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<mods:title id="5CAC7FC2E5ADE14BCE5485C5F5AE8E95">A striking sexually dimorphic new species of Castolus (Hemiptera: Heteroptera Reduviidae) from Colombia, with new records from Neotropical countries and taxonomic notes on the genus</mods:title>
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<mods:namePart id="9BEDC962FD36CD00852ED0AFF1BD19D3">Forero, Dimitri</mods:namePart>
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<mods:affiliation id="86E95E30906E1D297B8BEBD9E9B21852">Laboratorio de Entomología, UNESIS, Departamento de Biología, Pontificia Universidad Javeriana, Bogotá, Colombia. forero-i @ javeriana. edu. co; https: // orcid. org / 0000 - 0002 - 6358 - 757 X</mods:affiliation>
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<mods:namePart id="A6CBCB568F69E9DA57036E4C3F0ED744">Mejía-Soto, Andrés</mods:namePart>
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<mods:affiliation id="AD8F3DE84E73404BA7C903A1DF7E2DA7">Grupo de Entomología Universidad de Antioquia (GEUA), Universidad de Antioquia, Medellín, Colombia. andres. mejia 6 @ udea. edu. co; https: // orcid. org / 0000 - 0001 - 7897 - 6183</mods:affiliation>
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<treatment id="52663A498F19FFF3FF30F911FE860CB8" ID-DOI="http://doi.org/10.5281/zenodo.5570418" ID-GBIF-Taxon="188639457" ID-Zenodo-Dep="5570418" LSID="urn:lsid:plazi:treatment:52663A498F19FFF3FF30F911FE860CB8" httpUri="http://treatment.plazi.org/id/52663A498F19FFF3FF30F911FE860CB8" lastPageId="12" lastPageNumber="550" pageId="10" pageNumber="548">
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<paragraph id="DA708B5F8F19FFF5FF30F911FDD10A12" blockId="10.[151,586,1770,1833]" box="[151,586,1770,1797]" pageId="10" pageNumber="548">
<heading id="81383C338F19FFF5FF30F911FDD10A12" bold="true" box="[151,586,1770,1797]" fontSize="11" level="1" pageId="10" pageNumber="548" reason="1">
<taxonomicName id="1DCFF0DC8F19FFF5FF30F911FDD10A12" ID-CoL="RQPG" authority="Breddin, 1904" authorityName="Breddin" authorityYear="1904" box="[151,586,1770,1797]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="548" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E8BB574D8F19FFF5FF30F911FDD10A12" bold="true" box="[151,586,1770,1797]" pageId="10" pageNumber="548">
<emphasis id="E8BB574D8F19FFF5FF30F911FE080A12" bold="true" box="[151,403,1770,1797]" italics="true" pageId="10" pageNumber="548">Castolus nigriventris</emphasis>
<bibRefCitation id="BE5EF6AE8F19FFF5FE3DF911FDD10A12" author="Breddin, G." box="[410,586,1770,1796]" pageId="10" pageNumber="548" pagination="147 - 148" refId="ref15054" refString="Breddin, G. (1904) Neue Rhynchotenausbeute aus Sud-Amerika. Societas Entomologica, 20, 147 - 148 + 153 - 154 + 177 - 178. https: // www. biodiversitylibrary. org / page / 9279381" type="journal article" year="1904">Breddin, 1904</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="DA708B5F8F19FFF5FF30F8F5FF640A3F" blockId="10.[151,586,1770,1833]" box="[151,255,1806,1833]" pageId="10" pageNumber="548">
(
<figureCitation id="42F497DA8F19FFF5FF07F8F5FF630A3F" box="[160,248,1806,1833]" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[166,1435,309,1487]" captionTargetId="figure-84@11.[151,1435,309,1520]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal view of Castolus type specimens. A. holotype of Castolus bicolor Maldonado, 1976 (synonym of C. nigriventris Breddin, 1904); B. holotype of C. bolivari Brailovsky, 1982; C. holotype labels of C. bicolor Maldonado, 1976." figureDoi="http://doi.org/10.5281/zenodo.5556788" httpUri="https://zenodo.org/record/5556788/files/figure.png" pageId="10" pageNumber="548">Fig. 6A</figureCitation>
)
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<paragraph id="DA708B5F8F19FFF5FF30F8A8FD050A7D" blockId="10.[151,670,1875,1899]" box="[151,670,1875,1899]" pageId="10" pageNumber="548">
<taxonomicName id="1DCFF0DC8F19FFF5FF30F8A8FE600A7D" ID-CoL="5XF3P" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[151,507,1875,1899]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="548" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F19FFF5FF30F8A8FEA40A7D" box="[151,319,1875,1899]" italics="true" pageId="10" pageNumber="548">Castolus bicolor</emphasis>
<bibRefCitation id="BE5EF6AE8F19FFF5FEE1F8A8FE600A7D" author="Maldonado, J." box="[326,507,1875,1899]" pageId="10" pageNumber="548" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
,
<emphasis id="E8BB574D8F19FFF5FDA2F8A8FD050A7D" bold="true" box="[517,670,1875,1899]" pageId="10" pageNumber="548">
<taxonomicNameLabel id="F388EA368F19FFF5FDA2F8A8FD010A7D" box="[517,666,1875,1899]" pageId="10" pageNumber="548">new synonym</taxonomicNameLabel>
.
</emphasis>
</paragraph>
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<subSubSection id="92D5D8D48F19FFF3FF30F862FC110DEF" lastPageId="12" lastPageNumber="550" pageId="10" pageNumber="548" type="discussion">
<paragraph id="DA708B5F8F19FFF4FF30F862FD9E0C0B" blockId="10.[151,1437,1945,2043]" lastBlockId="11.[151,1437,151,285]" lastPageId="11" lastPageNumber="549" pageId="10" pageNumber="548">
<emphasis id="E8BB574D8F19FFF5FF30F862FE900AA5" bold="true" box="[151,267,1945,1971]" pageId="10" pageNumber="548">Remarks.</emphasis>
<bibRefCitation id="BE5EF6AE8F19FFF5FEB5F862FE790AA2" author="Maldonado, J." box="[274,482,1945,1972]" pageId="10" pageNumber="548" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado (1976)</bibRefCitation>
did not include
<taxonomicName id="1DCFF0DC8F19FFF5FD31F862FC7B0AA5" authority="Breddin, 1904" authorityName="Breddin" authorityYear="1904" box="[662,992,1945,1971]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="548" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E8BB574D8F19FFF5FD31F862FCA10AA5" box="[662,826,1945,1971]" italics="true" pageId="10" pageNumber="548">C. nigriventris</emphasis>
<bibRefCitation id="BE5EF6AE8F19FFF5FCE7F862FC7B0AA5" author="Breddin, G." box="[832,992,1945,1971]" pageId="10" pageNumber="548" pagination="147 - 148" refId="ref15054" refString="Breddin, G. (1904) Neue Rhynchotenausbeute aus Sud-Amerika. Societas Entomologica, 20, 147 - 148 + 153 - 154 + 177 - 178. https: // www. biodiversitylibrary. org / page / 9279381" type="journal article" year="1904">
Breddin,
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in his key or in his taxon treatment section. He stated that Breddins
<typeStatus id="057435FD8F19FFF5FE4FF846FDD10AC1" box="[488,586,1981,2007]" pageId="10" pageNumber="548" type="holotype">holotype</typeStatus>
seems to be lost, and apparently was not at the DEI. We recently contacted Dr. S. Blank (DEI) and he confirmed that Breddins type is not there. We also contacted Dr. M. Husemann (ZMUH) who stated as well that Breddins type is not there, so apparently Breddins type could be lost. Alternatively, Dr. Blank indicated that it might still be possible for Breddins type to be unlabeled as a type and be sitting along his undetermined material, as has been the case with other Breddin type material. Unfortunately, so far, we have not been able to find such specimen.
</paragraph>
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<paragraph id="DA708B5F8F18FFF4FF30F9FCFADE0B55" blockId="11.[151,1436,1543,1603]" pageId="11" pageNumber="549">
<emphasis id="E8BB574D8F18FFF4FF30F9FCFE890B09" bold="true" box="[151,274,1543,1567]" pageId="11" pageNumber="549">FIGURE 6.</emphasis>
Dorsal view of
<taxonomicName id="1DCFF0DC8F18FFF4FE14F9FCFD970B09" authorityName="Stal" authorityYear="1858" box="[435,524,1543,1567]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="E8BB574D8F18FFF4FE14F9FCFD970B09" box="[435,524,1543,1567]" italics="true" pageId="11" pageNumber="549">Castolus</emphasis>
</taxonomicName>
type specimens.
<emphasis id="E8BB574D8F18FFF4FD1FF9FCFD4A0B09" bold="true" box="[696,721,1543,1567]" pageId="11" pageNumber="549">A.</emphasis>
holotype of
<taxonomicName id="1DCFF0DC8F18FFF4FCF7F9FCFB340B09" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[848,1199,1543,1567]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F18FFF4FCF7F9FCFC6C0B09" box="[848,1015,1543,1567]" italics="true" pageId="11" pageNumber="549">Castolus bicolor</emphasis>
<bibRefCitation id="BE5EF6AE8F18FFF4FC5AF9FCFB340B09" author="Maldonado, J." box="[1021,1199,1543,1567]" pageId="11" pageNumber="549" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
(synonym of
<taxonomicName id="1DCFF0DC8F18FFF4FA9DF9F3FEF60B55" authority="Breddin, 1904" authorityName="Breddin" authorityYear="1904" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E8BB574D8F18FFF4FA9DF9F3FF4A0B55" italics="true" pageId="11" pageNumber="549">C. nigriventris</emphasis>
<bibRefCitation id="BE5EF6AE8F18FFF4FF70F9D0FEF60B55" author="Breddin, G." box="[215,365,1579,1603]" pageId="11" pageNumber="549" pagination="147 - 148" refId="ref15054" refString="Breddin, G. (1904) Neue Rhynchotenausbeute aus Sud-Amerika. Societas Entomologica, 20, 147 - 148 + 153 - 154 + 177 - 178. https: // www. biodiversitylibrary. org / page / 9279381" type="journal article" year="1904">Breddin, 1904</bibRefCitation>
</taxonomicName>
);
<emphasis id="E8BB574D8F18FFF4FED9F9D0FE0E0B55" bold="true" box="[382,405,1579,1603]" pageId="11" pageNumber="549">B.</emphasis>
holotype of
<taxonomicName id="1DCFF0DC8F18FFF4FDB1F9D7FCA10B55" authority="Brailovsky, 1982" authorityName="Brailovsky" authorityYear="1982" box="[534,826,1579,1603]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis id="E8BB574D8F18FFF4FDB1F9D7FD1F0B55" box="[534,644,1579,1603]" italics="true" pageId="11" pageNumber="549">C. bolivari</emphasis>
<bibRefCitation id="BE5EF6AE8F18FFF4FD2DF9D0FCA10B55" author="Brailovsky, H." box="[650,826,1579,1603]" pageId="11" pageNumber="549" pagination="223 - 229" refId="ref15001" refString="Brailovsky, H. (1982) Hemiptera-Heteroptera de Mexico: XX. Notas acerca del genero Castolus Stal, y descripcion de una nueva especie. (Reduviidae-Harpactorinae). Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico, Serie Zoologia, 1, 223 - 229." type="journal article" year="1982">Brailovsky, 1982</bibRefCitation>
</taxonomicName>
;
<emphasis id="E8BB574D8F18FFF4FCE3F9D0FCC60B55" bold="true" box="[836,861,1579,1603]" pageId="11" pageNumber="549">C.</emphasis>
holotype labels of
<taxonomicName id="1DCFF0DC8F18FFF4FBB8F9D7FAA40B55" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[1055,1343,1579,1603]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F18FFF4FBB8F9D7FB1D0B55" box="[1055,1158,1579,1603]" italics="true" pageId="11" pageNumber="549">C. bicolor</emphasis>
<bibRefCitation id="BE5EF6AE8F18FFF4FB2BF9D0FAA40B55" author="Maldonado, J." box="[1164,1343,1579,1603]" pageId="11" pageNumber="549" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="DA708B5F8F18FFF4FF60F98AFD620ABD" blockId="11.[151,1437,1649,2036]" pageId="11" pageNumber="549">
<bibRefCitation id="BE5EF6AE8F18FFF4FF60F98AFE0C0B9A" author="Maldonado, J." box="[199,407,1649,1676]" pageId="11" pageNumber="549" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado (1976)</bibRefCitation>
indicated that Breddins description was insufficient to identify this species, but we disagree.
<bibRefCitation id="BE5EF6AE8F18FFF4FF30F96EFEDE0BA6" author="Breddin, G." box="[151,325,1685,1712]" pageId="11" pageNumber="549" pagination="147 - 148" refId="ref15054" refString="Breddin, G. (1904) Neue Rhynchotenausbeute aus Sud-Amerika. Societas Entomologica, 20, 147 - 148 + 153 - 154 + 177 - 178. https: // www. biodiversitylibrary. org / page / 9279381" type="journal article" year="1904">Breddin (1904)</bibRefCitation>
described
<taxonomicName id="1DCFF0DC8F18FFF4FE64F96DFDF20BB9" authorityName="Breddin" authorityYear="1904" box="[451,617,1686,1712]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E8BB574D8F18FFF4FE64F96DFDF20BB9" box="[451,617,1686,1712]" italics="true" pageId="11" pageNumber="549">C. nigriventris</emphasis>
</taxonomicName>
as follows: “...a dull black, broad animal, in which only a longitudinal band of the posterior lobe of the pronotum, along with its broad posterior margin and the slender spicules of the humeral angles, as well as the hemelytra, and a very narrow margin of the abdomen [connexivum], are somewhat dull light yellow.” Following Breddins description it is clear that this particular and striking color pattern, being mostly black with yellow hemelytra and black and yellow posterior pronotal lobe, is exactly the same as the color pattern described for
<taxonomicName id="1DCFF0DC8F18FFF4FE2FF8B1FD190A75" authority="Maldonado" authorityName="Maldonado" authorityYear="1976" box="[392,642,1865,1892]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F18FFF4FE2FF8B1FE610A75" box="[392,506,1865,1892]" italics="true" pageId="11" pageNumber="549">C. bicolor</emphasis>
Maldonado
</taxonomicName>
(
<figureCitation id="42F497DA8F18FFF4FD35F8B2FD770A72" box="[658,748,1865,1892]" captionStart="FIGURE 6" captionStartId="11.[151,250,1543,1567]" captionTargetBox="[166,1435,309,1487]" captionTargetId="figure-84@11.[151,1435,309,1520]" captionTargetPageId="11" captionText="FIGURE 6. Dorsal view of Castolus type specimens. A. holotype of Castolus bicolor Maldonado, 1976 (synonym of C. nigriventris Breddin, 1904); B. holotype of C. bolivari Brailovsky, 1982; C. holotype labels of C. bicolor Maldonado, 1976." figureDoi="http://doi.org/10.5281/zenodo.5556788" httpUri="https://zenodo.org/record/5556788/files/figure.png" pageId="11" pageNumber="549">Fig. 6A</figureCitation>
). The only difference is that the pale areas in the
<typeStatus id="057435FD8F18FFF4FABCF8B2FAE60A75" box="[1307,1405,1865,1891]" pageId="11" pageNumber="549" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1DCFF0DC8F18FFF4FF30F895FE920A91" authorityName="Maldonado" authorityYear="1976" box="[151,265,1901,1928]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="549" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F18FFF4FF30F895FE920A91" box="[151,265,1901,1928]" italics="true" pageId="11" pageNumber="549">C. bicolor</emphasis>
</taxonomicName>
are rather orange than yellow. Therefore, because of the same coloration patter between the two species, we synonymize Maldonados species under Breddins.
</paragraph>
<paragraph id="DA708B5F8F18FFF3FF60F84EFC110DEF" blockId="11.[151,1437,1649,2036]" lastBlockId="12.[151,1437,151,430]" lastPageId="12" lastPageNumber="550" pageId="11" pageNumber="549">
<bibRefCitation id="BE5EF6AE8F18FFF4FF60F84EFE020AC6" author="Maldonado, J." box="[199,409,1973,2000]" pageId="11" pageNumber="549" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado (1976)</bibRefCitation>
also offered an intriguing observation, he mentioned that he had examined a female specimen from
<collectingCountry id="A2D8CBCF8F18FFF4FEAAF822FEF70AE5" box="[269,364,2009,2035]" name="Trinidad and Tobago" pageId="11" pageNumber="549">Trinidad</collectingCountry>
with less extensive black areas on the pronotum, but otherwise very similar to the
<typeStatus id="057435FD8F18FFF4FABCF822FAE60AE5" box="[1307,1405,2009,2035]" pageId="11" pageNumber="549" type="holotype">holotype</typeStatus>
of
<taxonomicName id="1DCFF0DC8F1FFFF3FF30FF6CFE910DA7" authorityName="Maldonado" authorityYear="1976" box="[151,266,151,177]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F1FFFF3FF30FF6CFE910DA7" box="[151,266,151,177]" italics="true" pageId="12" pageNumber="550">C. bicolor</emphasis>
</taxonomicName>
. He hypothesized that this might be the female of the specimen he described. Unfortunately, he did not illustrate this specimen. Similar to what happens in
<taxonomicName id="1DCFF0DC8F1FFFF3FD70FF40FCCB0DC3" authorityName="Maldonado" authorityYear="1976" box="[727,848,187,213]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="E8BB574D8F1FFFF3FD70FF40FCCB0DC3" box="[727,848,187,213]" italics="true" pageId="12" pageNumber="550">C. lineatus</emphasis>
</taxonomicName>
and other species, some color variation might occur, with an extreme sexually dimorphic condition in
<taxonomicName id="1DCFF0DC8F1FFFF3FD19FF24FCB10DEF" authority="Forero &amp; Mejía-Soto, 2021" authorityName="Forero &amp; Mejía-Soto" authorityYear="2021" box="[702,810,223,249]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="rafaeli" status="sp. nov.">
<emphasis id="E8BB574D8F1FFFF3FD19FF24FCB10DEF" box="[702,810,223,249]" italics="true" pageId="12" pageNumber="550">C. rafaeli</emphasis>
</taxonomicName>
<emphasis id="E8BB574D8F1FFFF3FC96FF24FC110DEF" bold="true" box="[817,906,223,249]" pageId="12" pageNumber="550">
<taxonomicNameLabel id="F388EA368F1FFFF3FC96FF24FC110DEF" box="[817,906,223,249]" pageId="12" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="92D5D8D48F1FFFF3FF60FEF9FA830C73" pageId="12" pageNumber="550" type="materials_examined">
<paragraph id="DA708B5F8F1FFFF3FF60FEF9FA830C73" blockId="12.[151,1437,151,430]" pageId="12" pageNumber="550">
<emphasis id="E8BB574D8F1FFFF3FF60FEF9FE310C0A" bold="true" box="[199,426,258,284]" pageId="12" pageNumber="550">Material examined</emphasis>
.
<emphasis id="E8BB574D8F1FFFF3FE1EFEF9FCCF0C0A" bold="true" box="[441,852,258,285]" pageId="12" pageNumber="550">
<taxonomicName id="1DCFF0DC8F1FFFF3FE1EFEF9FCD40C0A" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[441,847,258,285]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F1FFFF3FE1EFEF9FDED0C0B" bold="true" box="[441,630,258,285]" italics="true" pageId="12" pageNumber="550">Castolus bicolor</emphasis>
<bibRefCitation id="BE5EF6AE8F1FFFF3FDDAFEF8FCD40C0A" author="Maldonado, J." box="[637,847,258,285]" pageId="12" pageNumber="550" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
.
</emphasis>
<materialsCitation id="6AA781028F1FFFF3FCFBFEF9FB750C54" ID-GBIF-Occurrence="3383561395" collectingDate="1966-02" collectorName="J. G. Rozen" country="Trinidad and Tobago" elevation="305" location="Valley" pageId="12" pageNumber="550" specimenCount="1" specimenCount-male="1" stateProvince="Arima" typeStatus="holotype">
<emphasis id="E8BB574D8F1FFFF3FCFBFEF9FB010C0A" bold="true" box="[860,1178,258,284]" pageId="12" pageNumber="550">
<typeStatus id="057435FD8F1FFFF3FCFBFEF9FC640C0A" box="[860,1023,258,284]" pageId="12" pageNumber="550" type="holotype">HOLOTYPE</typeStatus>
:
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</emphasis>
[
<emphasis id="E8BB574D8F1FFFF3FB0DFEF9FAF30C0A" bold="true" box="[1194,1384,258,284]" pageId="12" pageNumber="550">
AND
<collectingCountry id="A2D8CBCF8F1FFFF3FB49FEF9FAF30C0A" box="[1262,1384,258,284]" name="Trinidad and Tobago" pageId="12" pageNumber="550">TOBAGO</collectingCountry>
</emphasis>
]—
<specimenCount id="CCC940D68F1FFFF3FA34FEF9FF340C57" pageId="12" pageNumber="550" type="male">1 ♂</specimenCount>
; W.I. [West Indies],
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<location id="DF10DD848F1FFFF3FE49FEDCFDA20C57" LSID="urn:lsid:plazi:treatment:52663A498F19FFF3FF30F911FE860CB8:DF10DD848F1FFFF3FE49FEDCFDA20C57" box="[494,569,295,321]" country="Trinidad and Tobago" name="Valley" pageId="12" pageNumber="550" stateProvince="Arima">Valley</location>
;
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<elevation id="51E26C6C8F1FFFF3FDE0FEDCFD470C57" box="[583,732,295,321]" metricMagnitude="2" metricUnit="m" metricValue="3.048" metricValueMax="3.6576" metricValueMin="2.4384" pageId="12" pageNumber="550" unit="ft" value="1000.0" valueMax="1200.0" valueMin="800.0">8001200 ft.</elevation>
</quantity>
;
<date id="AE71AD9F8F1FFFF3FD4DFEDCFCC20C57" box="[746,857,295,321]" pageId="12" pageNumber="550" value="1966-02">
<collectingDate id="BE3554778F1FFFF3FD4DFEDCFCC20C57" box="[746,857,295,321]" pageId="12" pageNumber="550" value="1966-02">Feb 1966</collectingDate>
</date>
;
<collectorName id="773AEE898F1FFFF3FCCFFEDCFC6B0C57" box="[872,1008,295,321]" pageId="12" pageNumber="550">J. G. Rozen</collectorName>
collector / (red label)
</materialsCitation>
<materialsCitation id="6AA781028F1FFFF3FB50FEDCFD890C70" ID-GBIF-Occurrence="3383561317" collectionCode="AMNH" country="Trinidad and Tobago" pageId="12" pageNumber="550" specimenCode="IZC 00343371" specimenCount="1" typeStatus="holotype">
<typeStatus id="057435FD8F1FFFF3FB50FEDCFA100C57" box="[1271,1419,295,321]" pageId="12" pageNumber="550" type="holotype">HOLOTYPE</typeStatus>
/
<collectionCode id="BCDE139A8F1FFFF3FF30FEB0FF760C73" box="[151,237,331,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="12" pageNumber="550">AMNH</collectionCode>
_
<specimenCode id="8A6923248F1FFFF3FF5CFEB0FE040C73" box="[251,415,331,357]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="12" pageNumber="550">IZC 00343371</specimenCode>
(barcode)
</materialsCitation>
/
<materialsCitation id="6AA781028F1FFFF3FD80FEB0FA8F0C73" ID-GBIF-Occurrence="3383561314" box="[551,1300,330,357]" collectionCode="AMNH" collectorName="J Maldonado C" country="Trinidad and Tobago" determinerName="Maldonado" pageId="12" pageNumber="550" specimenCount="1" typeStatus="holotype">
<taxonomicName id="1DCFF0DC8F1FFFF3FD94FEB0FD760C73" authorityName="Maldonado" authorityYear="1976" box="[563,749,331,357]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="E8BB574D8F1FFFF3FD94FEB0FD760C73" box="[563,749,331,357]" italics="true" pageId="12" pageNumber="550">Castolus bicolor</emphasis>
</taxonomicName>
det.
<determinerName id="2FAC679B8F1FFFF3FC83FEB0FC3C0C73" box="[804,935,331,357]" pageId="12" pageNumber="550">Maldonado</determinerName>
<collectorName id="773AEE898F1FFFF3FC10FEB0FBF80C73" box="[951,1123,331,357]" pageId="12" pageNumber="550">J Maldonado C</collectorName>
1975 (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="92D5D8D48F1FFFF3FF60FE95FE860CB8" pageId="12" pageNumber="550" type="distribution">
<paragraph id="DA708B5F8F1FFFF3FF60FE95FE860CB8" blockId="12.[151,1437,151,430]" pageId="12" pageNumber="550">
<emphasis id="E8BB574D8F1FFFF3FF60FE95FEC20C9E" bold="true" box="[199,345,366,392]" pageId="12" pageNumber="550">Distribution</emphasis>
:
<taxonomicName id="1DCFF0DC8F1FFFF3FECEFE94FDCC0C9E" authorityName="Breddin" authorityYear="1904" box="[361,599,367,393]" class="Insecta" family="Reduviidae" genus="Castolus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="550" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="E8BB574D8F1FFFF3FECEFE94FDCC0C9E" box="[361,599,367,393]" italics="true" pageId="12" pageNumber="550">Castolus nigriventris</emphasis>
</taxonomicName>
is known from
<collectingCountry id="A2D8CBCF8F1FFFF3FCAAFE94FCC40C9F" box="[781,863,367,393]" name="Bolivia" pageId="12" pageNumber="550">Bolivia</collectingCountry>
(
<bibRefCitation id="BE5EF6AE8F1FFFF3FCC8FE94FB970C9F" author="Breddin, G." box="[879,1036,367,393]" pageId="12" pageNumber="550" pagination="147 - 148" refId="ref15054" refString="Breddin, G. (1904) Neue Rhynchotenausbeute aus Sud-Amerika. Societas Entomologica, 20, 147 - 148 + 153 - 154 + 177 - 178. https: // www. biodiversitylibrary. org / page / 9279381" type="journal article" year="1904">Breddin 1904</bibRefCitation>
) and
<collectingCountry id="A2D8CBCF8F1FFFF3FBEAFE94FAA20C9F" box="[1101,1337,367,393]" name="Trinidad and Tobago" pageId="12" pageNumber="550">Trinidad and Tobago</collectingCountry>
(
<bibRefCitation id="BE5EF6AE8F1FFFF3FAEFFE94FE8A0CB8" author="Maldonado, J." pageId="12" pageNumber="550" pagination="435 - 446" refId="ref15942" refString="Maldonado, J. (1976) Three new species of Castolus and a key to the species (Hemiptera: Reduviidae). Proceedings of the Entomological Society of Washington, 78, 435 - 446. https: // www. biodiversitylibrary. org / part / 56099" type="journal article" year="1976">Maldonado 1976</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>