treatments-xml/data/1D/17/F6/1D17F65BBE47C64C85D9B0A88977FC19.xml
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<document id="A4EA38C05026488EA3CAF0877D22DDE4" ID-CLB-Dataset="7378" ID-DOI="10.11646/zootaxa.4952.1.12" ID-GBIF-Dataset="6374cf98-83ad-4dbb-823f-cdd94ea9cafb" ID-ISSN="1175-5326" ID-Zenodo-Dep="4671621" ID-ZooBank="7D2013F1-9C51-4581-B281-D117012E0ACC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617870706140" checkinUser="plazi" docAuthor="Mückstein, Petr" docDate="2021" docId="1D17F65BBE47C64C85D9B0A88977FC19" docLanguage="en" docName="zootaxa.4952.1.12.pdf" docOrigin="Zootaxa 4952 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protaetia (Macroliocola) dianae Muckstein 2021, new species" docType="treatment" docVersion="7" lastPageNumber="199" masterDocId="E12E8E23BE43C64B854EB50A8D13FFBC" masterDocTitle="A new species and a new country record in the genus Protaetia Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae) from Vietnam" masterLastPageNumber="200" masterPageNumber="192" pageNumber="196" updateTime="1698936179732" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FFB7B45D420305B5AED66AB45CF1C8A4">A new species and a new country record in the genus Protaetia Burmeister, 1842 (Coleoptera: Scarabaeidae: Cetoniinae) from Vietnam</mods:title>
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<mods:namePart id="ECB7F48214DA3311757C0E946B3B4EE6">Mückstein, Petr</mods:namePart>
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<mods:title id="4744606D3FC9F50F9A43170FE752ABF5">Zootaxa</mods:title>
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<mods:date id="4D458BE1563A67EC26593684DA713476">2021</mods:date>
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<mods:number id="8343CF74D33FC7DB70999BE3E2C0FA4F">2021-04-08</mods:number>
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<mods:identifier id="CF9EB56FD32CD938757830CC28796958" type="DOI">10.11646/zootaxa.4952.1.12</mods:identifier>
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<treatment id="1D17F65BBE47C64C85D9B0A88977FC19" ID-DOI="http://doi.org/10.5281/zenodo.4671625" ID-GBIF-Taxon="181003684" ID-Zenodo-Dep="4671625" LSID="urn:lsid:plazi:treatment:1D17F65BBE47C64C85D9B0A88977FC19" httpUri="http://treatment.plazi.org/id/1D17F65BBE47C64C85D9B0A88977FC19" lastPageId="7" lastPageNumber="199" pageId="4" pageNumber="196">
<subSubSection id="DDA414C6BE47C64F85D9B0A88E56FA00" box="[151,837,1442,1469]" pageId="4" pageNumber="196" type="nomenclature">
<paragraph id="9501474DBE47C64F85D9B0A88E56FA00" blockId="4.[151,837,1442,1505]" box="[151,837,1442,1469]" pageId="4" pageNumber="196">
<heading id="CE49F021BE47C64F85D9B0A88E56FA00" bold="true" box="[151,837,1442,1469]" fontSize="11" level="1" pageId="4" pageNumber="196" reason="1">
<emphasis id="A7CA9B5FBE47C64F85D9B0A88E56FA00" bold="true" box="[151,837,1442,1469]" pageId="4" pageNumber="196">
<taxonomicName id="52BE3CCEBE47C64F85D9B0A88FBBFA00" authority="Muckstein" authorityName="Muckstein" authorityYear="2021" box="[151,680,1442,1469]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="196" phylum="Arthropoda" rank="species" species="dianae" status="sp. nov." subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE47C64F85D9B0A88F08FA01" bold="true" box="[151,539,1442,1469]" italics="true" pageId="4" pageNumber="196">Protaetia (Macroliocola) dianae</emphasis>
Mückstein
</taxonomicName>
,
<taxonomicNameLabel id="BCF92624BE47C64F87FBB0A88E56FA00" box="[693,837,1442,1468]" pageId="4" pageNumber="196" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE47C64F85D0B0CC8C7EFA5D" box="[158,365,1478,1505]" pageId="4" pageNumber="196" type="description">
<paragraph id="9501474DBE47C64F85D0B0CC8C7EFA5D" blockId="4.[151,837,1442,1505]" box="[158,365,1478,1505]" pageId="4" pageNumber="196">
(
<figureCitation id="0D855BC8BE47C64F85E8B0CC8C01FA5C" box="[166,274,1478,1504]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="4" pageNumber="196">Figs. 16</figureCitation>
,
<figureCitation id="0D855BC8BE47C64F8450B0CC8C76FA5D" box="[286,357,1478,1505]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="4" pageNumber="196">1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE47C64F85D9B3048E66F929" pageId="4" pageNumber="196" type="materials_examined">
<paragraph id="9501474DBE47C64F85D9B3048E40F994" blockId="4.[151,1437,1550,2008]" box="[151,851,1550,1577]" pageId="4" pageNumber="196">
<materialsCitation id="25D64D10BE47C64F85D9B3048E43F994" ID-GBIF-Occurrence="3071291303" box="[151,848,1550,1577]" country="Vietnam" location="Northwest region" pageId="4" pageNumber="196" specimenCount="1" stateProvince="Lai Chau" typeStatus="holotype">
<emphasis id="A7CA9B5FBE47C64F85D9B3048C23F994" bold="true" box="[151,304,1550,1576]" pageId="4" pageNumber="196">
<typeStatus id="4A05F9EFBE47C64F85D9B3048DC1F994" box="[151,210,1550,1576]" pageId="4" pageNumber="196">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="577A89AFBE47C64F8470B3048F02F994" box="[318,529,1550,1576]" country="Vietnam" name="Lai Chau" pageId="4" pageNumber="196">Lai Châu province</collectingRegion>
,
<location id="90611196BE47C64F8752B3048FF1F994" LSID="urn:lsid:plazi:treatment:1D17F65BBE47C64C85D9B0A88977FC19:90611196BE47C64F8752B3048FF1F994" box="[540,738,1550,1576]" country="Vietnam" name="Northwest region" pageId="4" pageNumber="196" stateProvince="Lai Chau">Northwest region</location>
,
<collectingCountry id="EDA907DDBE47C64F87A3B3048E43F994" box="[749,848,1550,1576]" name="Vietnam" pageId="4" pageNumber="196">Vietnam</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="9501474DBE47C64F8589B3388E66F929" blockId="4.[151,1437,1550,2008]" pageId="4" pageNumber="196">
<emphasis id="A7CA9B5FBE47C64F8589B3388C7EF9F0" bold="true" box="[199,365,1586,1612]" pageId="4" pageNumber="196">Type material</emphasis>
(
<specimenCount id="83B88CC4BE47C64F8433B3388CADF9F1" box="[381,446,1586,1613]" pageId="4" pageNumber="196" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="83B88CC4BE47C64F8484B3388F1BF9F1" box="[458,520,1586,1613]" pageId="4" pageNumber="196" type="female">3 ♀♀</specimenCount>
).
<emphasis id="A7CA9B5FBE47C64F8751B3388F9FF9F0" bold="true" box="[543,652,1586,1612]" pageId="4" pageNumber="196">
<typeStatus id="4A05F9EFBE47C64F8751B3388F9FF9F0" box="[543,652,1586,1612]" pageId="4" pageNumber="196" type="holotype">Holotype</typeStatus>
</emphasis>
,
<materialsCitation id="25D64D10BE47C64F87D7B3398E1EF9CC" ID-GBIF-Occurrence="3071291307" collectingDate="2020-06" collectionCode="BMNH" country="Vietnam" location="Northwest region" pageId="4" pageNumber="196" specimenCount="1" specimenCount-male="1" stateProvince="Lai Chau" typeStatus="holotype">
<specimenCount id="83B88CC4BE47C64F87D7B3398FBDF9F1" box="[665,686,1587,1613]" pageId="4" pageNumber="196" type="male"></specimenCount>
(
<collectionCode id="F3AFDF88BE47C64F87F3B3388E06F9F0" box="[701,789,1586,1612]" pageId="4" pageNumber="196">BMNH</collectionCode>
,
<figureCitation id="0D855BC8BE47C64F866EB3388E99F9F0" box="[800,906,1586,1612]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="4" pageNumber="196">Figs. 13</figureCitation>
,
<figureCitation id="0D855BC8BE47C64F86DBB3388ECCF9F1" box="[917,991,1586,1613]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="4" pageNumber="196">1618</figureCitation>
), labelled: “
<collectingCountry id="EDA907DDBE47C64F812DB33889D6F9F0" box="[1123,1221,1586,1612]" name="Vietnam" pageId="4" pageNumber="196">Vietnam</collectingCountry>
/
<location id="90611196BE47C64F8197B338888EF9F0" LSID="urn:lsid:plazi:treatment:1D17F65BBE47C64C85D9B0A88977FC19:90611196BE47C64F8197B338888EF9F0" box="[1241,1437,1586,1612]" country="Vietnam" name="Northwest region" pageId="4" pageNumber="196" stateProvince="Lai Chau">Northwest region</location>
/
<collectingRegion id="577A89AFBE47C64F85E6B35C8C6DF9CC" box="[168,382,1622,1648]" country="Vietnam" name="Lai Chau" pageId="4" pageNumber="196">Lai Châu province</collectingRegion>
/
<date id="E100618DBE47C64F84D7B35D8F1EF9CC" box="[409,525,1622,1649]" pageId="4" pageNumber="196" value="2020-06">
<collectingDate id="F1449865BE47C64F84D7B35D8F1EF9CC" box="[409,525,1622,1649]" pageId="4" pageNumber="196" value="2020-06">June 2020</collectingDate>
</date>
/ local collector leg.”
</materialsCitation>
.
<materialsCitation id="25D64D10BE47C64F8654B35C8E63F929" ID-GBIF-Occurrence="3071291309" collectingDate="2020-06" collectionCode="PMCH" country="Vietnam" location="Northwest region" pageId="4" pageNumber="196" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Lai Chau" typeStatus="paratype">
<emphasis id="A7CA9B5FBE47C64F8654B35C8E81F9CC" bold="true" box="[794,914,1622,1648]" pageId="4" pageNumber="196">
<typeStatus id="4A05F9EFBE47C64F8654B35C8E81F9CC" box="[794,914,1622,1648]" pageId="4" pageNumber="196" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="83B88CC4BE47C64F86EDB35C8EC2F9CD" box="[931,977,1622,1649]" pageId="4" pageNumber="196" type="male">1 ♂</specimenCount>
,
<specimenCount id="83B88CC4BE47C64F86AEB35C8932F9CD" box="[992,1057,1622,1649]" pageId="4" pageNumber="196" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="F3AFDF88BE47C64F817DB35C8999F9CC" box="[1075,1162,1622,1648]" pageId="4" pageNumber="196">PMCH</collectionCode>
): same label data as the holotype, one of the females was labelled as
<emphasis id="A7CA9B5FBE47C64F87C3B3708FFFF928" bold="true" box="[653,748,1658,1684]" pageId="4" pageNumber="196">
<typeStatus id="4A05F9EFBE47C64F87C3B3708FFFF928" box="[653,748,1658,1684]" pageId="4" pageNumber="196" type="allotype">allotype</typeStatus>
</emphasis>
(
<figureCitation id="0D855BC8BE47C64F87B5B3708E7BF929" box="[763,872,1658,1685]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="4" pageNumber="196">Figs. 46</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE47C64C8589B3948EEFFD81" lastPageId="7" lastPageNumber="199" pageId="4" pageNumber="196" type="description">
<paragraph id="9501474DBE47C64F8589B394891CF8BC" blockId="4.[151,1437,1550,2008]" pageId="4" pageNumber="196">
<emphasis id="A7CA9B5FBE47C64F8589B3948F53F904" bold="true" box="[199,576,1694,1720]" pageId="4" pageNumber="196">Description of holotype (male)</emphasis>
(
<figureCitation id="0D855BC8BE47C64F871AB3948FD6F904" box="[596,709,1694,1720]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="4" pageNumber="196">Figs. 13</figureCitation>
). Robust and convex body, somewhat elongate and narrowed apically, with rather short and stout legs; color bright metallic golden-green with strong reflection. Head, pronotum, elytra, ventrum and pygidium completely lacking any noticeable white macula.
</paragraph>
<paragraph id="9501474DBE47C64F8589B2008F3CF864" blockId="4.[151,1437,1550,2008]" pageId="4" pageNumber="196">
<emphasis id="A7CA9B5FBE47C64F8589B2008C17F898" box="[199,260,1802,1828]" italics="true" pageId="4" pageNumber="196">Head</emphasis>
. Metallic green with strong reflection and rather straight lateral declivity of clypeus. Apical margin of clypeus raised, almost straight with a slight shallow depression in the middle. Surface of frons and clypeus completely bare without setation, covered only with simple scattered punctation, with somewhat coarse and denser punctures on the sides of clypeus. Mentum and gula with dense, moderately long, fulvous setae. Antennae normal size, scapus and antennal club greenish, rest of antenomeres blackish. Inner lamellae of antennal club with one row of short, brush-like, fulvous bristles.
</paragraph>
<paragraph id="9501474DBE46C64E8589B59D8CC8FEA1" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="197">
<emphasis id="A7CA9B5FBE46C64E8589B59D8C25FF0C" box="[199,310,151,176]" italics="true" pageId="5" pageNumber="197">Pronotum</emphasis>
. Metallic green to golden coloured, with strong reflection. Obtusely bordered, convex, narrowed apically, widest at base. Antescutellar and posterolateral margins strongly sinuated. Disc, medial line and base of pronotum smooth, almost without punctation. Anterolateral margin very coarsely punctured, with scattered deep and wide rounded punctures.
</paragraph>
<paragraph id="9501474DBE46C64E8589B42D886CFEFD" blockId="5.[151,1436,151,466]" box="[199,1407,295,321]" pageId="5" pageNumber="197">
<taxonomicName id="52BE3CCEBE46C64E8589B42D8C26FEFD" box="[199,309,295,321]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="5" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="A7CA9B5FBE46C64E8589B42D8C26FEFD" box="[199,309,295,321]" italics="true" pageId="5" pageNumber="197">Scutellum</emphasis>
</taxonomicName>
. Elongated, sides slightly arcuate, apex rounded, surface bare, impunctate and strongly reflective.
</paragraph>
<paragraph id="9501474DBE46C64E8589B4418833FE6D" blockId="5.[151,1436,151,466]" pageId="5" pageNumber="197">
<emphasis id="A7CA9B5FBE46C64E8589B4418C1EFED9" box="[199,269,331,357]" italics="true" pageId="5" pageNumber="197">Elytra</emphasis>
. Metallic green, with strong golden reflection. Smooth, with very sparse scattered thin punctures and indistinctly marked striolate line beside sutural ridge in anterior third. Subhumeral emargination shallow, from there gently narrowing to elytral apex. Apex rounded, apical and humeral calli obtuse. Sutural ridge flat throughout its length, slightly elevated in posterior half, with obtuse termination (without pointed end). Macropterous.
</paragraph>
<caption id="C1C117C5BE46C64E85D9B2598C04F864" ID-DOI="http://doi.org/10.5281/zenodo.4671637" ID-Zenodo-Dep="4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="5" pageNumber="197" startId="5.[151,264,1875,1899]" targetBox="[162,1426,504,1824]" targetPageId="5">
<paragraph id="9501474DBE46C64E85D9B2598C04F864" blockId="5.[151,1437,1875,2008]" pageId="5" pageNumber="197">
<emphasis id="A7CA9B5FBE46C64E85D9B2598C40F8D7" bold="true" box="[151,339,1875,1899]" pageId="5" pageNumber="197">FIGURES 1624.</emphasis>
Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618,
<taxonomicName id="52BE3CCEBE46C64E81D1B25E8C41F82C" authority="Muckstein" authorityName="Muckstein" authorityYear="2021" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="dianae" status="sp. nov." subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE46C64E81D1B25E89EDF8D7" box="[1183,1278,1876,1899]" italics="true" pageId="5" pageNumber="197">Protaetia</emphasis>
(
<emphasis id="A7CA9B5FBE46C64E8045B25E8887F8D0" box="[1291,1428,1876,1900]" italics="true" pageId="5" pageNumber="197">Macroliocola</emphasis>
)
<emphasis id="A7CA9B5FBE46C64E85D9B2728DCFF82C" box="[151,220,1912,1936]" italics="true" pageId="5" pageNumber="197">dianae</emphasis>
Mückstein
</taxonomicName>
,
<emphasis id="A7CA9B5FBE46C64E8413B2728CCFF833" bold="true" box="[349,476,1911,1936]" pageId="5" pageNumber="197">
<taxonomicNameLabel id="BCF92624BE46C64E8413B2728CCFF833" box="[349,476,1911,1936]" pageId="5" pageNumber="197" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype; 1921,
<taxonomicName id="52BE3CCEBE46C64E87D0B272895AF82C" authority="(Nonfried, 1892)" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[670,1097,1912,1936]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="montana" subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE46C64E87D0B2728FEEF833" box="[670,765,1912,1935]" italics="true" pageId="5" pageNumber="197">Protaetia</emphasis>
(
<emphasis id="A7CA9B5FBE46C64E8640B2728E39F833" box="[782,810,1912,1935]" italics="true" pageId="5" pageNumber="197">M.</emphasis>
)
<emphasis id="A7CA9B5FBE46C64E8677B2728E81F833" box="[825,914,1912,1935]" italics="true" pageId="5" pageNumber="197">montana</emphasis>
(
<bibRefCitation id="F12F3ABCBE46C64E86EFB2728951F82C" author="Nonfried, A. F." box="[929,1090,1912,1936]" pageId="5" pageNumber="197" pagination="359 - 380" refId="ref4098" refString="Nonfried, A. F. (1892) Weitere Beitrage zur Kaferfauna von Sudasien und Neuguinea. Berliner Entomologische Zeitschrift, 36, 359 - 380. https: // doi. org / 10.1002 / mmnd. 47918920107" type="journal article" year="1892">Nonfried, 1892</bibRefCitation>
)
</taxonomicName>
, Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224,
<taxonomicName id="52BE3CCEBE46C64E848FB2968E96F808" authority="Yawata, 1941" authorityName=", Yawata" authorityYear="1941" box="[449,901,1948,1972]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="197" phylum="Arthropoda" rank="species" species="nigropurpurea" subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE46C64E848FB2968F33F80F" box="[449,544,1948,1971]" italics="true" pageId="5" pageNumber="197">Protaetia</emphasis>
(
<emphasis id="A7CA9B5FBE46C64E877EB2968F5EF80F" box="[560,589,1948,1971]" italics="true" pageId="5" pageNumber="197">M.</emphasis>
)
<emphasis id="A7CA9B5FBE46C64E8712B2968FFCF80F" box="[604,751,1948,1971]" italics="true" pageId="5" pageNumber="197">nigropurpurea</emphasis>
<bibRefCitation id="F12F3ABCBE46C64E87B8B2968E96F808" author="Yawata, H." box="[758,901,1948,1972]" pageId="5" pageNumber="197" pagination="114 - 126" refId="ref4508" refString="Yawata, H. (1941) Synonymic notes on Japanese species belonging to the genus Protaetia (II). Kontyu, 15 (3), 114 - 126." type="journal article" year="1941">Yawata, 1941</bibRefCitation>
</taxonomicName>
, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale.
</paragraph>
</caption>
<caption id="C1C117C5BE45C64D85D9B32F8874F9DE" ID-DOI="http://doi.org/10.5281/zenodo.4671639" ID-Zenodo-Dep="4671639" httpUri="https://zenodo.org/record/4671639/files/figure.png" pageId="6" pageNumber="198" startId="6.[151,264,1573,1597]" targetBox="[186,1387,222,1523]" targetPageId="6">
<paragraph id="9501474DBE45C64D85D9B32F8874F9DE" blockId="6.[151,1437,1573,1634]" pageId="6" pageNumber="198">
<emphasis id="A7CA9B5FBE45C64D85D9B32F8C40F981" bold="true" box="[151,339,1573,1597]" pageId="6" pageNumber="198">FIGURES 2529.</emphasis>
Habitus of
<taxonomicName id="52BE3CCEBE45C64D8488B32C8EF6F982" authority="(Alexis &amp; Delpont, 1998)" baseAuthorityName="Alexis &amp; Delpont" baseAuthorityYear="1998" box="[454,997,1574,1598]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="198" phylum="Arthropoda" rank="species" species="miharai" subGenus="Liocola">
<emphasis id="A7CA9B5FBE45C64D8488B32C8F36F981" box="[454,549,1574,1597]" italics="true" pageId="6" pageNumber="198">Protaetia</emphasis>
(
<emphasis id="A7CA9B5FBE45C64D877CB32C8F6CF982" box="[562,639,1574,1598]" italics="true" pageId="6" pageNumber="198">Liocola</emphasis>
)
<emphasis id="A7CA9B5FBE45C64D87C3B32C8FCEF982" box="[653,733,1574,1598]" italics="true" pageId="6" pageNumber="198">miharai</emphasis>
(
<bibRefCitation id="F12F3ABCBE45C64D87A7B32C8ECDF982" author="Alexis, R. &amp; Delpont, M." box="[745,990,1574,1598]" pageId="6" pageNumber="198" pagination="163 - 180" refId="ref3699" refString="Alexis, R. &amp; Delpont, M. (1998) Contribution a l'etude des Cetoniinae (Coleoptera, Scarabaeoidea, Cetoniidae). Lambillionea, 98 (2), 163 - 180." type="journal article" year="1998">Alexis &amp; Delpont, 1998</bibRefCitation>
)
</taxonomicName>
, male, Vietnam, Yen Bai province (PMCH). 25, dorsal view; 26, ventral view; 27, lateral view; 28, aedeagus in dorsal view; 29, aedeagus in lateral view. Not to scale.
</paragraph>
</caption>
<paragraph id="9501474DBE45C64D8589B39A89B3F916" blockId="6.[151,1437,1680,2030]" box="[199,1184,1680,1706]" pageId="6" pageNumber="198">
<emphasis id="A7CA9B5FBE45C64D8589B39A8C21F916" box="[199,306,1680,1706]" italics="true" pageId="6" pageNumber="198">Pygidium</emphasis>
. Metallic dark green, with a very fine wrinkled structure on the entire surface.
</paragraph>
<paragraph id="9501474DBE45C64D8589B3BE8DC1F81A" blockId="6.[151,1437,1680,2030]" pageId="6" pageNumber="198">
<emphasis id="A7CA9B5FBE45C64D8589B3BE8C60F972" box="[199,371,1716,1742]" italics="true" pageId="6" pageNumber="198">Ventral surface</emphasis>
. Metallic green, with strong golden reflection (
<figureCitation id="0D855BC8BE45C64D86DEB3BE8EC4F972" box="[912,983,1716,1742]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="6" pageNumber="198">Fig. 2</figureCitation>
). Disc of metasternal ventrites glabrous and impunctate, metasternal sides with sparse punctation of wide horseshoe-shaped punctures, merged posteriorly. Abdominal ventrites smooth, bare and impuctate; except for the two posterior ventrites bearing very fine punctation and striolation laterally. Mesometasternal process wider than long, smooth and bare, impunctate, with obtusely rounded apex (
<figureCitation id="0D855BC8BE45C64D840AB24E8C88F8E3" box="[324,411,1860,1887]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="6" pageNumber="198">Fig. 16</figureCitation>
). Metepimeron green, with golden-red lustre and very fine striolation. Mesepimeron and metepisternum the same colour, with a few larger punctures. Anal sternites with shallow depression medially (
<figureCitation id="0D855BC8BE45C64D802AB2628DD6F81A" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="6" pageNumber="198">Figs. 23</figureCitation>
).
</paragraph>
<paragraph id="9501474DBE45C64C8589B2BB8DC4FF69" blockId="6.[151,1437,1680,2030]" lastBlockId="7.[151,1437,151,933]" lastPageId="7" lastPageNumber="199" pageId="6" pageNumber="198">
<emphasis id="A7CA9B5FBE45C64D8589B2BB8DEFF876" box="[199,252,1969,1994]" italics="true" pageId="6" pageNumber="198">Legs</emphasis>
. Rather short and stout, colored metallic bronze to purpureous red with golden/green reflection. Probia tridentate in both sexes, protibial teeth well developed, sharp, larger in females. Position of protibial teeth equidistant in both sexes. Mesotibia and metatibia with one simple carina in posterior half and brush of fulvous setation on inner sides.
</paragraph>
<paragraph id="9501474DBE44C64C8589B5D58DE3FE35" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B5D58C27FF45" box="[199,308,223,249]" italics="true" pageId="7" pageNumber="199">Aedeagus</emphasis>
(
<figureCitation id="0D855BC8BE44C64C840BB5D58CDCFF45" box="[325,463,223,249]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="7" pageNumber="199">Figs. 1718</figureCitation>
). Parameres elongate, symmetrical, glabrous, shorter than phallobase. Inner paramere rims wide flat, widest in basal half, then slightly narrowing, narrowing more before apex and terminated widely and obtuse at apical end. Outer paramere rims widest in the basal part, gradually narrowing in basal half, then the sclerotized part continues to well-developed internal sac terminated with lobes densely covered with short fulvous bristles.
</paragraph>
<paragraph id="9501474DBE44C64C8589B4988F36FE49" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B4988CAAFE10" bold="true" box="[199,441,402,428]" pageId="7" pageNumber="199">Sexual dimorphism.</emphasis>
Female similar to male (
<figureCitation id="0D855BC8BE44C64C8793B4998E58FE12" box="[733,843,403,430]" captionStart="FIGURES 16" captionStartId="1.[151,264,1936,1960]" captionTargetBox="[203,1385,351,1885]" captionTargetId="figure-75@1.[151,1436,309,1912]" captionTargetPageId="1" captionText="FIGURES 16. Habitus of Protaetia (Macroliocola) dianae Mückstein, new species. 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, allotype, dorsal view; 5, allotype, ventral view; 6, allotype, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671623" httpUri="https://zenodo.org/record/4671623/files/figure.png" pageId="7" pageNumber="199">Figs. 16</figureCitation>
), but differing in the following characters: antennal clubs shorter; protibia slightly wider with teeth on the outer margin distinctly larger and pointed; anal sternites without shallow medial depression.
</paragraph>
<paragraph id="9501474DBE44C64C8589B4F58890FDA5" blockId="7.[151,1437,151,933]" box="[199,1411,511,537]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B4F58C6DFDA5" bold="true" box="[199,382,511,537]" pageId="7" pageNumber="199">Measurements.</emphasis>
Body length 23.826.1 mm (
<typeStatus id="4A05F9EFBE44C64C8788B4F58E39FDA5" box="[710,810,511,537]" pageId="7" pageNumber="199" type="holotype">holotype</typeStatus>
26.1 mm), width 12.413.7 mm (
<typeStatus id="4A05F9EFBE44C64C81EAB4F5881BFDA5" box="[1188,1288,511,537]" pageId="7" pageNumber="199" type="holotype">holotype</typeStatus>
13.7 mm).
</paragraph>
<paragraph id="9501474DBE44C64C8589B7288EEFFD81" blockId="7.[151,1437,151,933]" box="[199,1020,546,573]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B7288C5EFD80" bold="true" box="[199,333,546,572]" pageId="7" pageNumber="199">Variability.</emphasis>
The body reflections can vary from golden green to coppery.
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE44C64C8589B74C8F31FD4D" pageId="7" pageNumber="199" type="diagnosis">
<paragraph id="9501474DBE44C64C8589B74C8F31FD4D" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B74C8CDEFDDC" bold="true" box="[199,461,582,608]" pageId="7" pageNumber="199">Differential diagnosis.</emphasis>
The newly described species is very similar to
<taxonomicName id="52BE3CCEBE44C64C8690B74D8980FDDC" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[990,1171,583,609]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="montana" subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE44C64C8690B74D8EE0FDDC" box="[990,1011,583,608]" italics="true" pageId="7" pageNumber="199">P.</emphasis>
(
<emphasis id="A7CA9B5FBE44C64C814CB74D8932FDDC" box="[1026,1057,583,608]" italics="true" pageId="7" pageNumber="199">M.</emphasis>
)
<emphasis id="A7CA9B5FBE44C64C817FB74D8980FDDC" box="[1073,1171,583,608]" italics="true" pageId="7" pageNumber="199">montana</emphasis>
</taxonomicName>
(
<figureCitation id="0D855BC8BE44C64C81ECB74D881FFDDD" box="[1186,1292,583,609]" captionStart="FIGURES 79" captionStartId="3.[151,264,1504,1528]" captionTargetBox="[177,1406,652,1464]" captionTargetId="figure-234@3.[151,1436,633,1481]" captionTargetPageId="3" captionText="FIGURES 79. Habitus of Protaetia (Macroliocola) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái, (PMCH). 7 male, dorsal view; 8, male, ventral view; 9, male, lateral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671627" httpUri="https://zenodo.org/record/4671627/files/figure.png" pageId="7" pageNumber="199">Figs. 79</figureCitation>
) from which can be separated by the following characters: pronotal sides with large, coarse punctation in both sexes; shape of mesometasternal process more rounded, without significant angles (
<figureCitation id="0D855BC8BE44C64C86C5B7858EF2FD15" box="[907,993,655,681]" captionStart="FIGURES 1015" captionStartId="4.[151,264,1300,1324]" captionTargetBox="[173,1419,391,1265]" captionTargetId="figure-85@4.[151,1436,380,1276]" captionTargetPageId="4" captionText="FIGURES 1015. Male holotype of Protaetia (M.) nigropurpurea Yawata, 1941 (NSMT). 10, dorsal view; 11, ventral view; 12, lateral view; 13, aedeagus in dorsal view; 14, aedeagus in lateral view; 15, labels. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671629" httpUri="https://zenodo.org/record/4671629/files/figure.png" pageId="7" pageNumber="199">Figs 13</figureCitation>
,
<figureCitation id="0D855BC8BE44C64C86A3B785891FFD15" box="[1005,1036,655,681]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="7" pageNumber="199">19</figureCitation>
); metasternal sides only with sparse punctation of large horseshoe-shaped punctures, merged posteriorly (without any striolation); clearly different shape of parameres (
<figureCitation id="0D855BC8BE44C64C8476B7DD8F05FD4D" box="[312,534,727,753]" captionStart="FIGURES 1624" captionStartId="5.[151,264,1875,1899]" captionTargetBox="[162,1426,504,1824]" captionTargetId="figure-140@5.[151,1435,489,1852]" captionTargetPageId="5" captionText="FIGURES 1624. Shape of mesometasternal process and aedeagus (in lateral and dorsal view): 1618, Protaetia (Macroliocola) dianae Mückstein, new species, holotype; 1921, Protaetia (M.) montana (Nonfried, 1892), Vietnam, Yen Bai province, Mú Cang Chái (PMCH); 2224, Protaetia (M.) nigropurpurea Yawata, 1941, Taiwan, Taoyuan county, Lala Shan Mts. (PMCH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4671637" httpUri="https://zenodo.org/record/4671637/files/figure.png" pageId="7" pageNumber="199">Figs. 1718, 2021</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE44C64C8589B7F18CC3FC85" pageId="7" pageNumber="199" type="etymology">
<paragraph id="9501474DBE44C64C8589B7F18CC3FC85" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B7F18C5DFCA9" bold="true" box="[199,334,763,789]" pageId="7" pageNumber="199">Etymology.</emphasis>
Matronymic, this new species is dedicated to my wife Diana, in thanks for her great tolerance of my entomological passions.
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE44C64C8589B6488CDEFCE1" box="[199,461,834,861]" pageId="7" pageNumber="199" type="distribution">
<paragraph id="9501474DBE44C64C8589B6488CDEFCE1" blockId="7.[151,1437,151,933]" box="[199,461,834,861]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B6488C72FCE0" bold="true" box="[199,353,834,860]" pageId="7" pageNumber="199">Distribution.</emphasis>
<collectingCountry id="EDA907DDBE44C64C8429B6498CD9FCE1" box="[359,458,835,861]" name="Vietnam" pageId="7" pageNumber="199">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DDA414C6BE44C64C8589B66D8977FC19" pageId="7" pageNumber="199" type="discussion">
<paragraph id="9501474DBE44C64C8589B66D8977FC19" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="199">
<emphasis id="A7CA9B5FBE44C64C8589B66D8C28FC3D" bold="true" box="[199,315,871,897]" pageId="7" pageNumber="199">Remarks.</emphasis>
Based on morphological characters and the shape of the male genitalia,
<taxonomicName id="52BE3CCEBE44C64C813BB66D8873FC3D" authority="Muckstein" authorityName="Muckstein" authorityYear="2021" box="[1141,1376,871,897]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="199" phylum="Arthropoda" rank="species" species="dianae" status="sp. nov.">
<emphasis id="A7CA9B5FBE44C64C813BB66D89CDFC3D" box="[1141,1246,871,897]" italics="true" pageId="7" pageNumber="199">P. dianae</emphasis>
Mückstein
</taxonomicName>
,
<emphasis id="A7CA9B5FBE44C64C8022B66D8DFAFC18" bold="true" pageId="7" pageNumber="199">
<taxonomicNameLabel id="BCF92624BE44C64C8022B66D8DFAFC18" pageId="7" pageNumber="199" rank="species">new species</taxonomicNameLabel>
</emphasis>
is classified in the subgenus
<taxonomicName id="52BE3CCEBE44C64C877EB6818973FC19" authority="Alexis &amp; Delpont, 1998" authorityName="Alexis &amp; Delpont" authorityYear="1998" box="[560,1120,907,933]" class="Insecta" family="Cetoniidae" genus="Protaetia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="199" phylum="Arthropoda" rank="subGenus" subGenus="Macroliocola">
<emphasis id="A7CA9B5FBE44C64C877EB6818F8AFC18" box="[560,665,907,932]" italics="true" pageId="7" pageNumber="199">Protaetia</emphasis>
(
<emphasis id="A7CA9B5FBE44C64C87E7B6818E53FC19" box="[681,832,907,933]" italics="true" pageId="7" pageNumber="199">Macroliocola</emphasis>
)
<bibRefCitation id="F12F3ABCBE44C64C8601B6818973FC19" author="Alexis, R. &amp; Delpont, M." box="[847,1120,907,933]" pageId="7" pageNumber="199" pagination="163 - 180" refId="ref3699" refString="Alexis, R. &amp; Delpont, M. (1998) Contribution a l'etude des Cetoniinae (Coleoptera, Scarabaeoidea, Cetoniidae). Lambillionea, 98 (2), 163 - 180." type="journal article" year="1998">Alexis &amp; Delpont, 1998</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>