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<document id="8FEFE8FC0F498D75B6F77EB6D22DCC17" ID-DOI="10.26107/RBZ-2020-0001" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577165" ID-ZooBank="4A76F694-DD52-48CC-A0D4-C1AD9D33B0A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784698889" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Reischütz, Alexander, Maassen, Wim J. M., Grego, Jozef &amp; Hunyadi, András" docDate="2020" docId="03EE1927C507140DFFA6F938FEF61B1D" docLanguage="en" docName="RafflesBZool.68.1-13.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Sinoennea angustistoma Pall-Gergely, A. Reischutz &amp; Maassen in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi 2020, new species" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="FFD7615FC5041409FFDAFFC0FF961E28" masterDocTitle="New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1698878575407" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5F54D6F3163A0167D3592373DF739B68" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DCADE068E9B5A37E84DA16A59384364">
<mods:title id="46F5B485A5A3A5A2F54ED718A3C1FB50">New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)</mods:title>
</mods:titleInfo>
<mods:name id="E110F947A5F6FDF8C4DED2D541011F2F" type="personal">
<mods:role id="208C857BA85730B1237AD015B02EA12B">
<mods:roleTerm id="C728EC835058F56CE07D3A5EC00172A4">Author</mods:roleTerm>
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<mods:namePart id="2370F3532C37AE3FDB30954C56989742">Páll-Gergely, Barna</mods:namePart>
</mods:name>
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<mods:namePart id="37E7EE94B26AC0FF64EC33488D9C97E2">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="C2665C5F705F7B8F509EEB932304FFA7" type="personal">
<mods:role id="D1D05264955B2D5FCA4BB0D066D7CDDE">
<mods:roleTerm id="3E783876FFE3E34FAD6D4A1A36E6E8C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD72765A9898831EDB5435F2DDF262AF">Maassen, Wim J. M.</mods:namePart>
</mods:name>
<mods:name id="94C3D88BEBE49E6584A05C73473401F6" type="personal">
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<mods:namePart id="900159143E6A3DB2DAA35AC4C8F2D619">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="714E5D85A214A0C2C72CAFDCFBFF485E" type="personal">
<mods:role id="65994448BE73DF6E2F0A57628D2AD0B3">
<mods:roleTerm id="48A90E715CE774EB55BB7C1D08628D5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="87A6C417E63686FD166B3E6F3B26F5FC">Hunyadi, András</mods:namePart>
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<mods:title id="FEDF0D1D4D51FCBD5A80FC78103EB90D">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="8757BA4EB74E7FE26906A02E4B5708D8">2020</mods:date>
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<mods:number id="35E164C779728F8448DC47E11C0E774D">2020-01-15</mods:number>
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<mods:number id="9713FEF58645E06AD7023221A90D22CA">68</mods:number>
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<mods:identifier id="FF5FD71DE48A275D79A52B05D8B12988" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03EE1927C507140DFFA6F938FEF61B1D" LSID="urn:lsid:plazi:treatment:03EE1927C507140DFFA6F938FEF61B1D" httpUri="http://treatment.plazi.org/id/03EE1927C507140DFFA6F938FEF61B1D" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="C35DFBBAC507140AFFA6F938FDB91907" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BF8A831C507140AFFA6F938FDB91907" blockId="3.[124,741,1783,1872]" pageId="3" pageNumber="4">
<heading id="D0B01F5DC507140AFFA6F938FDB91907" bold="true" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<emphasis id="B9337423C507140AFFA6F938FDB91907" bold="true" pageId="3" pageNumber="4">
<taxonomicName id="4C47D3B2C507140AFFA6F938FE0A1918" authority="Pall-Gergely, A. Reischutz &amp; Maassen" authorityName="Pall-Gergely, A. Reischutz &amp; Maassen in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="angustistoma" status="sp. nov.">
<emphasis id="B9337423C507140AFFA6F938FE1B1938" bold="true" box="[124,397,1784,1808]" italics="true" pageId="3" pageNumber="4">Sinoennea angustistoma</emphasis>
Páll-Gergely, A. Reischütz &amp; Maassen
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C507140AFE70F8D8FDB91907" box="[426,559,1815,1840]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35DFBBAC507140AFEB9F8F8FE691978" box="[355,511,1848,1872]" pageId="3" pageNumber="4" type="description">
<paragraph id="8BF8A831C507140AFEB9F8F8FE691978" blockId="3.[124,741,1783,1872]" box="[355,511,1848,1872]" pageId="3" pageNumber="4">
(
<figureCitation id="137CB4B4C507140AFEB1F8F8FE5D1978" box="[363,459,1848,1872]" captionStart="Fig" captionStartId="3.[113,148,1550,1572]" captionTargetBox="[113,1451,189,1534]" captionTargetId="figure-11@3.[113,1451,189,1534]" captionTargetPageId="3" captionText="Fig. 2. New Sinoennea Kobelt, 1904, species from Laos. A, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype); B, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577171" httpUri="https://zenodo.org/record/4577171/files/figure.png" pageId="3" pageNumber="4">Figs. 2B</figureCitation>
,
<figureCitation id="137CB4B4C507140AFE02F8F8FE611978" box="[472,503,1848,1872]" captionStart="Fig" captionStartId="10.[135,170,1193,1215]" captionTargetBox="[135,1473,190,1177]" captionTargetId="figure-112@10.[135,1473,190,1177]" captionTargetPageId="10" captionText="Fig. 6. Synoptic view of Sinoennea and Laoennea, new genus, species of Laos (all species shown) and northern Thailand. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species; B, Sinoennea infantilis Páll-Gergely &amp; Grego, new species; CE, Sinoennea variabilis Páll-Gergely &amp; Grego, new species; F, Sinoennea sutchariti Páll-Gergely &amp; Hunyadi, new species; G, Sinoennea panhai Páll-Gergely &amp; Hunyadi, new species; H, Sinoennea euryomphala Inkhavilay &amp; Panha, 2016; I, Sinoennea ljudmilena Páll-Gergely, new species; J, Sinoennea lizae Maassen, 2008; K, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species; L, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577179" httpUri="https://zenodo.org/record/4577179/files/figure.png" pageId="3" pageNumber="4">6L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC507140AFFABF8B8FAA71898" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BF8A831C507140AFFABF8B8FD661638" blockId="3.[113,753,1911,2064]" pageId="3" pageNumber="4">
<emphasis id="B9337423C507140AFFABF8B8FEB619A7" bold="true" box="[113,288,1911,1936]" pageId="3" pageNumber="4">Type material.</emphasis>
<materialsCitation id="3B2FA26CC507140AFEF0F8B8FD661638" collectionCode="H, D" country="Laos" elevation="300" latitude="17.5834" location="Gnommalat" longLatPrecision="1" longitude="105.17405" pageId="3" pageNumber="4" specimenCode="NHMW 113108" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="54FC1693C507140AFEF0F8B8FE1919B8" box="[298,399,1912,1936]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
(1 shell,
<collectionCode id="ED5630F4C507140AFDDAF8B8FD8119A7" box="[512,535,1912,1935]" pageId="3" pageNumber="4">H</collectionCode>
:
<quantity id="4CBF05D4C507140AFDFCF8B8FD1B19A7" box="[550,653,1912,1936]" metricMagnitude="-3" metricUnit="m" metricValue="2.91" pageId="3" pageNumber="4" unit="mm" value="2.91">2.91 mm</quantity>
,
<collectionCode id="ED5630F4C507140AFD41F8B8FD2419A7" box="[667,690,1912,1935]" pageId="3" pageNumber="4">D</collectionCode>
:
<quantity id="4CBF05D4C507140AFD1BF8B8FF361987" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="3" pageNumber="4" unit="mm" value="1.68">1.68 mm</quantity>
;
<figureCitation id="137CB4B4C507140AFF6BF858FE9A1998" box="[177,268,1944,1968]" captionStart="Fig" captionStartId="3.[113,148,1550,1572]" captionTargetBox="[113,1451,189,1534]" captionTargetId="figure-11@3.[113,1451,189,1534]" captionTargetPageId="3" captionText="Fig. 2. New Sinoennea Kobelt, 1904, species from Laos. A, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype); B, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species (holotype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577171" httpUri="https://zenodo.org/record/4577171/files/figure.png" pageId="3" pageNumber="4">Fig. 2B</figureCitation>
) (
<specimenCode id="DBE1004AC507140AFEF2F858FE7B1998" box="[296,493,1944,1968]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="4" type="Museum">NHMW 113108</specimenCode>
),
<collectingCountry id="F350E8A1C507140AFDDCF858FDD71987" box="[518,577,1944,1967]" name="Laos" pageId="3" pageNumber="4">Laos</collectingCountry>
,
<collectingRegion id="498366D3C507140AFD8BF858FF4019F8" country="Laos" name="Khammouan" pageId="3" pageNumber="4">Khammouane Province</collectingRegion>
, limestone hill in the rice paddies E of road 1E about
<quantity id="4CBF05D4C507140AFF62F818FE9719C7" box="[184,257,2008,2032]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="m" value="300.0">
<elevation id="006A4F02C507140AFF62F818FE9719C7" box="[184,257,2008,2032]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="m" value="300.0">300 m</elevation>
</quantity>
SE of
<location id="8E98FEEAC507140AFE8CF818FE4F19D8" LSID="urn:lsid:plazi:treatment:03EE1927C507140DFFA6F938FEF61B1D:8E98FEEAC507140AFE8CF818FE4F19D8" box="[342,473,2008,2032]" country="Laos" latitude="17.5834" longLatPrecision="1" longitude="105.17405" name="Gnommalat" pageId="3" pageNumber="4" stateProvince="Khammouan">Gnommalat</location>
,
<locationDeviation id="9A4AB113C507140AFE32F818FD0F19D8" box="[488,665,2008,2032]" location="Thakhek" pageId="3" pageNumber="4">NE of Thakhek</locationDeviation>
,
<elevation id="006A4F02C507140AFD72F818FF321638" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="4" unit="m" value="175.0">
<quantity id="4CBF05D4C507140AFD72F818FD7919C7" box="[680,751,2008,2032]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="3" pageNumber="4" unit="m" value="175.0">175 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE73CEF6C507140AFF69F836FEAA1627" box="[179,316,2038,2063]" degrees="17" direction="north" minutes="35.004" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="17.5834">17°35.004N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C507140AFE90F836FE481627" box="[330,478,2038,2063]" degrees="105" direction="east" minutes="10.443" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="105.17405">105°10.443E</geoCoordinate>
, at the base of limestone
</materialsCitation>
</paragraph>
<paragraph id="8BF8A831C507140AFCF7F9B8FAA71898" blockId="3.[813,1451,1656,1712]" pageId="3" pageNumber="4">
rocks, coll. A. Reischütz,
<date id="FFF98EF1C507140AFB8AF9B8FB4F18B8" box="[1104,1241,1656,1680]" pageId="3" pageNumber="4" value="2010-03">March 2010</date>
;
<materialsCitation id="3B2FA26CC507140AFB3DF9B8FAB81898" collectionCode="H" country="Laos" elevation="300" latitude="17.5834" location="Gnommalat" longLatPrecision="1" longitude="105.17405" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Khammouan" typeStatus="paratype">
<specimenCount id="9D4163B8C507140AFB3DF9B8FACC18B8" box="[1255,1370,1656,1680]" pageId="3" pageNumber="4" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(RE/1,
<collectionCode id="ED5630F4C507140AFCF7F958FCD21887" box="[813,836,1688,1711]" pageId="3" pageNumber="4">H</collectionCode>
:
<quantity id="4CBF05D4C507140AFC8AF958FC251887" box="[848,947,1688,1712]" metricMagnitude="-3" metricUnit="m" metricValue="3.02" pageId="3" pageNumber="4" unit="mm" value="3.02">3.02 mm</quantity>
,
<quantity id="4CBF05D4C507140AFC1AF958FBB21898" box="[960,1060,1688,1712]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="4" unit="mm" value="1.72">1.72 mm</quantity>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC507140AFCF7F917FAC91978" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BF8A831C507140AFCF7F917FAC91978" blockId="3.[813,1452,1751,1872]" pageId="3" pageNumber="4">
<emphasis id="B9337423C507140AFCF7F917FC3018C7" bold="true" box="[813,934,1751,1775]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
Shell ovoid with a conspicuously narrowed, elongate aperture and a deep, wide, internally grooved umbilicus. Aperture with short parietal lamella, and three denticles each on both palatal and columellar sides.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC507140DFCF7F8B7FD631D7E" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="8BF8A831C507140DFCF7F8B7FD631D7E" blockId="3.[811,1453,1911,2064]" lastBlockId="4.[135,776,189,854]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="B9337423C507140AFCF7F8B7FC2119A7" bold="true" box="[813,951,1911,1935]" pageId="3" pageNumber="4">Description.</emphasis>
Shell chalk white, probably slightly transparent when fresh. Shell ovoid with strongly sideways flattened aperture. Last and penultimate whorls are the widest from standard apertural view. Entire shell with 5.255.5 whorls. Protoconch consisting of two whorls, moderately glossy, very finely pitted. Teleoconch regularly ribbed. Ribs of upper whorls curved or slightly S-shaped for last whorl. Aperture elongate ovate, strongly compressed from both sides. Parietal callus is smeared onto preceding whorl. Apertural dentition seven-fold. Parietal lamella elevated, very short, rather straight, its outer end slightly turning to the palatal side, and its inner end slightly bent in direction of the columella. Sinulus relatively small, teardrop-shaped, opens slightly upwards. Palatal wall with three denticles, uppermost one small, pointed, facing parietal lamella, middle one slightly elongate inside, somewhat larger than first one, third one also slightly elongate obliquely in basal direction. Columellar wall also with three denticles, uppermost tiny, blunt, situated at the middle of parietal lamella, lower two ones situated along an inwards running ridge, lower one situated closer to the peristome edge. Both denticles slightly elongated inside. Outer side of body whorl slightly impressed in the position of the innermost palatal denticle, no swelling visible behind peristome. Umbilicus wide, rounded, showing all whorls, periumbilical keel prominent, sharp. Inner side of body whorl (i.e., inside umbilicus) with a conspicuous groove.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC500140DFF52FCBDFECC1AFD" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BF8A831C500140DFF52FCBDFECC1AFD" blockId="4.[134,775,893,1237]" pageId="4" pageNumber="5">
<emphasis id="B9337423C500140DFF52FCBDFEEF1DBD" bold="true" box="[136,377,893,917]" pageId="4" pageNumber="5">Differential diagnosis.</emphasis>
<taxonomicName id="4C47D3B2C500140DFEA4FCBEFDEF1DBD" authority="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi, 2020" authorityName="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi" authorityYear="2020" box="[382,633,894,917]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="angustistoma" status="sp. nov.">
<emphasis id="B9337423C500140DFEA4FCBEFDEF1DBD" box="[382,633,894,917]" italics="true" pageId="4" pageNumber="5">Sinoennea angustistoma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C500140DFD5FFCBEFC951DBD" box="[645,771,893,917]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
, is similar to
<emphasis id="B9337423C500140DFED9FC5EFE6B1D9D" box="[259,509,926,949]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="4C47D3B2C500140DFED9FC5EFEE61D9D" authorityName="Kobelt" authorityYear="1904" box="[259,368,926,949]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">Sinoennea</taxonomicName>
guiyangensis
</emphasis>
T.-C. Luo, D.-N. Chen &amp; G.-Q. Zhang, 1998, from
<collectingRegion id="498366D3C500140DFE48FC7DFDC61DFD" box="[402,592,957,981]" country="China" name="Guizhou" pageId="4" pageNumber="5">Guizhou Province</collectingRegion>
of
<collectingCountry id="F350E8A1C500140DFDABFC7DFD251DFD" box="[625,691,957,981]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
, but has a more rounded shell, denser ribs, narrower aperture, and an additional columellar denticle on the peristome. Furthermore, the umbilicus of the new species is probably wider than that of
<emphasis id="B9337423C500140DFF73FBFEFEC51A7D" box="[169,339,1086,1109]" italics="true" pageId="4" pageNumber="5">S. guiyangensis</emphasis>
, but it is not illustrated in the original description of that species.
<emphasis id="B9337423C500140DFE1AFB9EFD551A5D" box="[448,707,1117,1141]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="4C47D3B2C500140DFE1AFB9EFDA71A5D" authorityName="Kobelt" authorityYear="1904" box="[448,561,1118,1141]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">Sinoennea</taxonomicName>
maolanensis
</emphasis>
T.-C. Luo, W.-C. Zhou &amp; D.-N. Chen, 2004, also from
<collectingRegion id="498366D3C500140DFD7EFBBDFC951ABD" box="[676,771,1149,1173]" country="China" name="Guizhou" pageId="4" pageNumber="5">Guizhou</collectingRegion>
, has a less rounded shell, a narrower umbilicus, and more widely-spaced ribs.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC500140DFF52FB3DFEF61B1D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BF8A831C500140DFF52FB3DFEF61B1D" blockId="4.[136,775,1277,1333]" pageId="4" pageNumber="5">
<emphasis id="B9337423C500140DFF52FB3DFE9F1B3D" bold="true" box="[136,265,1277,1301]" pageId="4" pageNumber="5">Etymology.</emphasis>
The specific epithet refers to the narrow aperture of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>