233 lines
34 KiB
XML
233 lines
34 KiB
XML
<document id="1F49418BFBFBE89F6EB7F545B7A5FB0D" ID-DOI="10.11646/zootaxa.5450.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="11232500" ID-ZooBank="1B5D192F-1D5B-4460-9133-9AEAE9C920BF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281048833" checkinUser="plazi" docAuthor="Johnson, Robert G." docDate="2024" docId="03FD8227FF8AE376FF78F9A6FD6C85BC" docLanguage="en" docName="zootaxa.5450.1.1.pdf" docOrigin="Zootaxa 5450 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5450.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Pinnuloharpes chaperon Johnson 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="87" masterDocId="FFC4FA5FFFDFE320FFEFF87CFFB58771" masterDocTitle="Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco" masterLastPageNumber="185" masterPageNumber="1" pageNumber="86" updateTime="1716282690877" updateUser="ExternalLinkService">
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<mods:title id="71A9B77FB90090163B4D3E45341C21C0">Devonian Harpetidae from the central and eastern Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="AFDFEE4B31C821F690C2A6FDBCA66A2A">Johnson, Robert G.</mods:namePart>
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<mods:title id="AD5747EDFF38C67FDBFB6B7978A0FB25">Zootaxa</mods:title>
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<mods:part id="B8C2E51954A1E98FE55CA4DAE154D188">
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<mods:date id="5F6686228D14262F31D45987C7368C07">2024</mods:date>
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<mods:number id="E29295590A14F05DBD77D40C8BF13731">2024-05-15</mods:number>
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<mods:number id="174601078454F3BF71B54ACE08EE8181">5450</mods:number>
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<mods:url id="E010A708D666564AE3F3DCB9D4FE8D1F">http://dx.doi.org/10.11646/zootaxa.5450.1.1</mods:url>
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<mods:identifier id="01F56CF0FFD9333BAEE767AD0DFE2658" type="DOI">10.11646/zootaxa.5450.1.1</mods:identifier>
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<mods:identifier id="8B1EE7E391AFDF41C0AB6D2DF064DEB3" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03FD8227FF8AE376FF78F9A6FD6C85BC" ID-DOI="http://doi.org/10.5281/zenodo.11232630" ID-Zenodo-Dep="11232630" LSID="urn:lsid:plazi:treatment:03FD8227FF8AE376FF78F9A6FD6C85BC" httpUri="http://treatment.plazi.org/id/03FD8227FF8AE376FF78F9A6FD6C85BC" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86">
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<subSubSection id="C34E60BAFF8AE375FF78F9A6FDBB8684" box="[151,526,474,501]" pageId="85" pageNumber="86" type="nomenclature">
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<paragraph id="8BEB3331FF8AE375FF78F9A6FDBB8684" blockId="85.[151,526,474,537]" box="[151,526,474,501]" pageId="85" pageNumber="86">
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<heading id="D0A3845DFF8AE375FF78F9A6FDBB8684" bold="true" box="[151,526,474,501]" fontSize="11" level="2" pageId="85" pageNumber="86" reason="4">
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<emphasis id="B920EF23FF8AE375FF78F9A6FDBB8684" bold="true" box="[151,526,474,501]" pageId="85" pageNumber="86">
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<taxonomicName id="4C5448B2FF8AE375FF78F9A6FE718684" authority="Johnson, 2024" authorityName="Johnson" authorityYear="2024" box="[151,452,474,501]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="chaperon" status="sp. nov.">
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<emphasis id="B920EF23FF8AE375FF78F9A6FE718684" bold="true" box="[151,452,474,501]" italics="true" pageId="85" pageNumber="86">Pinnuloharpes chaperon</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A2135258FF8AE375FE24F9A7FDBB8684" box="[459,526,475,501]" pageId="85" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E60BAFF8AE375FF78F983FEAA8568" box="[151,287,511,537]" pageId="85" pageNumber="86" type="description">
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<paragraph id="8BEB3331FF8AE375FF78F983FEAA8568" blockId="85.[151,526,474,537]" box="[151,287,511,537]" pageId="85" pageNumber="86">
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<figureCitation id="136F2FB4FF8AE375FF78F983FEAA8568" box="[151,287,511,537]" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="85" pageNumber="86">Plate 41A–J</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E60BAFF8AE375FF78FA3AFE168464" pageId="85" pageNumber="86" type="diagnosis">
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<paragraph id="8BEB3331FF8AE375FF78FA3AFE168464" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
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<emphasis id="B920EF23FF8AE375FF78FA3AFEBF8511" bold="true" box="[151,266,582,608]" pageId="85" pageNumber="86">Diagnosis</emphasis>
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. Cephalon pyriform, strongly vaulted and widest at eye lobes. Glabella inset and joined to anterior boss. S1 not rising out of axial furrow. Genal areas are strongly convex (sag. and exsag.) and genal roll is steeply sloping, over
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<superScript id="7C219E79FF8AE375FF3FFAF3FF4185ED" attach="left" box="[208,244,654,681]" fontSize="6" pageId="85" pageNumber="86">600</superScript>
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anteriorly and over
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<superScript id="7C219E79FF8AE375FE36FAF3FE4885ED" attach="left" box="[473,509,654,681]" fontSize="6" pageId="85" pageNumber="86">700</superScript>
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laterally. Brim wide (sag.), 35% of cephalic length (sag.), barely sloping and brim width ratio very low, at 0.52. Perforations on brim very fine (<80 μm) and even finer on genal roll. Brim steepens quickly to near vertical on prolongations. In dorsal view, genal spine not following curve of exterior rim and curving upwards in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFF8AE375FF28FB63FB828448" box="[199,1079,799,825]" pageId="85" pageNumber="86" type="etymology">
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<paragraph id="8BEB3331FF8AE375FF28FB63FB828448" blockId="85.[151,1437,582,2013]" box="[199,1079,799,825]" pageId="85" pageNumber="86">
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<emphasis id="B920EF23FF8AE375FF28FB63FEFB8448" bold="true" box="[199,334,799,825]" pageId="85" pageNumber="86">Etymology.</emphasis>
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Similar to the shape of the chaperon hat worn in the Middle Ages.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E60BAFF8AE375FF28FB3EFC5D8360" pageId="85" pageNumber="86" type="materials_examined">
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<paragraph id="8BEB3331FF8AE375FF28FB3EFBAC84B8" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
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<emphasis id="B920EF23FF8AE375FF28FB3EFE49842C" bold="true" box="[199,508,834,861]" pageId="85" pageNumber="86">Material and occurrence.</emphasis>
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<materialsCitation id="3B3C396CFF8AE375FDE9FB3FFC1484F0" collectionCode="NHMUK" pageId="85" pageNumber="86" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8D93FF8AE375FDE9FB3FFDC6842C" box="[518,627,835,861]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED45ABF4FF8AE375FD6EFB3FFD5E842C" box="[641,747,835,861]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
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It 29332,
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<figureCitation id="136F2FB4FF8AE375FC87FB3FFC69842C" box="[872,988,835,861]" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="85" pageNumber="86">Pl. 41G–J</figureCitation>
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, from “
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<taxonomicName id="4C5448B2FF8AE375FBD6FB3FFA91842C" authority="Couche" authorityName="Couche" box="[1081,1316,835,861]" class="Trilobita" family="Lichidae" genus="Basseiarges" kingdom="Animalia" order="Lichida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
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<emphasis id="B920EF23FF8AE375FBD6FB3FFB0A842D" box="[1081,1215,835,860]" italics="true" pageId="85" pageNumber="86">Basseiarges</emphasis>
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Couche
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</taxonomicName>
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”, Eifelian, Fezna (
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<figureCitation id="136F2FB4FF8AE375FF04FB1BFEF084F0" box="[235,325,871,897]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="85" pageNumber="86">Fig. 2B</figureCitation>
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, Map 6, site 4), dorsal exoskeleton of cephalon only
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</materialsCitation>
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.
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<materialsCitation id="3B3C396CFF8AE375FC43FB1BFC2384D4" collectionCode="NHMUK" pageId="85" pageNumber="86" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54EF8D93FF8AE375FC43FB1BFBA884F0" box="[940,1053,871,897]" pageId="85" pageNumber="86" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED45ABF4FF8AE375FBC5FB1BFB2184F0" box="[1066,1172,871,897]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
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It 29331 (1 and 2),
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<figureCitation id="136F2FB4FF8AE375FA91FB1BFF5384D4" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="85" pageNumber="86">Pl. 41A–F</figureCitation>
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, from type locality and type horizon (
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<figureCitation id="136F2FB4FF8AE375FD7DFBF7FD5F84D4" box="[658,746,907,933]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="85" pageNumber="86">Fig. 2B</figureCitation>
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, Map 6, site 2)
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</materialsCitation>
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,
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<collectionCode id="ED45ABF4FF8AE375FC4CFBF7FBB884D4" box="[931,1037,907,933]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
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It 29331 (1), dorsal exoskeleton of cephalon only, and
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<collectionCode id="ED45ABF4FF8AE375FE9EFBD3FE6E84B8" box="[369,475,943,969]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
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It 29331 (2), ventral exoskeleton of cephalon only.
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</paragraph>
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<paragraph id="8BEB3331FF8AE375FF28FBAFFC5D8360" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
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Other material:
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<taxonomicName id="4C5448B2FF8AE375FE98FBAFFD1A849C" authorityName="Johnson" authorityYear="2024" box="[375,687,979,1005]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="chaperon">
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<emphasis id="B920EF23FF8AE375FE98FBAFFDA9849C" box="[375,540,979,1005]" italics="true" pageId="85" pageNumber="86">Pinnuloharpes</emphasis>
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cf.
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<emphasis id="B920EF23FF8AE375FDA9FBAFFD1A849C" box="[582,687,979,1005]" italics="true" pageId="85" pageNumber="86">chaperon</emphasis>
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</taxonomicName>
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:
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<collectionCode id="ED45ABF4FF8AE375FD53FBAFFC93849C" box="[700,806,979,1005]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
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It 29330,
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<figureCitation id="136F2FB4FF8AE375FC77FBAFFBA4849C" box="[920,1041,979,1005]" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="85" pageNumber="86">Pl. 41K–N</figureCitation>
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||
, from
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<taxonomicName id="4C5448B2FF8AE375FBB6FBAFFB68849C" authorityName="Johnson" authorityYear="2024" box="[1113,1245,979,1005]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="chaperon">
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<emphasis id="B920EF23FF8AE375FBB6FBAFFB68849C" box="[1113,1245,979,1005]" italics="true" pageId="85" pageNumber="86">P. chaperon</emphasis>
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</taxonomicName>
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<typeStatus id="54EF8D93FF8AE375FB0CFBAFFAA6849C" box="[1251,1299,979,1005]" pageId="85" pageNumber="86">type</typeStatus>
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||
locality and
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<typeStatus id="54EF8D93FF8AE375FF78FB8BFF728360" box="[151,199,1015,1041]" pageId="85" pageNumber="86">type</typeStatus>
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||
horizon (
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||
<figureCitation id="136F2FB4FF8AE375FEDCFB8BFE3E8360" box="[307,395,1015,1041]" captionStart="FIGURE 2" captionStartId="7.[152,255,1943,1967]" captionText="FIGURE 2A. Map locations in the Souss–Massa region" pageId="85" pageNumber="86">Fig. 2B</figureCitation>
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, Map 6, site 21), dorsal exoskeleton of cephalon only.
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C34E60BAFF8AE375FF28FC66FD0F8000" pageId="85" pageNumber="86" type="description">
|
||
<paragraph id="8BEB3331FF8AE375FF28FC66FBB68240" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
|
||
<emphasis id="B920EF23FF8AE375FF28FC66FEEC8345" bold="true" box="[199,345,1050,1076]" pageId="85" pageNumber="86">Description.</emphasis>
|
||
Cephalon widest at eye lobes and pyriform in shape. Genicranium narrow, width (tr.) 1.25 times length (sag.), strongly vaulted and widest at alae. Glabella inset, narrow, with rounded crest and tectiform flanks, no sculpture or pitting. S1 barely rising out of axial furrow. Axial furrows convex and weakly convergent anteriorly. Preglabellar furrow indistinct. Occipital furrow is broad, shallow and medially straight in dorsal view. Occipital ring same width (sag.) as posterior border (exsag.), and in lateral view, sloping posteriorly and same height as glabella anterior to S0. No sculpture or pitting on dorsal surface. Occipital node poorly inflated. Alae large (tr.), over 40% of width (tr.) of glabella at S0, just broader (tr.) than long (exsag.) and anterolaterally directed. Alar furrow shallow, well-defined. Posterior border moderately wide (tr.), and no preglabellar field.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF8AE375FF28FD47FCB9803C" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
|
||
Genal area moderately broad and without caeca, pits or tubercles. Eye lobes oval, large (exsag.), barely inflated and located away from inner margin of fringe. Each eye lobe with two lozenge-shaped eye lenses (
|
||
<figureCitation id="136F2FB4FF8AE375FAE7FD23FAD08208" box="[1288,1381,1375,1401]" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="85" pageNumber="86">Pl. 41D</figureCitation>
|
||
). No genal ridge, and eye ridge poorly defined. Inner margin of fringe not marked by bigger perforations as it crosses anterior boss. Anterior boss inflated, reaching down to weak girder kink and joined to glabella. Preaxial furrows converge. Genal roll steeply sloping
|
||
<superScript id="7C219E79FF8AE375FDD5FDB7FDEB82A9" attach="left" box="[570,606,1482,1509]" fontSize="6" pageId="85" pageNumber="86">620</superScript>
|
||
anteriorly, and over
|
||
<superScript id="7C219E79FF8AE375FCA2FDB7FCC482A9" attach="left" box="[845,881,1482,1509]" fontSize="6" pageId="85" pageNumber="86">720</superScript>
|
||
laterally, and in lateral view, narrowing strongly posteriorly. Perforations on genal roll are smaller than those on brim and no row of larger perforations above girder. Genal roll without caeca; girder moderately wide and brim wide (sag.), barely sloping and concave anteriorly, straight laterally. Brim width ratio low, at around 0.50. Brim perforations very fine, around 70µm, and no caeca on brim. Rim stout, dorsal surface not tilted outwards and without sculpture. Marginal band near vertical, straight, with tubercles but without ridges. Prolongations moderately short (exsag.), at around 80% of cephalic length (sag.). In lateral view, prolongations strongly tapered and internal rims concave. Internal rims subparallel in dorsal view, external rims curve adaxially posteriorly. Internal and external rims separated by a row of perforations from where they meet to end of prolongation, where they join. Genal spine long and set at an angle to curve of external rim. Tubercles on dorsal surface of internal rim, and extension of girder meeting internal rim 1/3 way down prolongation. Brims quickly steepen to near vertical on prolongations.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF8AE375FF28FF2BFD0F8000" blockId="85.[151,1437,582,2013]" box="[199,698,1879,1905]" pageId="85" pageNumber="86">Thorax, pygidium and hypostome unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E60BAFF8AE376FF28FF07FD6C85BC" lastPageId="86" lastPageNumber="87" pageId="85" pageNumber="86" type="discussion">
|
||
<paragraph id="8BEB3331FF8AE375FF28FF07FAEB80AC" blockId="85.[151,1437,582,2013]" pageId="85" pageNumber="86">
|
||
<emphasis id="B920EF23FF8AE375FF28FF07FE8E80E4" bold="true" box="[199,315,1915,1941]" pageId="85" pageNumber="86">Remarks.</emphasis>
|
||
<taxonomicName id="4C5448B2FF8AE375FEACFF07FDED80E4" authorityName="Johnson" authorityYear="2024" box="[323,600,1915,1941]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF8AE375FEACFF07FDED80E4" box="[323,600,1915,1941]" italics="true" pageId="85" pageNumber="86">Pinnuloharpes chaperon</emphasis>
|
||
</taxonomicName>
|
||
has cephalon that has a pyriform profile, a genicranium that is broadest at the alae rather than the posterior border and the course of the inner margin of the fringe cannot be determined from the material presented here, otherwise it has the other 8 diagnostic characters of the genus shown in
|
||
<tableCitation id="C6D6068AFF8AE375FB17FFBFFAEF80AC" box="[1272,1370,1986,2013]" captionStart="TABLE 5" captionStartId="182.[151,239,503,529]" captionText="TABLE 5b. Proposed diagnostic characters for Devonian harpetid genera (continued)" pageId="85" pageNumber="86">Table 5c</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF89E376FF28F8EBFD7A8568" blockId="86.[151,1437,150,717]" pageId="86" pageNumber="87">
|
||
<taxonomicName id="4C5448B2FF89E376FF28F8EBFE5587C0" authorityName="Johnson" authorityYear="2024" box="[199,480,151,177]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FF28F8EBFE5587C0" box="[199,480,151,177]" italics="true" pageId="86" pageNumber="87">Pinnuloharpes chaperon</emphasis>
|
||
</taxonomicName>
|
||
is quite similar to
|
||
<taxonomicName id="4C5448B2FF89E376FD2CF8EBFC5587C0" authority="(Lutke 1965)" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[707,992,150,177]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FD2CF8EBFC8A87C0" box="[707,831,151,177]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC54EC0FF89E376FCBEF8EBFC6D87C0" author="Lutke, F." box="[849,984,150,177]" pageId="86" pageNumber="87" pagination="151 - 236" refId="ref88811" refString="Lutke, F. (1965) Zur kenntnis herzynischer Trilobiten aus dem unter- und mittledevon des Harzes. Palaeontographica, Abteilung A, 124, 151 - 236." type="journal article" year="1965">Lütke 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but differs in having: a cephalon that is widest opposite eye lobes, not mid-point between the eye lobes and alae, and is more pyriform in shape; its genicranium widest at the alae and not the posterior border; the glabella inset and joined with the anterior boss; the axial furrows convex rather than straight; a wider occipital furrow; the occipital ring in lateral view the same height as the glabella anterior to S0, while that of
|
||
<taxonomicName id="4C5448B2FF89E376FD3DF95BFCF98630" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[722,844,295,321]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FD3DF95BFCF98630" box="[722,844,295,321]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
</taxonomicName>
|
||
is lower; a very shallow preglabellar furrow and no preglabellar field; alae with a width (tr.) 1.05 times their length (exsag.), while those of
|
||
<taxonomicName id="4C5448B2FF89E376FB80F937FB528614" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[1135,1255,331,357]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FB80F937FB528614" box="[1135,1255,331,357]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
</taxonomicName>
|
||
have a width of only around 0.75 times their length; a much broader (tr.) genal area, at 58% of its length (exsag.), rather than 47% as in
|
||
<taxonomicName id="4C5448B2FF89E376FF5CF9EFFE9F86DC" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[179,298,403,429]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FF5CF9EFFE9F86DC" box="[179,298,403,429]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
</taxonomicName>
|
||
; a wider (sag.) brim, being 36% of cephalic length (sag.), rather than 32% in
|
||
<taxonomicName id="4C5448B2FF89E376FB6FF9EFFB4D86DC" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[1152,1272,403,429]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FB6FF9EFFB4D86DC" box="[1152,1272,403,429]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
</taxonomicName>
|
||
; a much lower brim width ratio (0.49 rather than 0.76); shorter (exsag.) prolongations, which have a length (exsag.) that is 81% of cephalic length (sag.), compared with 89% in
|
||
<taxonomicName id="4C5448B2FF89E376FD78F9A7FCBB8684" baseAuthorityName="Lutke" baseAuthorityYear="1965" box="[663,782,475,501]" class="Trilobita" family="Harpetidae" genus="Harpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="hastatus">
|
||
<emphasis id="B920EF23FF89E376FD78F9A7FCBB8684" box="[663,782,475,501]" italics="true" pageId="86" pageNumber="87">P. hastatus</emphasis>
|
||
</taxonomicName>
|
||
; the extension of the brim on the prolongations steepening to near vertical, not just along their proximal edge.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF89E376FF28FA5FFBA385F4" blockId="86.[151,1437,150,717]" pageId="86" pageNumber="87">
|
||
<taxonomicName id="4C5448B2FF89E376FF28FA5FFE6B854C" authorityName="Johnson" authorityYear="2024" box="[199,478,547,573]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FF28FA5FFE6B854C" box="[199,478,547,573]" italics="true" pageId="86" pageNumber="87">Pinnuloharpes chaperon</emphasis>
|
||
</taxonomicName>
|
||
is different to other species of
|
||
<taxonomicName id="4C5448B2FF89E376FCA9FA5FFC5E854C" authorityName="Johnson" authorityYear="2024" box="[838,1003,547,573]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EF23FF89E376FCA9FA5FFC5E854C" box="[838,1003,547,573]" italics="true" pageId="86" pageNumber="87">Pinnuloharpes</emphasis>
|
||
</taxonomicName>
|
||
, in having a pyriform cephalon and a low brim width ratio. The pyriform cephalon is similar to the cephala of
|
||
<taxonomicName id="4C5448B2FF89E376FC51FA3BFB2A8510" box="[958,1183,583,609]" pageId="86" pageNumber="87">
|
||
<emphasis id="B920EF23FF89E376FC51FA3BFBF18510" box="[958,1092,583,609]" italics="true" pageId="86" pageNumber="87">Stoloharpes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, however, the brims of these species are sloping and convex, whereas that of
|
||
<taxonomicName id="4C5448B2FF89E376FD1CFA17FCC285F4" authorityName="Johnson" authorityYear="2024" box="[755,887,619,645]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FD1CFA17FCC285F4" box="[755,887,619,645]" italics="true" pageId="86" pageNumber="87">P. chaperon</emphasis>
|
||
</taxonomicName>
|
||
is almost flat.
|
||
</paragraph>
|
||
<paragraph id="8BEB3331FF89E376FF20FAF3FD6C85BC" blockId="86.[151,1437,150,717]" pageId="86" pageNumber="87">
|
||
<taxonomicName id="4C5448B2FF89E376FF20FAF3FDBF85D8" authorityName="Johnson" authorityYear="2024" box="[207,522,655,681]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" isUncertain="true" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FF20FAF3FEC185D8" box="[207,372,655,681]" italics="true" pageId="86" pageNumber="87">Pinnuloharpes</emphasis>
|
||
cf.
|
||
<emphasis id="B920EF23FF89E376FE4EFAF3FDBF85D8" box="[417,522,655,681]" italics="true" pageId="86" pageNumber="87">chaperon</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2FB4FF89E376FDF5FAF3FD2085D8" box="[538,661,655,681]" captionStart="PLATE 41" captionStartId="147.[151,234,1679,1703]" captionTargetBox="[236,1351,181,1655]" captionTargetId="figure-15@147.[236,1351,181,1655]" captionTargetPageId="147" captionText="PLATE 41. A–J Pinnuloharpes chaperon n. sp., “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. A–F exoskeleton of cephalon only, paratypes, NHMUK It 29331 (1 and 2), (Fig. 2B, Map 6, site 2). A–E NHMUK It 29331 (1). A, dorsal, B, lateral and C, anterior dorsal view. D, lateral view of eye lobe (uncoated) and E, dorsal view of brim perforations. F, NHMUK It 29331 (2), ventral exoskeleton of upper lamella. G–J dorsal exoskeleton of cephalon, holotype, NHMUK It 29332 (Fig. 2B, Map 6, site 4). G, dorsal, H, anterior and I, lateral view. J, dorsal view of brim perforations. K–N Pinnuloharpes cf. chaperon, “Basseiarges Couche”, El Otfal Formation, Eifelian, Fezna. Dorsal exoskeleton of partial cephalon only, NHMUK It 29330 (Map 2B, site 21). K, dorsal, L, anterior and M, lateral view. N, dorsal view of brim perforations. Scale bars shown represent 1 cm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.11232668" httpUri="https://zenodo.org/record/11232668/files/figure.png" pageId="86" pageNumber="87">Pl. 41K–N</figureCitation>
|
||
) differs from
|
||
<taxonomicName id="4C5448B2FF89E376FCDCFAF3FC0C85D8" authorityName="Johnson" authorityYear="2024" box="[819,953,655,681]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FCDCFAF3FC0C85D8" box="[819,953,655,681]" italics="true" pageId="86" pageNumber="87">P. chaperon</emphasis>
|
||
</taxonomicName>
|
||
in that its cephalon is not pyriform and its prolongations are shorter than those of
|
||
<taxonomicName id="4C5448B2FF89E376FDA1FACFFD6785BC" authorityName="Johnson" authorityYear="2024" box="[590,722,691,717]" class="Trilobita" family="Harpetidae" genus="Pinnuloharpes" kingdom="Animalia" order="Harpetida" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="chaperon">
|
||
<emphasis id="B920EF23FF89E376FDA1FACFFD6785BC" box="[590,722,691,717]" italics="true" pageId="86" pageNumber="87">P. chaperon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |