
Zerny, 1914: 334-335.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="68" type="type locality">
<paragraph pageId="17" pageNumber="68">Type locality.</paragraph>
<paragraph pageId="17" pageNumber="68">Mongolia, central Altai mountains.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="68" type="materials_examined">
<paragraph pageId="17" pageNumber="68">Material examined.</paragraph>
<paragraph pageId="17" pageNumber="68">
Type specimens. Lectotype ♀ &quot;Altai centr. | mont.&quot;, &quot;Stgr. | [handwritten] 1914&quot;,
<normalizedToken originalValue="“667.”">&quot;667.&quot;</normalizedToken>
, [handwritten] &quot;P. austriacalis | v. altaica | Zerny ♀ [in red] Type&quot;, Mally prep. no. 1084 (NMW); Paralectotype ♂ (abdomen lost) &quot;Altai centr. | mont.&quot;, &quot;Stgr. | [handwritten] 1914&quot;,
<normalizedToken originalValue="“666.”">&quot;666.&quot;</normalizedToken>
, [handwritten] &quot;P. austriacalis | v. altaica | Zerny ♂ [in red] Type&quot; (NMW). - Additional material. MONGOLIA. 3♂ 2♀
<normalizedToken originalValue="“Altai”">&quot;Altai&quot;</normalizedToken>
, one of the ♂ also with [handwritten] &quot;Alticolalis | BH i L&quot;, Mally prep. no. 1099 (♀), 1100 (♀), 1117-1119 (♂) (ZMHB); 1♂ 1♀ [handwritten] &quot;Pyrausta | Alticolalis | Altai BH&quot;, Mally prep. no. 1090 (♂) &amp; 1098 (♀) (ZMHB).
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="68" type="diagnosis">
<paragraph pageId="17" pageNumber="68">Diagnosis.</paragraph>
<paragraph pageId="17" pageNumber="68">
Proximal outer spur of hindleg minute (as in
<taxonomicName lsidName="U. alpinalis" pageId="17" pageNumber="68" rank="species" species="alpinalis">U. alpinalis</taxonomicName>
,
<taxonomicName lsidName="U. juldusalis" pageId="17" pageNumber="68" rank="species" species="juldusalis">U. juldusalis</taxonomicName>
and
<taxonomicName lsidName="U. plumbalis" pageId="17" pageNumber="68" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
), whereas in
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
,
<taxonomicName lsidName="U. cretacea" pageId="17" pageNumber="68" rank="species" species="cretacea">U. cretacea</taxonomicName>
,
<taxonomicName lsidName="U. donzelalis" pageId="17" pageNumber="68" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
and
<taxonomicName lsidName="U. uliginosalis" pageId="17" pageNumber="68" rank="species" species="uliginosalis">U. uliginosalis</taxonomicName>
it is ca. half to two thirds the length of the proximal inner spur. The wing maculation of
<taxonomicName lsidName="U. altaica" pageId="17" pageNumber="68" rank="species" species="altaica">U. altaica</taxonomicName>
can be confused with that of
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
,
<taxonomicName lsidName="U. cretacea" pageId="17" pageNumber="68" rank="species" species="cretacea">U. cretacea</taxonomicName>
,
<taxonomicName lsidName="U. donzelalis" pageId="17" pageNumber="68" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
,
<taxonomicName lsidName="U. juldusalis" pageId="17" pageNumber="68" rank="species" species="juldusalis">U. juldusalis</taxonomicName>
,
<taxonomicName lsidName="U. plumbalis" pageId="17" pageNumber="68" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
,
<taxonomicName lsidName="U. uliginosalis" pageId="17" pageNumber="68" rank="species" species="uliginosalis">U. uliginosalis</taxonomicName>
and untypically maculated specimens of
<taxonomicName lsidName="U. alpinalis" pageId="17" pageNumber="68" rank="species" species="alpinalis">U. alpinalis</taxonomicName>
(see Fig. 4 in
<bibRefCitation author="Panigaj, L" journalOrPublisher="Vestnik zoologii, Kyiv" pageId="33" pageNumber="84" pagination="41 - 45" title="Distribution and bionomics of Udeaalpinalis (Lepidoptera, Pyralidae) in western Carpathians (Slovakia)." url="https://doi.org/10.2478/v10058-012-0024-y" volume="46" year="2012">Panigaj and Kulfan 2012</bibRefCitation>
), but it can be distinguished from all those species by the more or less distinct proximal brown section of the postmedial line on the ventral side of the forewing in both sexes (Figs 16, 18); in males, the proximal subterminal area of the ventral forewing side is as light brown as the central area (Fig. 16), whereas in the other species it is darker than the central area; on the hindwing ventral side, the subterminal area is only faintly darker than the central wing area in both sexes (Figs 16, 18), whereas the other species have a darker subterminal area, at least in the apex. In male genitalia only distinguishable from
<taxonomicName lsidName="U. cretacea" pageId="17" pageNumber="68" rank="species" species="cretacea">U. cretacea</taxonomicName>
and
<taxonomicName lsidName="U. uliginosalis" pageId="17" pageNumber="68" rank="species" species="uliginosalis">U. uliginosalis</taxonomicName>
by the small dentate ridge-like process on the posterior phallus apodeme, whereas in
<taxonomicName lsidName="U. cretacea" pageId="17" pageNumber="68" rank="species" species="cretacea">U. cretacea</taxonomicName>
the sclerotisation at posterior phallus apodeme is a slim, elongate, apically dentate process emerging from the posteriormost end (Fig. 35), in
<taxonomicName lsidName="U. uliginosalis" pageId="17" pageNumber="68" rank="species" species="uliginosalis">U. uliginosalis</taxonomicName>
a large hooked spine. In the female genitalia, the main signum is 3.1- to 3.4-times as long as broad, whereas in
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
the main signum is 3.4- to 4.4-times longer and in
<taxonomicName lsidName="U. donzelalis" pageId="17" pageNumber="68" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
4.1- to 5.1-times longer than its maximum width (Tab. 1). The antrum is conical, widening posteriorly and is about twice as long as broad (Fig. 47), whereas in
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
and
<taxonomicName lsidName="U. donzelalis" pageId="17" pageNumber="68" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
the sclerotised antrum is predominantly tubular and 1- to 1.5-times as long as broad (Figs 45-46).
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="69" pageId="17" pageNumber="68" type="redescription">
<paragraph pageId="17" pageNumber="68">Redescription.</paragraph>
<paragraph pageId="17" pageNumber="68">
Head. As for
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
, part from: frons and vertex with cream-white scales; distal half of labial palps brown on the outside, basal half and inner sides cream white; maxillary palps cream-white with some brown scales mixed in; antennal length approx. 60 % of forewing length in males, approx. 70 % in females.
</paragraph>
<paragraph lastPageId="18" lastPageNumber="69" pageId="17" pageNumber="68">
Thorax. As for
<taxonomicName lsidName="U. austriacalis" pageId="17" pageNumber="68" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
, apart from: legs cream-white on inner and outer sides; proximal pair of metatarsal spurs with outer spur minute and inner spur long,
<pageBreakToken pageId="18" pageNumber="69" start="start">distal</pageBreakToken>
pair ca. half the length of the proximal inner spur, distal inner spur a bit longer than distal outer spur.
</paragraph>
<paragraph pageId="18" pageNumber="69">Wings. Forewing length 14 mm in males, 11 mm in females. Males and females with one frenulum bristle. Female forewing with more acute apex due to the straight costa (distally curved in males), and with outer wing margin (termen) of hindwing cut straight. Forewing upper side cream white with brown scales interspersed, giving it a dirty appearance; brownish subcostal line along the basal two third of the forewing; cell margin facing the forewing centre demarcated by a thin brown line, less prominent in females; outer medial area with a transverse cream white band lacking the interspersed brown scales; outer margin of cream white band delimited by diffuse grey postmedial line which leaves the costa in a right angle, bends inward at vein M1 and parallels the termen until the line reaches the dorsum; postmedial area homogenous greyish white, apex with more or less darker streak; termen with a slim brown margin and long, cream-white fringes. Hindwing upper side in males pale yellowish brown with diffuse light brown apex, in females light brown with a faint, slightly brighter medial band. Forewing underside in females vivid brown, with a darker, diffuse outer cellular spot and a darker postmedial area, demarcated by the postmedial line, maculation paler in males; slim whitish subcostal line along the basal two third of the forewing; fringes cream-white. Hindwing underside cream white with the subcostal and terminal areas tinted slightly brownish, the postmedial line more or less prominent.</paragraph>
<paragraph pageId="18" pageNumber="69">Abdomen. Pale grey dorsally, slightly darker grey ventrally; distal segment margins greyish white, scales on terminal segment pale yellowish. Tympanum without broad short lobulus.</paragraph>
<paragraph pageId="18" pageNumber="69">
Male genitalia. (Figs 36-37) As for
<taxonomicName lsidName="U. austriacalis" pageId="18" pageNumber="69" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
, apart from: juxta nearly rhombical to broad drop-shaped, with small indention on each side dorsal of its greatest width, apex sharply bifid with narrow V-shaped medial incision ca. 1/4 of juxta length; ventral valva edge convex, with a slight bulge in the area to which the fibula is pointing; valva apex evenly rounded. An elongate triangular, apically tapering, strongly sclerotised fibula directed towards the distal sacculus, apical half narrowed to pointed, ventrad curved claw, ventral side of claw with flat
<normalizedToken originalValue="blade">'blade'</normalizedToken>
; fibula emerging from an oval sclerotised lobe near base of costa which is very sparsely studded with thin long simple setae; posterior phallus apodeme dorsally and ventrally with elongate unsclerotised strip, right posterior phallus apodeme forming a sclerotised spatulate lobe with a medially protruding ridge containing at the posterior end two equal-sized teeth and a third tiny posterior-most tooth.
</paragraph>
<paragraph pageId="18" pageNumber="69">
Female genitalia. (Fig. 47) As for
<taxonomicName lsidName="U. austriacalis" pageId="18" pageNumber="69" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
, apart from: signum 3.1- to 3.4-times as long as broad (n = 4); auxillary signum 52-67 % length of main signum (n=4); antrum conical, widening posteriorly, sclerotised section about twice as long as its diameter, with or without a narrowly V-shaped unsclerotised longitudinal indentation in the
<normalizedToken originalValue="antrums">antrum's</normalizedToken>
dorsodistal sclerotisation; apophyses posteriores slender, approx. 60-80 % the length of the apophyses anteriores.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="69" type="immature stages">
<paragraph pageId="18" pageNumber="69">Immature stages.</paragraph>
<paragraph pageId="18" pageNumber="69">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="70" type="distribution">
<paragraph pageId="19" pageNumber="70">
<pageBreakToken pageId="19" pageNumber="70" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="19" pageNumber="70">
The species is known from the central Altai mountains in NW Mongolia, the
<normalizedToken originalValue="Küngöy">Kuengoey</normalizedToken>
Ala-Too (Kungey Alatau) Range in Kazakhstan and Kyrgyzstan, and the Yulduz mountains in NW China (see Fig. 6); the
<normalizedToken originalValue="Küngöy">Kuengoey</normalizedToken>
Ala-Too and Yulduz mountain ranges are part of the Tian Shan mountains.
</paragraph>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figure 6. Distribution of investigated specimens of
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea juldusalis" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="juldusalis">Udea juldusalis</taxonomicName>
(blue),
<taxonomicName lsidName="U. altaica" pageId="19" pageNumber="70" rank="species" species="altaica">U. altaica</taxonomicName>
(red) and
<taxonomicName lsidName="U. plumbalis" pageId="19" pageNumber="70" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
(yellow) in Central Asia; altitudes ³ 1,000 m are marked in increasingly darker grey shades every 500 m, altitudes ³ 4,000 m are in black. Note that the eastern locality of
<taxonomicName lsidName="U. juldusalis" pageId="19" pageNumber="70" rank="species" species="juldusalis">U. juldusalis</taxonomicName>
and the localities of
<taxonomicName lsidName="U. altaica" pageId="19" pageNumber="70" rank="species" species="altaica">U. altaica</taxonomicName>
and
<taxonomicName lsidName="U. plumbalis" pageId="19" pageNumber="70" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
are only approximations of the type localities.
</paragraph>
</caption>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figures 7-14. Adult specimens of
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="genus">Udea</taxonomicName>
species. 7-10
<taxonomicName lsidName="U. austriacalis" pageId="19" pageNumber="70" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
7-8 male, dorsal (7) and ventral (8) 9-10 female, dorsal (9) and ventral (10), abdomen removed 11-14
<taxonomicName lsidName="U. donzelalis" pageId="19" pageNumber="70" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
11-12 male, dorsal (11) and ventral (12) 13-14 female, dorsal (13) and ventral (14), abdomen removed. Scale bars: 500
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figures 15-22. Adult specimens of
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="genus">Udea</taxonomicName>
species. 15-18
<taxonomicName lsidName="U. altaica" pageId="19" pageNumber="70" rank="species" species="altaica">U. altaica</taxonomicName>
15-16 male, dorsal (15) and ventral (16) 17-18 Lectotype (NHMW) female, dorsal (17) and ventral (18) 19-20
<taxonomicName lsidName="U. juldusalis" pageId="19" pageNumber="70" rank="species" species="juldusalis">U. juldusalis</taxonomicName>
Lectotype (NHMW) male, dorsal (19) and ventral (20) 21-22
<taxonomicName lsidName="U. plumbalis" pageId="19" pageNumber="70" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
Holotype (NHMW) male, dorsal (21) and ventral (22). Scale bar: 500
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figures 23-35. Male genitalia of the
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea austriacalis" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="austriacalis">Udea austriacalis</taxonomicName>
species complex. 23-28
<taxonomicName lsidName="U. austriacalis" pageId="19" pageNumber="70" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
23 male genitalia (Mally prep. 1092) 24-28 posterior phallus apodeme 24 Mally prep. 1042 25 Mally prep. 1043 26 Mally prep. 1044 27 Mally prep. 1045 28 Mally prep. 1046 29-33
<taxonomicName lsidName="U. donzelalis" pageId="19" pageNumber="70" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
29 male genitalia (Mally prep. 1024) 30-33 posterior phallus apodeme 30 Mally prep. 1024 31 Mally prep. 866 32 Mally prep. 867 33 Mally prep. 1022 34-35
<taxonomicName lsidName="U. cretacea" pageId="19" pageNumber="70" rank="species" species="cretacea">U. cretacea</taxonomicName>
(Mally prep. 523) 34 male genitalia 35 posterior phallus apodeme; 500
<normalizedToken originalValue="µm">µm</normalizedToken>
scale bar refers to male genitalia, 200
<normalizedToken originalValue="µm">µm</normalizedToken>
scale bar to posterior phallus apodemes.
</paragraph>
</caption>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figures 36-44. Male genitalia of
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="genus">Udea</taxonomicName>
species. 36-37
<taxonomicName lsidName="U. altaica" pageId="19" pageNumber="70" rank="species" species="altaica">U. altaica</taxonomicName>
(Mally prep. 1090) 36 male genitalia 37 posterior phallus apodeme 38-41
<taxonomicName lsidName="U. juldusalis" pageId="19" pageNumber="70" rank="species" species="juldusalis">U. juldusalis</taxonomicName>
38 male genitalia, Paralectotype (Mally prep. 1081) 39-41 posterior phallus apodeme 39 Paralectotype (Mally prep. 1081) 40 Lectotype (Mally prep. 1082) 41 (Mally prep. 1089) 42-44
<taxonomicName lsidName="U. plumbalis" pageId="19" pageNumber="70" rank="species" species="plumbalis">U. plumbalis</taxonomicName>
42 male genitalia, Holotype (Mally prep. 1083) 43-44 posterior phallus apodeme 43 Holotype (Mally prep. 1083) 44 (Mally prep. 1094); 500
<normalizedToken originalValue="µm">µm</normalizedToken>
scale bar refers to male genitalia, 200
<normalizedToken originalValue="µm">µm</normalizedToken>
scale bar to posterior phallus apodemes.
</paragraph>
</caption>
<caption pageId="19" pageNumber="70">
<paragraph pageId="19" pageNumber="70">
Figures 45-47. Female genitalia of
<taxonomicName class="Insecta" family="Crambidae" genus="Udea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udea" order="Lepidoptera" pageId="19" pageNumber="70" phylum="Arthropoda" rank="genus">Udea</taxonomicName>
species. 45
<taxonomicName lsidName="U. austriacalis" pageId="19" pageNumber="70" rank="species" species="austriacalis">U. austriacalis</taxonomicName>
(Mally prep. 1047) 46
<taxonomicName lsidName="U. donzelalis" pageId="19" pageNumber="70" rank="species" species="donzelalis">U. donzelalis</taxonomicName>
(Mally prep. 1023) 47
<taxonomicName lsidName="U. altaica" pageId="19" pageNumber="70" rank="species" species="altaica">U. altaica</taxonomicName>
(Mally prep. 1084).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="70" type="food plants">
<paragraph pageId="19" pageNumber="70">Food plants.</paragraph>
<paragraph pageId="19" pageNumber="70">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="70" type="dna data">
<paragraph pageId="19" pageNumber="70">DNA data.</paragraph>
<paragraph pageId="19" pageNumber="70">Unavailable.</paragraph>
</subSubSection>
</treatment>
</document>