188 lines
17 KiB
XML
188 lines
17 KiB
XML
<document id="1BA62360C26EB8CFB1F09BA3D6E9A8BF" ID-CLB-Dataset="55854" ID-DOI="10.11646/zootaxa.5128.1.6" ID-GBIF-Dataset="1d4bdfae-b87b-4824-afa7-049177f8fad0" ID-ISSN="1175-5326" ID-Zenodo-Dep="6479636" ID-ZooBank="6BB58C2A-323F-4DE4-A0CA-3140428D1D3B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650719839255" checkinUser="plazi" docAuthor="Teoh, Vincent Yong Jian & Woo, Sau Pinn" docDate="2022" docId="4150A955FF84A56DFF2CFF18E0ADFD7D" docLanguage="en" docName="zootaxa.5128.1.6.pdf" docOrigin="Zootaxa 5128 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acaudina H. L. Clark 1908" docType="key" docVersion="6" lastPageNumber="117" masterDocId="BD69D12DFF8EA567FFBBFF8FE530FFDE" masterDocTitle="Two new species of sea cucumbers (Echinodermata: Holothuroidea) from the seagrass meadow of Penang, Malaysia" masterLastPageNumber="118" masterPageNumber="107" pageNumber="117" updateTime="1699329046161" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="73483D69F908A09723B239A816208056" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3AF25FAE453E8969A3C7ADFFFC47C710">
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<mods:title id="27BCA29DC7974E604A444729E8772623">Two new species of sea cucumbers (Echinodermata: Holothuroidea) from the seagrass meadow of Penang, Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="A3A962A29665565AA5FFAB2AD9B7F435" type="personal">
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<mods:roleTerm id="AC0E430C6151409B7628AACD9BFFCCDE">Author</mods:roleTerm>
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<mods:namePart id="576B804F3CE275687E1D4B4AC698AABF">Teoh, Vincent Yong Jian</mods:namePart>
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<mods:name id="A99D8C26C526D8DD735AC546C9CF205C" type="personal">
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<mods:roleTerm id="316D8174297BAA0051F9049CDBD49E06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6E250BDDDF8F94BCC42F37E243ED42FC">Woo, Sau Pinn</mods:namePart>
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<mods:affiliation id="5EA257FAFFDEAB1793EACC1649D9ED56">Corresponding: woosaupinn @ usm. my</mods:affiliation>
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<mods:typeOfResource id="DD681664F1ECE948657FF0037A87E006">text</mods:typeOfResource>
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<mods:titleInfo id="412D0B60408FE1122A740B0289D5320B">
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<mods:title id="4E1668A6053F9E37C48492321E54DB11">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8399BAEF2CE524FB2CC4DA8C4A9E6455">
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<mods:date id="C952B5A73D78694C0570F9B9C019B3F7">2022</mods:date>
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<mods:detail id="CAE1F706B090E75630C3A903939591AA" type="pubDate">
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<mods:number id="A3DF814606941E872D4A185D91CDBE99">2022-04-19</mods:number>
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<mods:detail id="71C7637D35C2B947ADB20B84B869CC7A" type="volume">
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<mods:number id="B56FCA6F4F6402332BB36542C0EE3966">5128</mods:number>
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<mods:detail id="5BBCA2ED6DCCBB19C7F20A05274FBD32" type="issue">
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<mods:number id="BBA2D0A1FDD10518630C304ADA0EC45C">1</mods:number>
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<mods:start id="A780AE6D65BD169E1B60B3B4050CB013">107</mods:start>
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<mods:classification id="7910D14874A58667C1AE044CFE505F78">journal article</mods:classification>
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<mods:identifier id="A17AC4128D4CE3B5B6630095541F5038" type="CLB-Dataset">55854</mods:identifier>
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<mods:identifier id="7A1F4BAF4B063A55D499AD2C1609D13E" type="DOI">10.11646/zootaxa.5128.1.6</mods:identifier>
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<mods:identifier id="C86A85AF3684BC08AB0DCC34F8184A9E" type="GBIF-Dataset">1d4bdfae-b87b-4824-afa7-049177f8fad0</mods:identifier>
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<mods:identifier id="E91F3EA93C8BBE0D9F830417BE6FA602" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4150A955FF84A56DFF2CFF18E0ADFD7D" ID-DOI="http://doi.org/10.5281/zenodo.6498167" ID-GBIF-Taxon="194847497" ID-Zenodo-Dep="6498167" LSID="urn:lsid:plazi:treatment:4150A955FF84A56DFF2CFF18E0ADFD7D" httpUri="http://treatment.plazi.org/id/4150A955FF84A56DFF2CFF18E0ADFD7D" lastPageNumber="117" pageId="10" pageNumber="117">
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<subSubSection id="81E34BC8FF84A56DFF2CFF18E7BEFF6C" box="[151,654,151,178]" pageId="10" pageNumber="117" type="nomenclature">
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<paragraph id="C9461843FF84A56DFF2CFF18E7BEFF6C" blockId="10.[151,654,151,178]" box="[151,654,151,178]" pageId="10" pageNumber="117">
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<heading id="920EAF2FFF84A56DFF2CFF18E7BEFF6C" bold="true" box="[151,654,151,178]" fontSize="11" level="1" pageId="10" pageNumber="117" reason="1">
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<emphasis id="FB8DC451FF84A56DFF2CFF18E7BEFF6C" bold="true" box="[151,654,151,178]" pageId="10" pageNumber="117">
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Identification key to the genus
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<taxonomicName id="0EF963C0FF84A56DFDA2FF18E7BEFF6C" ID-CoL="7NMPM" authorityName="H. L. Clark" authorityYear="1908" box="[537,654,151,178]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="genus">
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<emphasis id="FB8DC451FF84A56DFDA2FF18E7BEFF6C" bold="true" box="[537,654,151,178]" italics="true" pageId="10" pageNumber="117">Acaudina</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="81E34BC8FF84A56DFF2CFF59E0ADFD7D" pageId="10" pageNumber="117" type="key">
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<key id="BD684F72FF84A56DFF2CFF59E0ADFD7D" pageId="10" pageNumber="117">
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<keyStep id="720D00E6FF84A56DFF2CFF59E0ACFED7" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFF59E0ACFF35" blockId="10.[151,1437,213,675]" box="[151,1436,213,235]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFF59E0ACFF35" box="[151,1436,213,235]" pageId="10" pageNumber="117">1. Presence of doughnut-shaped bodies….................................................................... 2</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFF7CE0ACFED7" blockId="10.[151,1437,213,675]" box="[151,1436,243,265]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFF7CE0ACFED7" box="[151,1436,243,265]" pageId="10" pageNumber="117">- Absence of doughnut-shaped bodies….................................................................... 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFE9FE0ACFE9D" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFE9FE0ADFEF8" blockId="10.[151,1437,213,675]" box="[151,1437,272,294]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE9FE0ADFEF8" box="[151,1437,272,294]" pageId="10" pageNumber="117">
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2. Doughnut-shaped bodies spinose........................................................ …
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<taxonomicName id="0EF963C0FF84A56DFB67FE9FE07CFEFB" authority="Teoh & Woo, 2022" authorityName="Teoh & Woo" authorityYear="2022" box="[1244,1356,272,293]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="spinifera" status="sp. nov.">
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<emphasis id="FB8DC451FF84A56DFB67FE9FE07CFEFB" box="[1244,1356,272,293]" italics="true" pageId="10" pageNumber="117">A. spinifera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0BE792AFF84A56DFAE9FE9EE0ADFEF8" box="[1362,1437,273,294]" pageId="10" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFEA1E0ACFE9D" blockId="10.[151,1437,213,675]" box="[151,1436,301,323]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFEA1E0ACFE9D" box="[151,1436,301,323]" pageId="10" pageNumber="117">
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<emphasis id="FB8DC451FF84A56DFF2CFEA1E5AFFE9D" bold="true" box="[151,159,302,323]" pageId="10" pageNumber="117">-</emphasis>
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Doughnut-shaped bodies smooth…....................................................................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFEC4E0ACFEA3" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFEC4E0ACFEBE" blockId="10.[151,1437,213,675]" box="[151,1436,331,353]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFEC4E0ACFEBE" box="[151,1436,331,353]" pageId="10" pageNumber="117">
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3. Perforated plates either spinose or smooth…....................................................
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<taxonomicName id="0EF963C0FF84A56DFABBFEC4E0ACFEBE" ID-CoL="93KJ" box="[1280,1436,331,352]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="molpadioides">
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<emphasis id="FB8DC451FF84A56DFABBFEC4E0ACFEBE" box="[1280,1436,331,352]" italics="true" pageId="10" pageNumber="117">A. molpadioides</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFEE6E0ACFEA3" blockId="10.[151,1437,213,675]" box="[151,1436,360,382]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFEE6E0ACFEA3" box="[151,1436,360,382]" pageId="10" pageNumber="117">- Perforated plates always smooth…....................................................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFE09E0ACFE66" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFE09E0ACFE45" blockId="10.[151,1437,213,675]" box="[151,1436,389,411]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE09E0ACFE45" box="[151,1436,389,411]" pageId="10" pageNumber="117">
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4. Rod ossicles may have branched ends…...........................................................
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<taxonomicName id="0EF963C0FF84A56DFA95FE09E0ACFE45" ID-CoL="93KL" box="[1326,1436,390,411]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="punctata">
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<emphasis id="FB8DC451FF84A56DFA95FE09E0ACFE45" box="[1326,1436,390,411]" italics="true" pageId="10" pageNumber="117">A. punctata</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFE2CE0ACFE66" blockId="10.[151,1437,213,675]" box="[151,1436,419,441]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE2CE0ACFE66" box="[151,1436,419,441]" pageId="10" pageNumber="117">
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<emphasis id="FB8DC451FF84A56DFF2CFE2CE5AFFE66" box="[151,159,419,440]" italics="true" pageId="10" pageNumber="117">-</emphasis>
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Rod ossicles without branched ends…..........................................................
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<taxonomicName id="0EF963C0FF84A56DFAABFE2CE0ACFE66" ID-CoL="93KH" box="[1296,1436,419,440]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="leucoprocta">
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<emphasis id="FB8DC451FF84A56DFAABFE2CE0ACFE66" box="[1296,1436,419,440]" italics="true" pageId="10" pageNumber="117">A. leucoprocta</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFE4FE0ACFE2D" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFE4FE0ACFE0B" blockId="10.[151,1437,213,675]" box="[151,1436,448,470]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE4FE0ACFE0B" box="[151,1436,448,470]" pageId="10" pageNumber="117">
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5. Ossicles absent…............................................................................
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<taxonomicName id="0EF963C0FF84A56DFA93FE4FE0ACFE0B" ID-CoL="93KK" box="[1320,1436,448,469]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="pellucida">
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<emphasis id="FB8DC451FF84A56DFA93FE4FE0ACFE0B" box="[1320,1436,448,469]" italics="true" pageId="10" pageNumber="117">A. pellucida</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFE51E0ACFE2D" blockId="10.[151,1437,213,675]" box="[151,1436,477,499]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE51E0ACFE2D" box="[151,1436,477,499]" pageId="10" pageNumber="117">- Ossicles present.................................................................................... …6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFE74E0ACFDF0" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFE74E0ACFDCE" blockId="10.[151,1437,213,675]" box="[151,1436,507,529]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFE74E0ACFDCE" box="[151,1436,507,529]" pageId="10" pageNumber="117">
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6. Rosette present in body wall…....................................................................
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<taxonomicName id="0EF963C0FF84A56DFA87FE74E0ACFDCE" ID-CoL="93KM" box="[1340,1436,507,528]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="rosettis">
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<emphasis id="FB8DC451FF84A56DFA87FE74E0ACFDCE" box="[1340,1436,507,528]" italics="true" pageId="10" pageNumber="117">A. rosettis</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFD96E0ACFDF0" blockId="10.[151,1437,213,675]" box="[151,1436,536,558]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFD96E0ACFDF0" box="[151,1436,536,558]" pageId="10" pageNumber="117">
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<emphasis id="FB8DC451FF84A56DFF2CFD96E5AFFDF0" box="[151,159,537,558]" italics="true" pageId="10" pageNumber="117">-</emphasis>
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Rosette absent in body wall…........................................................................... 7
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFDB9E0ACFDB6" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFDB9E0ACFD94" blockId="10.[151,1437,213,675]" box="[151,1436,565,587]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFDB9E0ACFD94" box="[151,1436,565,587]" pageId="10" pageNumber="117">
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7. Spinose subspherical ossicles present in body wall….................................................
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<taxonomicName id="0EF963C0FF84A56DFA8FFDB9E0ACFD94" ID-CoL="93KF" box="[1332,1436,565,587]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="demissa">
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<emphasis id="FB8DC451FF84A56DFA8FFDB9E0ACFD94" box="[1332,1436,565,587]" italics="true" pageId="10" pageNumber="117">A. demissa</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFDDCE0ACFDB6" blockId="10.[151,1437,213,675]" box="[151,1436,595,617]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFDDCE0ACFDB6" box="[151,1436,595,617]" pageId="10" pageNumber="117">
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<emphasis id="FB8DC451FF84A56DFF2CFDDCE5AFFDB6" box="[151,159,595,616]" italics="true" pageId="10" pageNumber="117">-</emphasis>
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Spinose subspherical ossicles absent in body wall….......................................................... 8
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="720D00E6FF84A56DFF2CFDFFE0ADFD7D" pageId="10" pageNumber="117">
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<paragraph id="C9461843FF84A56DFF2CFDFFE0ADFD5B" blockId="10.[151,1437,213,675]" box="[151,1437,624,646]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFDFFE0ADFD5B" box="[151,1437,624,646]" pageId="10" pageNumber="117">
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8. Rod ossicles dominant in body wall; may be smooth or spinose........................................ …
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<taxonomicName id="0EF963C0FF84A56DFA84FDFFE0ADFD5B" ID-CoL="64FTW" box="[1343,1437,624,645]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="bacilla">
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<emphasis id="FB8DC451FF84A56DFA84FDFFE0ADFD5B" box="[1343,1437,624,645]" italics="true" pageId="10" pageNumber="117">A. bacilla</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="C9461843FF84A56DFF2CFD01E0ADFD7D" blockId="10.[151,1437,213,675]" box="[151,1437,653,675]" pageId="10" pageNumber="117">
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<keyLead id="7208BB76FF84A56DFF2CFD01E0ADFD7D" box="[151,1437,653,675]" pageId="10" pageNumber="117">
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<emphasis id="FB8DC451FF84A56DFF2CFD01E5AFFD7D" box="[151,159,654,675]" italics="true" pageId="10" pageNumber="117">-</emphasis>
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Spinose perforated plates dominant in body wall; rod ossicles always finely spinose…......................
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<taxonomicName id="0EF963C0FF84A56DFA94FD01E0ADFD7D" ID-CoL="93KN" box="[1327,1437,654,675]" class="Holothuroidea" family="Caudinidae" genus="Acaudina" kingdom="Animalia" order="Molpadida" pageId="10" pageNumber="117" phylum="Echinodermata" rank="species" species="suspecta">
|
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<emphasis id="FB8DC451FF84A56DFA94FD01E0ADFD7D" box="[1327,1437,654,675]" italics="true" pageId="10" pageNumber="117">A. suspecta</emphasis>
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</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</key>
|
|
</subSubSection>
|
|
</treatment>
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</document> |