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<document id="FDBABD1AF364DAD62C6A70E108739F8C" ID-DOI="10.5733/afin.049.0207" ID-ISSN="2305-2562" ID-Zenodo-Dep="7917296" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683578241543" checkinUser="felipe" docAuthor="Barraclough, D. A." docDate="2008" docId="582487C3A80E844D17D31BABFE70B060" docLanguage="en" docName="AfrInvertebr.49.2.127-130.pdf" docOrigin="African Invertebrates 49 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.049.0207" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Afrocamilla Barraclough 1992" docType="treatment" docVersion="4" lastPageNumber="129" masterDocId="A41DFFBBA80E844F16481F4EFFCFB407" masterDocTitle="Modified fore femora in a new species of Afrocamilla Barraclough: a morphological innovation in the Camillidae (Diptera)" masterLastPageNumber="127" masterPageNumber="127" pageNumber="127" updateTime="1699499045202" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="61D2C3345304A9E8AB070E7777360344">Modified fore femora in a new species of Afrocamilla Barraclough: a morphological innovation in the Camillidae (Diptera)</mods:title>
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<mods:namePart id="06A80240018167A45CDA47F663CF45F1">Barraclough, D. A.</mods:namePart>
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<mods:date id="B25DB8DAF48DE5578A74D0947B99090A">2008</mods:date>
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<heading id="8B7A81B9A80E844F17D31BABFC8AB0F9" box="[411,837,1253,1278]" centered="true" fontSize="36" level="2" pageId="0" pageNumber="127" reason="6">
Genus
<taxonomicName id="178D4D56A80E844F17A11BABFC8AB0F9" ID-CoL="629PJ" ID-ENA="2952855" authority="Barraclough, 1992" authorityName="Barraclough" authorityYear="1992" box="[489,837,1253,1278]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="E2F9EAC7A80E844F17A11BABFDA1B0FA" box="[489,622,1253,1277]" italics="true" pageId="0" pageNumber="127">Afrocamilla</emphasis>
<bibRefCitation id="B41C4B24A80E844F143C1BABFC8AB0F9" author="BARRACLOUGH, D. A." box="[628,837,1253,1278]" pageId="0" pageNumber="127" pagination="37 - 49" refId="ref1748" refString="BARRACLOUGH, D. A. 1992. Afrocamilla stuckenbergi, a new African genus and species of Camillidae (Diptera), with comments on its behaviour and biology. Annals of the Natal Museum 33 (1): 37 - 49." type="journal article" year="1992">Barraclough, 1992</bibRefCitation>
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</paragraph>
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Type species:
<taxonomicName id="178D4D56A80E844F17671A43FC05B121" authority="Barraclough, 1992" authorityName="Barraclough" authorityYear="1992" box="[303,970,1293,1318]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="E2F9EAC7A80E844F17671A43FD86B122" box="[303,585,1293,1317]" italics="true" pageId="0" pageNumber="127">Afrocamilla stuckenbergi</emphasis>
<bibRefCitation id="B41C4B24A80E844F14191A43FCEDB121" author="BARRACLOUGH, D. A." box="[593,802,1293,1318]" pageId="0" pageNumber="127" pagination="37 - 49" refId="ref1748" refString="BARRACLOUGH, D. A. 1992. Afrocamilla stuckenbergi, a new African genus and species of Camillidae (Diptera), with comments on its behaviour and biology. Annals of the Natal Museum 33 (1): 37 - 49." type="journal article" year="1992">Barraclough, 1992</bibRefCitation>
(Afrotropical)
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="D03236D5A80E844F17851A29FCDDB187" blockId="0.[144,876,1384,1565]" box="[461,786,1383,1408]" pageId="0" pageNumber="127">
<taxonomicName id="178D4D56A80E844F17851A29FD1FB187" authority="Barraclough, 2008" authorityName="Barraclough" authorityYear="2008" box="[461,720,1383,1408]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="femorata" status="sp. nov.">
<emphasis id="E2F9EAC7A80E844F17851A29FD1FB187" bold="true" box="[461,720,1383,1408]" pageId="0" pageNumber="127">Afrocamilla femorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9CA57BCA80E844F149E1A29FCDDB187" box="[726,786,1383,1408]" pageId="0" pageNumber="127" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="D03236D5A80E844F14771AC1FD6FB1AF" blockId="0.[144,876,1384,1565]" box="[575,672,1423,1448]" pageId="0" pageNumber="127">
<figureCitation id="48B62A50A80E844F14771AC1FD6FB1AF" box="[575,672,1423,1448]" captionStart="Figs 1, 2" captionStartId="1.[182,221,1623,1643]" captionTargetBox="[208,1115,694,1602]" captionTargetId="figure-216@1.[208,1115,694,1602]" captionTargetPageId="1" captionText="Figs 1, 2. Detail of fore femur in Afrocamilla femorata sp. n.: (1)Anterior face showing position of cuticular protrusion with spine; (2) Slender, cone-like protrusion with stout dagger-like spine (same leg)." figureDoi="http://doi.org/10.5281/zenodo.7917298" httpUri="https://zenodo.org/record/7917298/files/figure.png" pageId="0" pageNumber="127">Figs 1, 2</figureCitation>
</paragraph>
<paragraph id="D03236D5A80E844F16D81AF9FCA3B1D7" blockId="0.[144,876,1384,1565]" box="[144,876,1463,1488]" pageId="0" pageNumber="127">Etymology: The species name refers to the modified fore femora.</paragraph>
<paragraph id="D03236D5A80E844F16D81A91FED4B1FF" blockId="0.[144,876,1384,1565]" box="[144,283,1503,1528]" pageId="0" pageNumber="127">Description:</paragraph>
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<emphasis id="E2F9EAC7A80E844F16D81946FF2BB227" box="[144,228,1544,1568]" italics="true" pageId="0" pageNumber="127">Female</emphasis>
.
<emphasis id="E2F9EAC7A80E844F16D8197EFE91B24F" box="[144,350,1584,1608]" italics="true" pageId="0" pageNumber="127">Colour/Pollinosity</emphasis>
: Head mainly brown to black, but anterior half of occiput (sometimes only slightly so), antenna (arista excepted) and part of gena yellow to yellow-brown; pollinosity silver to brown, relatively dense and mainly restricted to occiput. Thorax mainly dark brown, but humeral callus and adjacent cuticle (dorsally and ventrally) yellow-brown; pollinosity dense and brown on scutellum, more difficult to discern but visible along posteroventral margin of sternopleuron (silver) and between hypopleuron and scutellum (ranging from silver below to pale brown above). Legs entirely pale yellow. Wing membrane hyaline; transverse basal fascia not evident. Abdomen black with indistinct purple metallic reflections;
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2 with brown pollinosity.
</paragraph>
<paragraph id="D03236D5A80F844D16FD1EDAFC05B540" blockId="1.[181,1142,206,652]" lastBlockId="2.[144,1105,206,1127]" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
<emphasis id="E2F9EAC7A80F844E16FD1EDAFF3DB5AB" box="[181,242,404,428]" italics="true" pageId="1" pageNumber="128">Head</emphasis>
: Eye margin smoothly rounded anterodorsally in profile. Frons moderately short and broad in dorsal view, length at midline about 3/4 maximum width. Width of face and parafacials together, at mid-height, 0.8–1.3× length of antenna. Antenna with short ventral rays along apical 1/2 to 3/4 of arista, longest dorsal rays 0.8–1.0× length of third antennal segment. Upper orbital plate obviously developed anteriorly (ending coincident with anterior reclinate bristle), here falling short of ptilinal suture by about 1.5× length of ocellar triangle, maximum width (near vertex) 0.7–0.8× width of ocellar triangle. Postocellar bristles moderately developed, length about 0.9–1.2× that of ocellar triangle. Ocellar triangle moderately raised above upper eye margin in profile, distance 0.7– 0.8× length of third antennal segment. Proclinate fronto-orbital bristle absent, anterior reclinate fronto-orbital present, being 1/8 to 1/10 length of posterior bristle.Two vibrissae present. Gena height in profile 0.4–0.7× length of third antennal segment.
</paragraph>
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<paragraph id="D03236D5A80F844E16FE1919FBB8B284" blockId="1.[182,1143,1623,1667]" pageId="1" pageNumber="128">
Figs 1, 2. Detail of fore femur in
<taxonomicName id="178D4D56A80F844E17901919FD55B26C" authority="Barraclough, 2008" authorityName="Barraclough" authorityYear="2008" box="[472,666,1623,1643]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="femorata" status="sp. nov.">
<emphasis id="E2F9EAC7A80F844E17901919FD55B26C" box="[472,666,1623,1643]" italics="true" pageId="1" pageNumber="128">Afrocamilla femorata</emphasis>
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: (1)Anterior face showing position of cuticular protrusion with spine; (2) Slender, cone-like protrusion with stout dagger-like spine (same leg).
</paragraph>
</caption>
<paragraph id="D03236D5A80C844D16D81E1AFBD5B66B" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
<emphasis id="E2F9EAC7A80C844D16D81E1AFF2EB56B" box="[144,225,340,364]" italics="true" pageId="2" pageNumber="129">Thorax</emphasis>
: Apical scutellar marginal bristles well developed, 0.9–1.1× length of basal marginals. Fore femur markedly swollen relative to other femora (most prominently so along posterior face), maximum width and depth about twice that of other femora; slender, ventrally directed cone-like protrusion of tubercle evident posteroventrally at about mid-length or just beyond, protrusion 1/4 to 1/3 femur depth in length; apex of protrusion bearing a stout dagger-like spine (length 1/2 to 3/4 depth of femur), which is anterolaterally directed beneath and well beyond ventral margin of femur such that it is obvious beyond anterior femoral face in dorsal view (
<figureCitation id="48B62A50A80C844D154A1D7DFCA1B64B" box="[770,878,563,588]" captionStart="Figs 1, 2" captionStartId="1.[182,221,1623,1643]" captionTargetBox="[208,1115,694,1602]" captionTargetId="figure-216@1.[208,1115,694,1602]" captionTargetPageId="1" captionText="Figs 1, 2. Detail of fore femur in Afrocamilla femorata sp. n.: (1)Anterior face showing position of cuticular protrusion with spine; (2) Slender, cone-like protrusion with stout dagger-like spine (same leg)." figureDoi="http://doi.org/10.5281/zenodo.7917298" httpUri="https://zenodo.org/record/7917298/files/figure.png" pageId="2" pageNumber="129">Figs 1, 2</figureCitation>
). Mid tibia without dorsal pre-apical bristle; apicoventral bristle present. Wing not markedly slender.
</paragraph>
<paragraph id="D03236D5A80C844D16D81D37FCB2B6B0" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
<emphasis id="E2F9EAC7A80C844D16D81D37FF37B696" box="[144,248,633,657]" italics="true" pageId="2" pageNumber="129">Abdomen</emphasis>
:
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2 median marginal bristles moderately developed, 1.2–1.5× length of
<collectionCode id="B69CAE10A80C844D15B41D37FBC0B695" box="[1020,1039,633,658]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="129">T</collectionCode>
1+
<collectionCode id="B69CAE10A80C844D12621D37FBF2B695" box="[1066,1085,633,658]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="129">T</collectionCode>
2.
<emphasis id="E2F9EAC7A80C844D16D81DD0FEFDB6B1" box="[144,306,670,694]" italics="true" pageId="2" pageNumber="129">Measurements</emphasis>
(in mm): Body length 2.7–3.8, wing length 2.5–3.8.
</paragraph>
<paragraph id="D03236D5A80C844D16D81D8CFD08B6E9" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
<materialsCitation id="60E53C88A80C844D16D81D8CFD08B6E9" collectingDate="2000-02-02" collectionCode="YAROM" country="Ethiopia" county="Forest" elevation="2800" location="Bale" municipality="Harenna" pageId="2" pageNumber="129" specimenCount="1" typeStatus="holotype">
<typeStatus id="0F368877A80C844D16D81D8CFF27B6D1" box="[144,232,706,726]" pageId="2" pageNumber="129" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="A89A7645A80C844D16BB1D8CFE90B6D1" box="[243,351,706,726]" name="Ethiopia" pageId="2" pageNumber="129">ETHIOPIA</collectingCountry>
:
<emphasis id="E2F9EAC7A80C844D17231D8CFE46B6D1" box="[363,393,706,726]" italics="true" pageId="2" pageNumber="129">^“</emphasis>
<location id="D552600EA80C844D17C21D8CFE7AB6D1" LSID="urn:lsid:plazi:treatment:582487C3A80E844D17D31BABFE70B060:D552600EA80C844D17C21D8CFE7AB6D1" box="[394,437,706,726]" country="Ethiopia" county="Forest" municipality="Harenna" name="Bale" pageId="2" pageNumber="129">Bale</location>
/
<location id="D552600EA80C844D17841D8CFD90B6D1" LSID="urn:lsid:plazi:treatment:582487C3A80E844D17D31BABFE70B060:D552600EA80C844D17841D8CFD90B6D1" box="[460,607,706,726]" country="Ethiopia" county="Forest" municipality="Harenna" name="Bale Mountains" pageId="2" pageNumber="129">Bale Mountains</location>
/
<location id="D552600EA80C844D143F1D8CFD76B6D1" LSID="urn:lsid:plazi:treatment:582487C3A80E844D17D31BABFE70B060:D552600EA80C844D143F1D8CFD76B6D1" box="[631,697,706,726]" country="Ethiopia" county="Forest" municipality="Harenna" name="Harena" pageId="2" pageNumber="129">Harena</location>
[=
<collectingMunicipality id="3056ACAFA80C844D14961D8CFCE2B6D1" box="[734,813,706,726]" pageId="2" pageNumber="129">Harenna</collectingMunicipality>
]
<collectingCounty id="39534E59A80C844D15731D8CFCBBB6D1" box="[827,884,706,726]" pageId="2" pageNumber="129">Forest</collectingCounty>
/
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<elevation id="5BA0D1E6A80C844D15C31D8CFC04B6D1" box="[907,971,706,726]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="129" unit="m" value="2800.0">2800m</elevation>
</quantity>
<date id="A4331015A80C844D159B1D8CFBECB6D1" box="[979,1059,706,726]" pageId="2" pageNumber="129" value="2000-02-02">
<collectingDate id="B477E9FDA80C844D159B1D8CFBECB6D1" box="[979,1059,706,726]" pageId="2" pageNumber="129" value="2000-02-02">2.ii.2000</collectingDate>
</date>
/ A. FRIEDBERG / &amp; I.
<collectionCode id="B69CAE10A80C844D17011D94FE52B6E9" box="[329,413,730,750]" pageId="2" pageNumber="129">YAROM</collectionCode>
” (Tel Aviv University Museum).
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</paragraph>
<paragraph id="D03236D5A80C844D16D81DB4FEE6B721" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
Note: The Harenna Forest is an Afromontane ecosystem on the southern slopes of the Bale Mtns in southeastern
<collectingCountry id="A89A7645A80C844D169D1C5CFEEAB721" box="[213,293,786,806]" name="Ethiopia" pageId="2" pageNumber="129">Ethiopia</collectingCountry>
.
</paragraph>
<paragraph id="D03236D5A80C844D16D81C61FCB1B744" blockId="2.[144,1105,206,1127]" box="[144,894,815,835]" pageId="2" pageNumber="129">
<materialsCitation id="60E53C88A80C844D16D81C61FCB5B744" box="[144,890,815,835]" collectingDate="2000-02-02" collectionCode="YAROM" country="South Africa" county="Forest" elevation="2800" location="Natal Museum" municipality="Harenna" pageId="2" pageNumber="129" specimenCount="1" typeStatus="paratype">
<typeStatus id="0F368877A80C844D16D81C61FF23B744" box="[144,236,815,835]" pageId="2" pageNumber="129" type="paratype">Paratypes</typeStatus>
:
<emphasis id="E2F9EAC7A80C844D16BC1C61FE9FB744" box="[244,336,815,835]" italics="true" pageId="2" pageNumber="129">10^same</emphasis>
data as holotype, except one in
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,
<collectingCountry id="A89A7645A80C844D154C1C61FCB5B744" box="[772,890,815,835]" name="South Africa" pageId="2" pageNumber="129">South Africa</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="D03236D5A80C844D16D81C00FC52B7C0" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
Comparison:
<taxonomicName id="178D4D56A80C844D17601C00FE61B761" authorityName="Barraclough" authorityYear="2008" box="[296,430,846,870]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="E2F9EAC7A80C844D17601C00FE61B761" box="[296,430,846,870]" italics="true" pageId="2" pageNumber="129">A. femorata</emphasis>
</taxonomicName>
is immediately distinguished from all other congeners, and indeed all
<taxonomicName id="178D4D56A80C844D16B71C20FEB4B780" authorityName="Frey" authorityYear="1921" box="[255,379,878,903]" class="Insecta" family="Camillidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="family">Camillidae</taxonomicName>
, by the swollen fore femora and the armature thereon. Head profile and wing shape are unremarkable and are consequently not figured; reference should be made to the morphological detail presented in Barraclough (1997).
</paragraph>
<paragraph id="D03236D5A80C844D16E21C80FE70B060" blockId="2.[144,1105,206,1127]" pageId="2" pageNumber="129">
The absence of the transverse basal fascia on the wing is a noteworthy character state. In the well-studied South African fauna it is absent in only one of the nine described species (
<taxonomicName id="178D4D56A80C844D16BA1B40FD89B020" authority="Barraclough, 1997" authorityName="Barraclough" authorityYear="1997" box="[242,582,1038,1063]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="E2F9EAC7A80C844D16BA1B40FEA6B021" box="[242,361,1038,1062]" italics="true" pageId="2" pageNumber="129">A. confusa</emphasis>
Barraclough, 1997
</taxonomicName>
), although
<taxonomicName id="178D4D56A80C844D148D1B40FC85B021" authorityName="Barraclough" authorityYear="2008" box="[709,842,1038,1062]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="E2F9EAC7A80C844D148D1B40FC85B021" box="[709,842,1038,1062]" italics="true" pageId="2" pageNumber="129">A. femorata</emphasis>
</taxonomicName>
and
<taxonomicName id="178D4D56A80C844D15CA1B40FC35B021" authorityName="Barraclough" authorityYear="1997" box="[898,1018,1038,1062]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="E2F9EAC7A80C844D15CA1B40FC35B021" box="[898,1018,1038,1062]" italics="true" pageId="2" pageNumber="129">A. confusa</emphasis>
</taxonomicName>
are not closely related.
<taxonomicName id="178D4D56A80C844D17731B60FE61B041" authorityName="Barraclough" authorityYear="1997" box="[315,430,1070,1094]" class="Insecta" family="Camillidae" genus="Afrocamilla" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="E2F9EAC7A80C844D17731B60FE61B041" box="[315,430,1070,1094]" italics="true" pageId="2" pageNumber="129">A. confusa</emphasis>
</taxonomicName>
differs in having a proclinate fronto-orbital bristle and largely pallid thoracic colouration.
</paragraph>
</subSubSection>
</treatment>
</document>
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