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<document id="5D43EC36CE99EDC35FDDCDE8F0D43F8D" ID-DOI="10.1093/zoolinnean/zlad121" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241136" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1716363944006" checkinUser="plazi" docAuthor="Bronner, Gary N., Mynhardt, Samantha, Bennett, Nigel C., Cohen, Lientjie, Crumpton, Nick, Hofreiter, Michael, Arnold, Patrick &amp; Asher, Robert J." docDate="2024" docId="582487E9FFEBFFACD5EB6476FD1522F3" docLanguage="en" docName="zlad121.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docSource="https://www.mendeley.com/catalogue/7d928dc7-e8de-32a4-8d69-64eca580d036/" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Tenrecidae Gray 1821" docType="treatment" docVersion="1" lastPageNumber="203" masterDocId="A41DFF91FFF8FFBFD4616629FFF4275F" masterDocTitle="Phylogenetic history of golden moles and tenrecs (Mammalia: Afrotheria)" masterLastPageNumber="213" masterPageNumber="184" pageNumber="203" updateTime="1716819710427" updateUser="julia">
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<mods:title id="3AEF570116E3DEBA64E0542F048974AB">Phylogenetic history of golden moles and tenrecs (Mammalia: Afrotheria)</mods:title>
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<mods:namePart id="5EE969838C48C1FD9AED3D8887CDBA75">Mynhardt, Samantha</mods:namePart>
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<taxonomicName id="178D4D7CFFEBFFACD5EB6476FDF42526" ID-CoL="GZG" authority="Gray, 1825" authorityName="Gray" authorityYear="1821" box="[394,512,607,633]" class="Mammalia" family="Tenrecidae" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="family">
<emphasis id="E2F9EAEDFFEBFFACD5EB6476FDF42526" bold="true" box="[394,512,607,633]" pageId="19" pageNumber="203">Tenrecidae</emphasis>
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<paragraph id="D03236FFFFEBFFACD4E064ACFD18238E" blockId="19.[129,778,607,1453]" pageId="19" pageNumber="203">
Our results for tenrecids at the Linnean rank of family and below are generally consistent with the taxonomy used by Bronner and Jenkins (2005) and
<bibRefCitation id="B41C4B0EFFEBFFACD5E664EDFD6D2583" author="Asher RJ &amp; Helgen KM" box="[391,665,708,732]" pageId="19" pageNumber="203" pagination="102" refId="ref18777" refString="Asher RJ, Helgen KM. Nomenclature and placental mammal phylogeny. BMC Evolutionary Biology 2010; 10: 102." type="journal article" year="2010">Asher and Helgen (2010)</bibRefCitation>
. This supports division of tenrecids into monophyletic potamogalines and tenrecines, the web-footed tenrec (
<taxonomicName id="178D4D7CFFEBFFACD622652AFEF42465" authority="Major, 1896" authorityName="Major" authorityYear="1896" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="mergulus">
<emphasis id="E2F9EAEDFFEBFFACD622652AFCFE2444" box="[579,778,771,795]" italics="true" pageId="19" pageNumber="203">Microgale mergulus</emphasis>
Major, 1896
</taxonomicName>
) as part of the genus
<taxonomicName id="178D4D7CFFEBFFACD5B7650BFDC12465" authorityName="Thomas" authorityYear="1882" box="[470,565,802,826]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD5B7650BFDC12465" box="[470,565,802,826]" italics="true" pageId="19" pageNumber="203">Microgale</emphasis>
</taxonomicName>
(as first proposed by Olson and Goodman 2003), a
<taxonomicName id="178D4D7CFFEBFFACD5D96568FD0F2406" authority="Thomas, 1884" authorityName="Thomas" authorityYear="1884" box="[440,763,833,857]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="dobsoni">
<emphasis id="E2F9EAEDFFEBFFACD5D96568FD922406" box="[440,614,833,857]" italics="true" pageId="19" pageNumber="203">Microgale dobsoni</emphasis>
Thomas, 1884
</taxonomicName>
<emphasis id="E2F9EAEDFFEBFFACD69D656BFCFE2406" box="[764,778,834,857]" italics="true" pageId="19" pageNumber="203"></emphasis>
<taxonomicName id="178D4D7CFFEBFFACD4E06548FE582426" authority="Major, 1896" authorityName="Major" authorityYear="1896" box="[129,428,865,889]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="talazaci">
<emphasis id="E2F9EAEDFFEBFFACD4E06548FEDA2426" box="[129,302,865,889]" italics="true" pageId="19" pageNumber="203">Microgale talazaci</emphasis>
Major, 1896
</taxonomicName>
clade as sister taxon to all other species of
<taxonomicName id="178D4D7CFFEBFFACD4A665A9FED224C7" authorityName="Thomas" authorityYear="1882" box="[199,294,896,920]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD4A665A9FED224C7" box="[199,294,896,920]" italics="true" pageId="19" pageNumber="203">Microgale</emphasis>
</taxonomicName>
, and an
<taxonomicName id="178D4D7CFFEBFFACD51A65A9FD6924C7" authority="Grandidier, 1870" authorityName="Grandidier" authorityYear="1870" box="[379,669,896,920]" class="Mammalia" family="Tenrecidae" genus="Oryzorictes" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD51A65A9FE1E24C8" box="[379,490,896,919]" italics="true" pageId="19" pageNumber="203">Oryzorictes</emphasis>
Grandidier, 1870
</taxonomicName>
<taxonomicName id="178D4D7CFFEBFFACD6CA65A9FCFE24C7" authorityName="Thomas" authorityYear="1882" box="[683,778,896,920]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD6CA65A9FCFE24C7" box="[683,778,896,920]" italics="true" pageId="19" pageNumber="203">Microgale</emphasis>
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clade, which we call Oryzorictini. This clade comprises the sister taxon of
<taxonomicName id="178D4D7CFFEBFFACD4BD6596FD292488" authority="Milne-Edwards &amp; Grandidier, 1871" authorityName="Milne-Edwards &amp; Grandidier" authorityYear="1871" box="[220,733,959,983]" class="Mammalia" family="Tenrecidae" genus="Geogale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="aurita">
<emphasis id="E2F9EAEDFFEBFFACD4BD6596FE9E2489" box="[220,362,959,983]" italics="true" pageId="19" pageNumber="203">Geogale aurita</emphasis>
Milne-Edwards &amp; Grandidier, 1871
</taxonomicName>
and collectively forms the Oryzorictinae (
<figureCitation id="48B62A7AFFEBFFACD66F65F7FDA824A9" box="[526,604,990,1014]" captionStart="Figure 10" captionStartId="16.[113,178,1739,1763]" captionTargetBox="[116,1457,145,1708]" captionTargetId="graphics-7@16.[116,1457,145,1709]" captionTargetPageId="16" captionText="Figure 10. Topologies resulting from Bayesian-BIC (A, left) and parsimony implied weighting K = 4 (B, right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a) and indels.Bayesian branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are in red, tenrecids green.Circled and coloured letters adjacent to nodes indicate high-level taxonomy: Am, Amblysominae; Cd, Chrysochloridae; Cn, Chrysochlorinae; Cx, Chrysospalacinae; Gi, Geogalini; Mg, Microgale; Oi, Oryzorictini; Or, Oryzorictinae; Po, Potamogalinae; Sa, Setiferina; Ta, Tenrecina; Td, Tenrecidae; Tn, Tenrecinae; Ti, Tenrecini." figureDoi="http://doi.org/10.5281/zenodo.11241156" httpUri="https://zenodo.org/record/11241156/files/figure.png" pageId="19" pageNumber="203">Figs 10</figureCitation>
,
<figureCitation id="48B62A7AFFEBFFACD60A65F7FD7124A9" box="[619,645,990,1014]" captionStart="Figure 11" captionStartId="17.[129,194,1821,1845]" captionTargetBox="[131,1471,147,1792]" captionTargetId="graphics-8@17.[131,1472,147,1792]" captionTargetPageId="17" captionText="Figure 11. Topologies resulting from Bayesian-BIC (A; left) and parsimony implied weighting K = 4 (B; right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a), indels and morphology.Branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are red, tenrecids green. Circled and coloured letters adjacent to nodes indicate high-level taxonomy, as shown in Figure 10." figureDoi="http://doi.org/10.5281/zenodo.11241158" httpUri="https://zenodo.org/record/11241158/files/figure.png" pageId="19" pageNumber="203">11</figureCitation>
). The taxon Geogalini is available to encompass fossil species more closely related to
<taxonomicName id="178D4D7CFFEBFFACD48C6234FEBD236B" authorityName="Milne-Edwards &amp; Grandidier" authorityYear="1871" box="[237,329,1053,1076]" class="Mammalia" family="Tenrecidae" genus="Geogale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="aurita">
<emphasis id="E2F9EAEDFFEBFFACD48C6234FEBD236B" box="[237,329,1053,1076]" italics="true" pageId="19" pageNumber="203">G. aurita</emphasis>
</taxonomicName>
than to other tenrecids, such as the fossil taxa
<taxonomicName id="178D4D7CFFEBFFACD4D36215FED0230B" authorityName="Butler" authorityYear="1984" box="[178,292,1084,1108]" class="Mammalia" family="Tenrecidae" genus="Parageogale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD4D36215FED0230B" box="[178,292,1084,1108]" italics="true" pageId="19" pageNumber="203">Parageogale</emphasis>
</taxonomicName>
and
<taxonomicName id="178D4D7CFFEBFFACD53B6215FE2B230B" authorityName="Butler &amp; Hopwood" authorityYear="1957" box="[346,479,1084,1108]" class="Mammalia" family="Tenrecidae" genus="Erythrozootes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD53B6215FE2B230B" box="[346,479,1084,1108]" italics="true" pageId="19" pageNumber="203">Erythrozootes</emphasis>
</taxonomicName>
(
<figureCitation id="48B62A7AFFEBFFACD5906215FDCD230B" box="[497,569,1084,1108]" captionStart="Figure 12" captionStartId="18.[113,174,1850,1874]" captionTargetBox="[127,1447,150,1829]" captionTargetId="graphics-7@18.[127,1446,161,1816]" captionTargetPageId="18" captionText="Figure 12.Topologies resultingfrom Bayesian-BIC (A; left)and parsimonyimplied weightingK = 4 (B;right) analyses of extanttaxasampled for six genes (cytB, ND2, GHR, vWF, 12S andstat5a)and indels,plus fossilsand morphologicaldata sampled forall.Bayesian branch lengthsreflect the scaleat the top left;parsimonybranch lengthsare arbitrary.Only chrysochloridsand Procavia are knownfor stat5a(andother regionsof missing data are shown in Table1). Bayesian nodes have a posterior probabilityof oneunlessindicated otherwise.Numbersadjacentto parsimonynodes represent bootstrap values,calculatedwith 250pseudoreplicates of a TNTsearchusing mult = tbr 10 replicateseach andreported onlyat orover50.Chrysochlorids are red, tenrecidsgreen;fossils are darkershadesof each.Circled and colouredlettersadjacent tonodes indicate high-leveltaxonomy,as shown in Figure10." figureDoi="http://doi.org/10.5281/zenodo.11241160" httpUri="https://zenodo.org/record/11241160/files/figure.png" pageId="19" pageNumber="203">Fig. 12</figureCitation>
). Tenrecini consists of
<taxonomicName id="178D4D7CFFEBFFACD4FC6272FE9F232C" authority="Froriep, 1806" authorityName="Froriep" authorityYear="1806" box="[157,363,1115,1139]" class="Mammalia" family="Tenrecidae" genus="Setifer" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD4FC6272FF2D232C" box="[157,217,1115,1139]" italics="true" pageId="19" pageNumber="203">Setifer</emphasis>
Froriep, 1806
</taxonomicName>
<emphasis id="E2F9EAEDFFEBFFACD50D6272FE8D232C" box="[364,377,1115,1139]" italics="true" pageId="19" pageNumber="203"></emphasis>
<taxonomicName id="178D4D7CFFEBFFACD5196272FDAB232C" authority="Martin, 1838" authorityName="Martin" authorityYear="1838" box="[376,607,1115,1139]" class="Mammalia" family="Tenrecidae" genus="Echinops" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD5196272FE3B232C" box="[376,463,1115,1139]" italics="true" pageId="19" pageNumber="203">Echinops</emphasis>
Martin, 1838
</taxonomicName>
(Setiferina) and
<taxonomicName id="178D4D7CFFEBFFACD4E06252FE9F23CC" authority="Lacepede, 1799" authorityName="Lacepede" authorityYear="1799" box="[129,363,1146,1171]" class="Mammalia" family="Tenrecidae" genus="Tenrec" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD4E06252FF4B23CD" box="[129,191,1147,1170]" italics="true" pageId="19" pageNumber="203">Tenrec</emphasis>
Lacépède, 1799
</taxonomicName>
<emphasis id="E2F9EAEDFFEBFFACD50A6252FE8C23CD" box="[363,376,1147,1170]" italics="true" pageId="19" pageNumber="203"></emphasis>
<taxonomicName id="178D4D7CFFEBFFACD5196252FD7323CC" authority="Mivart, 1871" authorityName="Mivart" authorityYear="1871" box="[376,647,1146,1171]" class="Mammalia" family="Tenrecidae" genus="Hemicentetes" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="genus">
<emphasis id="E2F9EAEDFFEBFFACD5196252FE0223CD" box="[376,502,1147,1170]" italics="true" pageId="19" pageNumber="203">Hemicentetes</emphasis>
Mivart, 1871
</taxonomicName>
(Tenrecina) and is collectively the sister taxon to all other extant Malagasy tenrecs in the Oryzorictinae (i.e., geogalins plus oryzorictins).
</paragraph>
<paragraph id="D03236FFFFEBFFACD4FD62F0FD1522F3" blockId="19.[129,778,607,1453]" pageId="19" pageNumber="203">
Our equally weighted parsimony analysis diverged from
<bibRefCitation id="B41C4B0EFFEBFFACD4E062D1FEA2224F" author="Everson KM &amp; Soarimalala V &amp; Goodman SM" box="[129,342,1272,1296]" pageId="19" pageNumber="203" pagination="890 - 909" refId="ref19590" refString="Everson KM, Soarimalala V, Goodman SM et al. Multiple loci and complete taxonomic sampling resolve the phylogeny and biogeographic history of tenrecs (Mammalia: Tenrecidae) and reveal higher speciation rates in Madagascar's humid forests. Systematic Biology 2016; 65: 890 - 909." type="journal article" year="2016">
Everson
<emphasis id="E2F9EAEDFFEBFFACD4BD62D0FEFA224F" box="[220,270,1272,1296]" italics="true" pageId="19" pageNumber="203">et al.</emphasis>
(2016
</bibRefCitation>
: fig. 3) in reconstructing
<taxonomicName id="178D4D7CFFEBFFACD60B62D0FD18224F" authorityName="Major" authorityYear="1896" box="[618,748,1272,1296]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="mergulus">
<emphasis id="E2F9EAEDFFEBFFACD60B62D0FD18224F" box="[618,748,1272,1296]" italics="true" pageId="19" pageNumber="203">Mi. mergulus</emphasis>
</taxonomicName>
as sister to a
<emphasis id="E2F9EAEDFFEBFFACD4866331FE2F2270" box="[231,475,1303,1327]" italics="true" pageId="19" pageNumber="203">
<taxonomicName id="178D4D7CFFEBFFACD4866331FEA82270" authorityName="Thomas" authorityYear="1884" box="[231,348,1303,1327]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="dobsoni">Mi. dobsoni</taxonomicName>
<taxonomicName id="178D4D7CFFEBFFACD50B633EFE2F2270" authorityName="Major" authorityYear="1896" box="[362,475,1303,1327]" class="Mammalia" family="Tenrecidae" genus="Microgale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="talazaci">Mi. talazaci</taxonomicName>
</emphasis>
clade and in placing
<taxonomicName id="178D4D7CFFEBFFACD6D06331FCFE2270" authorityName="Milne-Edwards &amp; Grandidier" authorityYear="1871" box="[689,778,1304,1327]" class="Mammalia" family="Tenrecidae" genus="Geogale" kingdom="Animalia" order="Afrosoricida" pageId="19" pageNumber="203" phylum="Chordata" rank="species" species="aurita">
<emphasis id="E2F9EAEDFFEBFFACD6D06331FCFE2270" box="[689,778,1304,1327]" italics="true" pageId="19" pageNumber="203">G. aurita</emphasis>
</taxonomicName>
as sister to Tenrecini rather than Oryzorictinae. We regard these signals as artefacts and instead draw our conclusions based on the Bayesian (
<figureCitation id="48B62A7AFFEBFFACD571635CFE9D22D2" box="[272,361,1397,1421]" captionStart="Figure 10" captionStartId="16.[113,178,1739,1763]" captionTargetBox="[116,1457,145,1708]" captionTargetId="graphics-7@16.[116,1457,145,1709]" captionTargetPageId="16" captionText="Figure 10. Topologies resulting from Bayesian-BIC (A, left) and parsimony implied weighting K = 4 (B, right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a) and indels.Bayesian branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are in red, tenrecids green.Circled and coloured letters adjacent to nodes indicate high-level taxonomy: Am, Amblysominae; Cd, Chrysochloridae; Cn, Chrysochlorinae; Cx, Chrysospalacinae; Gi, Geogalini; Mg, Microgale; Oi, Oryzorictini; Or, Oryzorictinae; Po, Potamogalinae; Sa, Setiferina; Ta, Tenrecina; Td, Tenrecidae; Tn, Tenrecinae; Ti, Tenrecini." figureDoi="http://doi.org/10.5281/zenodo.11241156" httpUri="https://zenodo.org/record/11241156/files/figure.png" pageId="19" pageNumber="203">Figs 10A</figureCitation>
,
<figureCitation id="48B62A7AFFEBFFACD514635CFE5422D2" box="[373,416,1397,1421]" captionStart="Figure 11" captionStartId="17.[129,194,1821,1845]" captionTargetBox="[131,1471,147,1792]" captionTargetId="graphics-8@17.[131,1472,147,1792]" captionTargetPageId="17" captionText="Figure 11. Topologies resulting from Bayesian-BIC (A; left) and parsimony implied weighting K = 4 (B; right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a), indels and morphology.Branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are red, tenrecids green. Circled and coloured letters adjacent to nodes indicate high-level taxonomy, as shown in Figure 10." figureDoi="http://doi.org/10.5281/zenodo.11241158" httpUri="https://zenodo.org/record/11241158/files/figure.png" pageId="19" pageNumber="203">11A</figureCitation>
,
<figureCitation id="48B62A7AFFEBFFACD5CA635CFE2222D2" box="[427,470,1397,1421]" captionStart="Figure 12" captionStartId="18.[113,174,1850,1874]" captionTargetBox="[127,1447,150,1829]" captionTargetId="graphics-7@18.[127,1446,161,1816]" captionTargetPageId="18" captionText="Figure 12.Topologies resultingfrom Bayesian-BIC (A; left)and parsimonyimplied weightingK = 4 (B;right) analyses of extanttaxasampled for six genes (cytB, ND2, GHR, vWF, 12S andstat5a)and indels,plus fossilsand morphologicaldata sampled forall.Bayesian branch lengthsreflect the scaleat the top left;parsimonybranch lengthsare arbitrary.Only chrysochloridsand Procavia are knownfor stat5a(andother regionsof missing data are shown in Table1). Bayesian nodes have a posterior probabilityof oneunlessindicated otherwise.Numbersadjacentto parsimonynodes represent bootstrap values,calculatedwith 250pseudoreplicates of a TNTsearchusing mult = tbr 10 replicateseach andreported onlyat orover50.Chrysochlorids are red, tenrecidsgreen;fossils are darkershadesof each.Circled and colouredlettersadjacent tonodes indicate high-leveltaxonomy,as shown in Figure10." figureDoi="http://doi.org/10.5281/zenodo.11241160" httpUri="https://zenodo.org/record/11241160/files/figure.png" pageId="19" pageNumber="203">12A</figureCitation>
; Supporting Information,
<figureCitation id="48B62A7AFFEBFFACD682635CFF6F22F2" captionStart="Figure 1" captionStartId="1.[129,194,1898,1922]" captionTargetBox="[150,1461,696,1851]" captionTargetId="figure-328@1.[129,1473,676,1871]" captionTargetPageId="1" captionText="Figure 1. Right forelimb and hand skeleton of Amblysomus corriae in dorsal view (ZM 42553).Roman numerals indicate digital rays. Abbreviations:dt, deltoid trough; ef, entepicondylar foramen; fp, flexor process; hh, humeral head; me, medial epicondyle; oft, ossified flexor tendon; op, olecranon process; rfc, radius flexor canal; sc, supinator crest. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11241138" httpUri="https://zenodo.org/record/11241138/files/figure.png" pageId="19" pageNumber="203">Fig. S1</figureCitation>
) and implied-weighting (
<figureCitation id="48B62A7AFFEBFFACD5C063BDFE0C22F2" box="[417,504,1428,1453]" captionStart="Figure 10" captionStartId="16.[113,178,1739,1763]" captionTargetBox="[116,1457,145,1708]" captionTargetId="graphics-7@16.[116,1457,145,1709]" captionTargetPageId="16" captionText="Figure 10. Topologies resulting from Bayesian-BIC (A, left) and parsimony implied weighting K = 4 (B, right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a) and indels.Bayesian branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are in red, tenrecids green.Circled and coloured letters adjacent to nodes indicate high-level taxonomy: Am, Amblysominae; Cd, Chrysochloridae; Cn, Chrysochlorinae; Cx, Chrysospalacinae; Gi, Geogalini; Mg, Microgale; Oi, Oryzorictini; Or, Oryzorictinae; Po, Potamogalinae; Sa, Setiferina; Ta, Tenrecina; Td, Tenrecidae; Tn, Tenrecinae; Ti, Tenrecini." figureDoi="http://doi.org/10.5281/zenodo.11241156" httpUri="https://zenodo.org/record/11241156/files/figure.png" pageId="19" pageNumber="203">Figs 10B</figureCitation>
,
<figureCitation id="48B62A7AFFEBFFACD66263BCFDD822F2" box="[515,556,1429,1453]" captionStart="Figure 11" captionStartId="17.[129,194,1821,1845]" captionTargetBox="[131,1471,147,1792]" captionTargetId="graphics-8@17.[131,1472,147,1792]" captionTargetPageId="17" captionText="Figure 11. Topologies resulting from Bayesian-BIC (A; left) and parsimony implied weighting K = 4 (B; right) analyses of extant taxa sampled for six genes (cytB, ND2, GHR, vWF, 12S and stat5a), indels and morphology.Branch lengths reflect the scale at the top left; parsimony branch lengths are arbitrary.Only chrysochlorids and Procavia are known for stat5a (for other missing loci, see Table 1). Bayesian nodes have a posterior probability of one unless indicated otherwise. Numbers adjacent to parsimony nodes represent bootstrap values, calculated with 500 pseudoreplicates of a TNT search using mult = tbr 100 replicates each and reported only at or over 50. Chrysochlorids are red, tenrecids green. Circled and coloured letters adjacent to nodes indicate high-level taxonomy, as shown in Figure 10." figureDoi="http://doi.org/10.5281/zenodo.11241158" httpUri="https://zenodo.org/record/11241158/files/figure.png" pageId="19" pageNumber="203">11B</figureCitation>
,
<figureCitation id="48B62A7AFFEBFFACD65663BCFD9422F2" box="[567,608,1429,1453]" captionStart="Figure 12" captionStartId="18.[113,174,1850,1874]" captionTargetBox="[127,1447,150,1829]" captionTargetId="graphics-7@18.[127,1446,161,1816]" captionTargetPageId="18" captionText="Figure 12.Topologies resultingfrom Bayesian-BIC (A; left)and parsimonyimplied weightingK = 4 (B;right) analyses of extanttaxasampled for six genes (cytB, ND2, GHR, vWF, 12S andstat5a)and indels,plus fossilsand morphologicaldata sampled forall.Bayesian branch lengthsreflect the scaleat the top left;parsimonybranch lengthsare arbitrary.Only chrysochloridsand Procavia are knownfor stat5a(andother regionsof missing data are shown in Table1). Bayesian nodes have a posterior probabilityof oneunlessindicated otherwise.Numbersadjacentto parsimonynodes represent bootstrap values,calculatedwith 250pseudoreplicates of a TNTsearchusing mult = tbr 10 replicateseach andreported onlyat orover50.Chrysochlorids are red, tenrecidsgreen;fossils are darkershadesof each.Circled and colouredlettersadjacent tonodes indicate high-leveltaxonomy,as shown in Figure10." figureDoi="http://doi.org/10.5281/zenodo.11241160" httpUri="https://zenodo.org/record/11241160/files/figure.png" pageId="19" pageNumber="203">12B</figureCitation>
) topologies.
</paragraph>
</subSubSection>
</treatment>
</document>