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<document id="CAA5E600971CBBBA6DC3FFB01ECF5843" ID-CLB-Dataset="46514" ID-DOI="10.5281/zenodo.203270" ID-GBIF-Dataset="356611de-8d26-46da-a75f-f2c07aea11d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="203270" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460379449012" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Komai, Tomoyuki" docDate="2011" docId="5240BA109763FFC64DDBF8ADFAFE255D" docLanguage="en" docName="zt02934p067.pdf" docOrigin="Zootaxa 2934" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pedroplax Ng &amp; Komai, 2011, new genus" docType="treatment" docVersion="10" lastPageNumber="62" masterDocId="AE79C2689763FFC74D4CFF90FFAA2030" masterDocTitle="A new genus for Psopheticus megalops Takeda, 1989 (Crustacea: Brachyura: Goneplacidae)" masterLastPageNumber="67" masterPageNumber="61" pageNumber="61" updateTime="1698296645123" updateUser="ExternalLinkService">
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<mods:titleInfo id="6822E836CE5B32E2F068D1C367BB1139">
<mods:title id="ED791C17634BB5440071CB7F2CF009CB">A new genus for Psopheticus megalops Takeda, 1989 (Crustacea: Brachyura: Goneplacidae)</mods:title>
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<mods:namePart id="A67341B43CB0748DBD2592C380E114B1">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="3EF28E8F30830F34C494F433AAEB44F8">Komai, Tomoyuki</mods:namePart>
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<mods:title id="5D42519A050DBFE72CF1938FC6312E63">Zootaxa</mods:title>
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<mods:date id="3911E0661564A73BEC3BB9BB319B2D9B">2011</mods:date>
<mods:detail id="37E5A5136D2C25C54CC04E913D957E10" type="volume">
<mods:number id="DF9FCCFD52D6E5644ACEEBCAA7653904">2934</mods:number>
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<mods:classification id="19CFCC9DCCD07856EE4F7CBD3D925337">journal article</mods:classification>
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<mods:identifier id="360E17547133951F61E035FFAC16BC1D" type="DOI">10.5281/zenodo.203270</mods:identifier>
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<treatment id="5240BA109763FFC64DDBF8ADFAFE255D" ID-DOI="http://doi.org/10.5281/zenodo.6194159" ID-GBIF-Taxon="119413861" ID-Zenodo-Dep="6194159" LSID="urn:lsid:plazi:treatment:5240BA109763FFC64DDBF8ADFAFE255D" httpUri="http://treatment.plazi.org/id/5240BA109763FFC64DDBF8ADFAFE255D" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
<subSubSection id="92F3588D9763FFC74DDBF8ADFE362767" box="[151,412,1853,1879]" pageId="0" pageNumber="61" type="nomenclature">
<paragraph id="DA560B069763FFC74DDBF8ADFE362767" blockId="0.[151,412,1853,1879]" box="[151,412,1853,1879]" pageId="0" pageNumber="61">
<heading id="811EBC6A9763FFC74DDBF8ADFE362767" bold="true" box="[151,412,1853,1879]" fontSize="11" level="1" pageId="0" pageNumber="61" reason="1">
<emphasis id="E89DD7149763FFC74DDBF8ADFE362767" bold="true" box="[151,412,1853,1879]" pageId="0" pageNumber="61">
<taxonomicName id="1DE970859763FFC74DDBF8ADFEB92767" ID-CoL="7PSTT" box="[151,275,1853,1879]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E89DD7149763FFC74DDBF8ADFEB92767" bold="true" box="[151,275,1853,1879]" italics="true" pageId="0" pageNumber="61">Pedroplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3AE6A6F9763FFC74C56F8ADFE362767" box="[282,412,1853,1879]" pageId="0" pageNumber="61" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92F3588D9763FFC64DDBF813FC712242" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61" type="diagnosis">
<paragraph id="DA560B069763FFC64DDBF813FC712242" blockId="0.[151,1436,1923,2016]" lastBlockId="1.[151,1436,151,1390]" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
<emphasis id="E89DD7149763FFC74DDBF813FEBB27AC" bold="true" box="[151,273,1923,1948]" pageId="0" pageNumber="61">Diagnosis.</emphasis>
Carapace slightly wider than long, with single anterolateral tooth (
<figureCitation id="42D217839763FFC7495DF813FBCA27AB" box="[1041,1120,1923,1947]" captionStart="FIGURE 1" captionStartId="3.[151,250,1952,1975]" captionTargetBox="[243,1344,195,1928]" captionTargetId="figure@3.[243,1345,195,1928]" captionTargetPageId="3" captionText="FIGURE 1. Pedroplax megalops (Takeda, 1989) new combination. A, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); B, male (5.2 × 4.4 mm) (CBM-ZC 5936); C, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297)." httpUri="https://zenodo.org/record/203271/files/figure.png" pageId="0" pageNumber="61">Figs. 1</figureCitation>
,
<figureCitation id="42D217839763FFC7493CF813FBD427AB" box="[1136,1150,1923,1947]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="0" pageNumber="61">2</figureCitation>
A, 3A). Orbit short; eyes large, protruding lateral margin of orbit (
<figureCitation id="42D217839763FFC74F21F836FD16278E" box="[621,700,1958,1982]" captionStart="FIGURE 1" captionStartId="3.[151,250,1952,1975]" captionTargetBox="[243,1344,195,1928]" captionTargetId="figure@3.[243,1345,195,1928]" captionTargetPageId="3" captionText="FIGURE 1. Pedroplax megalops (Takeda, 1989) new combination. A, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); B, male (5.2 × 4.4 mm) (CBM-ZC 5936); C, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297)." httpUri="https://zenodo.org/record/203271/files/figure.png" pageId="0" pageNumber="61">Figs. 1</figureCitation>
,
<figureCitation id="42D217839763FFC74F81F836FD71278E" box="[717,731,1958,1982]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="0" pageNumber="61">2</figureCitation>
A, 3A). Antennules proportionately large, fossa asymmetrical (
<figureCitation id="42D217839763FFC74DD3F858FF4C27D0" box="[159,230,1992,2016]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="0" pageNumber="61">Fig. 2</figureCitation>
A). Epistome with anterior part triangular; posterior margin sinuous with 3 low median lobes, separated from lateral parts by deep concavity (
<figureCitation id="42D217839762FFC64F08FF07FD23209F" box="[580,649,151,175]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
A). Endostomial ridges strong. Third maxilliped with merus about as long as broad, anterolateral angle rounded, not auriculiform; ischium subrectangular (
<figureCitation id="42D217839762FFC64937FF2AFB6720E2" box="[1147,1229,186,210]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
C, 3B). Chelipeds homochelous; chela stout, fingers laterally flattened (
<figureCitation id="42D217839762FFC64E4DFF4DFCF920C5" box="[769,851,221,245]" captionStart="FIGURE 1" captionStartId="3.[151,250,1952,1975]" captionTargetBox="[243,1344,195,1928]" captionTargetId="figure@3.[243,1345,195,1928]" captionTargetPageId="3" captionText="FIGURE 1. Pedroplax megalops (Takeda, 1989) new combination. A, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); B, male (5.2 × 4.4 mm) (CBM-ZC 5936); C, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297)." httpUri="https://zenodo.org/record/203271/files/figure.png" pageId="1" pageNumber="62">Figs. 1</figureCitation>
,
<figureCitation id="42D217839762FFC64E28FF4DFCD820C5" box="[868,882,221,245]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1907]" captionTargetBox="[186,1387,205,1838]" captionTargetId="figure@5.[178,1391,193,1846]" captionTargetPageId="5" captionText="FIGURE 3. Pedroplax megalops (Takeda, 1989) new combination, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754). A, dorsal view of carapace; B, left third maxilliped; C, dorsal view of left cheliped; D, outer view of left chela; E, right third ambulatory leg; F, anterior thoracic sternum and abdominal somites 3 – 6 with telson; G, posterior thoracic sternum and abdominal somites 1 – 3; H, left G 1; I, distal part of left G 1; J, left G 2; K, distal part of left G 2. Abbreviations: a 1 – a 6 = male abdominal somites 1 – 6; c 5 = coxae of fifth ambulatory leg; st 4 – 8 = thoracic sternites 4 – 8; t = telson; v = vulva. (After Takeda 1989: figs. 16, 17)." httpUri="https://zenodo.org/record/203273/files/figure.png" pageId="1" pageNumber="62">3</figureCitation>
D). Ambulatory legs slender, smooth, unarmed; propodus of fourth leg subfoliaceous; dactylus of fourth leg subspatuliform (
<figureCitation id="42D217839762FFC64944FF6FFBE02127" box="[1032,1098,255,279]" captionStart="FIGURE 1" captionStartId="3.[151,250,1952,1975]" captionTargetBox="[243,1344,195,1928]" captionTargetId="figure@3.[243,1345,195,1928]" captionTargetPageId="3" captionText="FIGURE 1. Pedroplax megalops (Takeda, 1989) new combination. A, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); B, male (5.2 × 4.4 mm) (CBM-ZC 5936); C, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297)." httpUri="https://zenodo.org/record/203271/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
B, C). Thoracic sternites 1, 2 completely fused, forming prominent triangular plate (
<figureCitation id="42D217839762FFC64FB4FEB2FCEC210A" box="[760,838,290,314]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
C, D); sternites 3, 4 fused, only lateral sutures visible; sutures between sternites 4–8 medially interrupted; longitudinal groove present between sternites 6–8. Press button of abdominal locking mechanism on anterior half of sternite 5 (
<figureCitation id="42D217839762FFC64E10FEF7FC00214F" box="[860,938,359,383]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
D). Coxo-sternal condition, with penis coxal but opening to base of G1 through narrow channel formed by episternite 7, sternite 8 (
<figureCitation id="42D217839762FFC64910FE1AFB352192" box="[1116,1183,394,418]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
E). Male abdomen relatively broadly T-shaped; somite 2 not reaching coxae of fourth pair of ambulatory legs, sternite 8 clearly exposed; somite 3 transversely trapezoidal, almost reaching coxae of fourth ambulatory legs except for narrow part of sternite 8 (
<figureCitation id="42D217839762FFC64D9BFE62FE8F223A" box="[215,293,498,522]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
B, 3G). G1 stout, gently curved outwards, distal quarter tapering with 2 stout subdistal spines on outer margin (
<figureCitation id="42D217839762FFC64DB8FD85FE9C221D" box="[244,310,533,557]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1907]" captionTargetBox="[186,1387,205,1838]" captionTargetId="figure@5.[178,1391,193,1846]" captionTargetPageId="5" captionText="FIGURE 3. Pedroplax megalops (Takeda, 1989) new combination, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754). A, dorsal view of carapace; B, left third maxilliped; C, dorsal view of left cheliped; D, outer view of left chela; E, right third ambulatory leg; F, anterior thoracic sternum and abdominal somites 3 – 6 with telson; G, posterior thoracic sternum and abdominal somites 1 – 3; H, left G 1; I, distal part of left G 1; J, left G 2; K, distal part of left G 2. Abbreviations: a 1 – a 6 = male abdominal somites 1 – 6; c 5 = coxae of fifth ambulatory leg; st 4 – 8 = thoracic sternites 4 – 8; t = telson; v = vulva. (After Takeda 1989: figs. 16, 17)." httpUri="https://zenodo.org/record/203273/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
H, I); G2 about 1.3 times length of G1, median part dilated with numerous short spines on outer margin, distal part with bifurcated tip, subdistal surface with 4 or 5 spines (
<figureCitation id="42D217839762FFC64E81FDA7FBBB227F" box="[973,1041,567,591]" captionStart="FIGURE 3" captionStartId="5.[151,250,1884,1907]" captionTargetBox="[186,1387,205,1838]" captionTargetId="figure@5.[178,1391,193,1846]" captionTargetPageId="5" captionText="FIGURE 3. Pedroplax megalops (Takeda, 1989) new combination, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754). A, dorsal view of carapace; B, left third maxilliped; C, dorsal view of left cheliped; D, outer view of left chela; E, right third ambulatory leg; F, anterior thoracic sternum and abdominal somites 3 – 6 with telson; G, posterior thoracic sternum and abdominal somites 1 – 3; H, left G 1; I, distal part of left G 1; J, left G 2; K, distal part of left G 2. Abbreviations: a 1 – a 6 = male abdominal somites 1 – 6; c 5 = coxae of fifth ambulatory leg; st 4 – 8 = thoracic sternites 4 – 8; t = telson; v = vulva. (After Takeda 1989: figs. 16, 17)." httpUri="https://zenodo.org/record/203273/files/figure.png" pageId="1" pageNumber="62">Fig. 3</figureCitation>
J, K). Vulva prominent, occupying most of anterior half of sternite 6, no visible sternal vulvar cover (
<figureCitation id="42D217839762FFC64E35FDCAFC162242" box="[889,956,602,626]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="92F3588D9762FFC64D89FDEDFAD02518" pageId="1" pageNumber="62" type="discussion">
<paragraph id="DA560B069762FFC64D89FDEDFB7C239A" blockId="1.[151,1436,151,1390]" pageId="1" pageNumber="62">
<emphasis id="E89DD7149762FFC64D89FDEDFE9322A5" bold="true" box="[197,313,637,661]" pageId="1" pageNumber="62">Remarks.</emphasis>
Although the carapace is superficially similar to species of
<taxonomicName id="1DE970859762FFC64EBDFDEDFACC22A5" authority="Wood-Mason, 1892" authorityName="Wood-Mason" authorityYear="1892" box="[1009,1382,637,661]" class="Malacostraca" family="Goneplacidae" genus="Psopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64EBDFDEDFBD222A5" box="[1009,1144,637,661]" italics="true" pageId="1" pageNumber="62">Psopheticus</emphasis>
<bibRefCitation id="BE7876F79762FFC649CFFDEDFACC22A5" author="Wood-Mason" box="[1155,1382,637,661]" pageId="1" pageNumber="62" refString="Wood-Mason, J. (1892) Illustrations of the Zoology of H. M. Indian Marine Surveying Steamer Investigator, under the command of Commander A. Carpenter, R. N., D. S. O., and of Commander R. F. Hoskyn, R. N. Part I. Crustaceans. Plates I - V. Director of the Royal Indian Marine, Calcutta, [five unnumbered pages for Explanation of Plates], pls. I - V." type="book" year="1892">Wood-Mason, 1892</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="1DE970859762FFC64DDBFD30FE112287" authority="Castro, 2007" authorityName="Castro" authorityYear="2007" box="[151,443,671,696]" class="Malacostraca" family="Goneplacidae" genus="Exopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64DDBFD30FE8A2288" box="[151,288,672,696]" italics="true" pageId="1" pageNumber="62">Exopheticus</emphasis>
<bibRefCitation id="BE7876F79762FFC64C64FD0FFE112287" author="Castro" box="[296,443,671,695]" pageId="1" pageNumber="62" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro, 2007</bibRefCitation>
</taxonomicName>
, it is distinctly less broad proportionately and appears paedomorphic, resembling juveniles of species of those latter genera. This is also true of its disproportionately large eyes. However, there is no doubt
<taxonomicName id="1DE970859762FFC64DADFD75FE6022CD" box="[225,458,741,765]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="E89DD7149762FFC64DADFD75FE6022CD" box="[225,458,741,765]" italics="true" pageId="1" pageNumber="62">Pedroplax megalops</emphasis>
</taxonomicName>
is an adult despite its small size; the gonopods are fully developed and three of the females examined are already ovigerous. Although
<bibRefCitation id="BE7876F79762FFC64F95FC98FCDF2310" author="Castro" box="[729,885,776,800]" pageId="1" pageNumber="62" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro (2007)</bibRefCitation>
and Ng
<emphasis id="E89DD7149762FFC64E99FC99FBA72310" box="[981,1037,776,800]" italics="true" pageId="1" pageNumber="62">et al.</emphasis>
(2008) questioned its classification in the
<taxonomicName id="1DE970859762FFC64DADFCBAFD9D2372" authority="MacLeay, 1838" authorityName="MacLeay" authorityYear="1838" box="[225,567,810,835]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="family">
Goneplacidae
<bibRefCitation id="BE7876F79762FFC64CC8FCBBFD9D2372" author="MacLeay" box="[388,567,810,835]" pageId="1" pageNumber="62" refString="MacLeay, W. S. (1838) On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In A. Smith, Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa': together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe, Published under the Authority of the Lords Commissioners of Her Majesty's Treasury, Invertebratae: [1849] pp. 53 - 71, pls 2, 3." type="journal article" year="1838">MacLeay, 1838</bibRefCitation>
</taxonomicName>
,
<emphasis id="E89DD7149762FFC64F08FCBCFD792373" box="[580,723,812,835]" italics="true" pageId="1" pageNumber="62">sensu stricto</emphasis>
, there is little doubt it is a goneplacid as currently defined by
<bibRefCitation id="BE7876F79762FFC64DDBFCDDFE992355" author="Castro" box="[151,307,845,869]" pageId="1" pageNumber="62" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro (2007)</bibRefCitation>
, Ng &amp;
<bibRefCitation id="BE7876F79762FFC64CCFFCDDFDD72355" box="[387,637,845,869]" pageId="1" pageNumber="62" refString="Ng, P. K. L. &amp; Manuel-Santos, M. R. (2007) Establishment of the Vultocinidae, a new family for an unusual new genus and new species of Indo-West Pacific crab (Crustacea: Decapoda: Brachyura: Goneplacoidea), with comments on the taxonomy of the Goneplacidae. Zootaxa, 1558, 39 - 68." type="journal article">Manuel-Santos (2007)</bibRefCitation>
and
<bibRefCitation id="BE7876F79762FFC64FFEFCDDFC272355" box="[690,909,845,869]" pageId="1" pageNumber="62" refString="Castro, P., Guinot, D. &amp; Ng, P. K. L. (2009) A new family for Sotoplax robertsi Guinot, 1984, with a diagnosis and key to the Goneplacoidea MacLeay, 1838 (Crustacea: Decapoda: Brachyura). Zootaxa, 2356, 36 - 56." type="journal article">
Castro
<emphasis id="E89DD7149762FFC64E4EFCDEFC932355" box="[770,825,845,869]" italics="true" pageId="1" pageNumber="62">et al.</emphasis>
(2009)
</bibRefCitation>
. The structures of the thoracic sternum (including the arrangement and extent of sutures) and male abdomen as well as the position of the penis and press-button of abdominal locking mechanism of
<taxonomicName id="1DE970859762FFC64F7EFC03FD0C239B" box="[562,678,915,939]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E89DD7149762FFC64F7EFC03FD0C239B" box="[562,678,915,939]" italics="true" pageId="1" pageNumber="62">Pedroplax</emphasis>
</taxonomicName>
<emphasis id="E89DD7149762FFC64FE1FC03FC82239B" bold="true" box="[685,808,915,939]" pageId="1" pageNumber="62">
<taxonomicNameLabel id="F3AE6A6F9762FFC64FE1FC03FC82239B" box="[685,808,915,939]" pageId="1" pageNumber="62" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
are consistent with other goneplacids.
</paragraph>
<paragraph id="DA560B069762FFC64D89FC25FBAE24F0" blockId="1.[151,1436,151,1390]" pageId="1" pageNumber="62">
The most unusual features of
<taxonomicName id="1DE970859762FFC64F5BFC25FD2123FD" box="[535,651,949,973]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E89DD7149762FFC64F5BFC25FD2123FD" box="[535,651,949,973]" italics="true" pageId="1" pageNumber="62">Pedroplax</emphasis>
</taxonomicName>
<emphasis id="E89DD7149762FFC64FDFFC25FCBA23FD" bold="true" box="[659,784,949,973]" pageId="1" pageNumber="62">
<taxonomicNameLabel id="F3AE6A6F9762FFC64FDFFC25FCBA23FD" box="[659,784,949,973]" pageId="1" pageNumber="62" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
are clearly the very peculiar G1 and G2 structures which are unique for the family. In fact, the unusual G1s and G2s were the main reason why its placement in the
<taxonomicName id="1DE970859762FFC6481AFC48FF5E2422" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
was doubted by
<bibRefCitation id="BE7876F79762FFC64CF4FC6AFDFF2422" author="Castro" box="[440,597,1018,1042]" pageId="1" pageNumber="62" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro (2007)</bibRefCitation>
and Ng
<emphasis id="E89DD7149762FFC64FF5FC6CFD592423" box="[697,755,1019,1043]" italics="true" pageId="1" pageNumber="62">et al.</emphasis>
(2008), but as discussed above, all the other key characters argue for its retention in the
<taxonomicName id="1DE970859762FFC64CAFFB8DFD2B2405" box="[483,641,1053,1077]" class="Malacostraca" family="Goneplacidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="family">Goneplacidae</taxonomicName>
. The gonopod structures, together with the atypical carapace features argue for the recognition of a distinct genus for the species. Another key difference of
<taxonomicName id="1DE970859762FFC64927FBD0FB742468" box="[1131,1246,1088,1112]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E89DD7149762FFC64927FBD0FB742468" box="[1131,1246,1088,1112]" italics="true" pageId="1" pageNumber="62">Pedroplax</emphasis>
</taxonomicName>
<emphasis id="E89DD7149762FFC649AAFBD0FAC92468" bold="true" box="[1254,1379,1088,1112]" pageId="1" pageNumber="62">
<taxonomicNameLabel id="F3AE6A6F9762FFC649AAFBD0FAC92468" box="[1254,1379,1088,1112]" pageId="1" pageNumber="62" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
with
<taxonomicName id="1DE970859762FFC64DDBFBF3FEB4244B" box="[151,286,1123,1147]" class="Malacostraca" family="Goneplacidae" genus="Psopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64DDBFBF3FEB4244B" box="[151,286,1123,1147]" italics="true" pageId="1" pageNumber="62">Psopheticus</emphasis>
</taxonomicName>
and
<taxonomicName id="1DE970859762FFC64C15FBF3FE48244B" box="[345,482,1123,1147]" class="Malacostraca" family="Goneplacidae" genus="Exopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64C15FBF3FE48244B" box="[345,482,1123,1147]" italics="true" pageId="1" pageNumber="62">Exopheticus</emphasis>
</taxonomicName>
is the relative narrowness of male abdominal somite 2 that exposes a large part of thoracic sternite 8 (
<figureCitation id="42D217839762FFC64C3CFB15FE1424AD" box="[368,446,1157,1181]" captionStart="FIGURE 2" captionStartId="4.[151,250,1735,1758]" captionTargetBox="[151,1436,193,1713]" captionTargetId="figure@4.[151,1436,193,1714]" captionTargetPageId="4" captionText="FIGURE 2. Pedroplax megalops (Takeda, 1989) new combination. A-E, holotype male (5.6 × 4.8 mm) (NSMT-Cr 9754); F, paratype female (5.1 × 4.4 mm) (NSMT-Cr 5297). A, frontal view of carapace; B, posterior view of carapace showing anterior abdominal somites; C, anterior part of thoracic sternum showing posterior abdominal somites; D, anterior part of thoracic sternum showing sterno-abdominal cavity and press-button; E, left side of posterior thoracic sternum showing position of penis; F, anterior thoracic sternum of sternum showing vulvae. Abbreviations: a 1 – a 4 = male abdominal somites 1 – 4; c 3 – c 5 = coxae of third to fifth ambulatory legs; es 7 = episternite 7; p = penis; pb = press-button of abdominal locking mechanism; st 6 – 8 = thoracic sternites 6 – 8; v = vulva." httpUri="https://zenodo.org/record/203272/files/figure.png" pageId="1" pageNumber="62">Figs. 2</figureCitation>
B, 3G). In
<taxonomicName id="1DE970859762FFC64F7BFB15FD1424AD" box="[567,702,1157,1181]" class="Malacostraca" family="Goneplacidae" genus="Psopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64F7BFB15FD1424AD" box="[567,702,1157,1181]" italics="true" pageId="1" pageNumber="62">Psopheticus</emphasis>
</taxonomicName>
and
<taxonomicName id="1DE970859762FFC64FB9FB15FCD424AD" box="[757,894,1157,1181]" class="Malacostraca" family="Goneplacidae" genus="Exopheticus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="E89DD7149762FFC64FB9FB15FCD424AD" box="[757,894,1157,1181]" italics="true" pageId="1" pageNumber="62">Exopheticus</emphasis>
</taxonomicName>
, the male abdominal somite is broader and only a relatively small part of sternite 8 is exposed, if at all (
<bibRefCitation id="BE7876F79762FFC64FB3FB38FC5D24F0" author="Castro" box="[767,1015,1192,1216]" pageId="1" pageNumber="62" refString="Castro, P. (2007) A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species. Zoosystema, 29 (4), 609 - 773." type="journal article" year="2007">Castro 2007: fig. 48B</bibRefCitation>
).
</paragraph>
<paragraph id="DA560B069762FFC64D89FB5AFAD02518" blockId="1.[151,1436,151,1390]" pageId="1" pageNumber="62">
With regard to the small adult size and disproportionately large eyes and general form of the last ambulatory legs,
<taxonomicName id="1DE970859762FFC64D9FFB7DFE132535" box="[211,441,1261,1285]" class="Malacostraca" family="Goneplacidae" genus="Pedroplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="megalops">
<emphasis id="E89DD7149762FFC64D9FFB7DFE132535" box="[211,441,1261,1285]" italics="true" pageId="1" pageNumber="62">Pedroplax megalops</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="1DE970859762FFC64F80FB7DFB2B2535" authority="Castro &amp; Ng, 2010" authorityName="Castro &amp; Ng" authorityYear="2010" box="[716,1153,1261,1286]" class="Malacostraca" family="Goneplacidae" genus="Guinoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="apheles">
<emphasis id="E89DD7149762FFC64F80FB7DFC0A2535" box="[716,928,1261,1285]" italics="true" pageId="1" pageNumber="62">Guinoplax apheles</emphasis>
<bibRefCitation id="BE7876F79762FFC64EEBFB7DFB2B2535" author="Castro" box="[935,1153,1261,1286]" pageId="1" pageNumber="62" refString="Castro, P. &amp; Ng, P. K. L. (2010) A new genus and species of goneplacid crab (Decapoda, Brachyura, Goneplacidae) from the western Pacific. Crustaceana Monographs, 11, 51 - 60." type="journal article" year="2010">Castro &amp; Ng, 2010</bibRefCitation>
</taxonomicName>
, but the two differ markedly in the characters of the carapace, chela, abdomen, thoracic sternum and gonopods (see
<bibRefCitation id="BE7876F79762FFC649D7FA80FAC42518" author="Castro" box="[1179,1390,1296,1321]" pageId="1" pageNumber="62" refString="Castro, P. &amp; Ng, P. K. L. (2010) A new genus and species of goneplacid crab (Decapoda, Brachyura, Goneplacidae) from the western Pacific. Crustaceana Monographs, 11, 51 - 60." type="journal article" year="2010">Castro &amp; Ng 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="92F3588D9762FFC64D89FAA3FAFE255D" pageId="1" pageNumber="62" type="etymology">
<paragraph id="DA560B069762FFC64D89FAA3FAFE255D" blockId="1.[151,1436,151,1390]" pageId="1" pageNumber="62">
<emphasis id="E89DD7149762FFC64D89FAA3FEE1257B" bold="true" box="[197,331,1331,1355]" pageId="1" pageNumber="62">Etymology.</emphasis>
We take pleasure in naming the genus after Peter Castro, known to his carcinologist friends as “Pedro. The name is used in combination with -
<emphasis id="E89DD7149762FFC64FE2FAC5FD74255D" box="[686,734,1365,1389]" italics="true" pageId="1" pageNumber="62">plax</emphasis>
, a suffix for many goneplacid genera. Gender feminine.
</paragraph>
</subSubSection>
</treatment>
</document>
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