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<document id="7ABA55780B22A52957958A4E2363AB49" ID-DOI="10.26107/RBZ-2020-0001" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577165" ID-ZooBank="4A76F694-DD52-48CC-A0D4-C1AD9D33B0A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784698889" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Reischütz, Alexander, Maassen, Wim J. M., Grego, Jozef &amp; Hunyadi, András" docDate="2020" docId="03EE1927C501140FFF7AFA0CFE851DFD" docLanguage="en" docName="RafflesBZool.68.1-13.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Sinoennea ljudmilena Pall-Gergely in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi 2020, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="FFD7615FC5041409FFDAFFC0FF961E28" masterDocTitle="New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="6" updateTime="1698878575407" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BA0BED549E174D3D5E500D182A268A61" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="149B75C67D0A310F5D954CF24BC9ACF3">
<mods:title id="3487D649E49B2CD6822AD18E93484B15">New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)</mods:title>
</mods:titleInfo>
<mods:name id="289E97661DEDD4D988D88F4DF2B1F3E8" type="personal">
<mods:role id="BB136C9BB251058719D3949814D33B07">
<mods:roleTerm id="C8C80A6B3DDBADEB5A314A0252FC8F07">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DCC80666F8743D10657FDD1885F958D0">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="7ABEB530A64CDB21BCF0C80056DE07B0" type="personal">
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<mods:namePart id="874E8105F37EF3850CDAC2F0123FD47B">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="09FB4508EE9EA2CF72238FF96DB99400" type="personal">
<mods:role id="913849B8DE332582FC97855D8460271F">
<mods:roleTerm id="382D11F3BB8D8325038ACA15A20DBC10">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB85B9C35B5C5EE59BA8996D57C5F93B">Maassen, Wim J. M.</mods:namePart>
</mods:name>
<mods:name id="E602D2565D9D3A4AEBA85BE431AB4B61" type="personal">
<mods:role id="8B5C014A34D6A648C18E655971358B56">
<mods:roleTerm id="ECC1ADB7C9E906149E63890E0CAC41A8">Author</mods:roleTerm>
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<mods:namePart id="217113CCD329DF31819466769A246AEF">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="A554ADF96E7AD2A2F8590BF4997083BF" type="personal">
<mods:role id="9E7E00EAEFA5433E58CD4ECFC5CBCF5A">
<mods:roleTerm id="41FF7DD61D2F5D869BB10227B52540AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF795CDE4753AD89FAEF08A9105B2925">Hunyadi, András</mods:namePart>
</mods:name>
<mods:typeOfResource id="424D0788BDF8AFA9565617391CD6D595">text</mods:typeOfResource>
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<mods:title id="0532F1490E76B4FF0E1A5973F94DE308">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="4536CF97AC8688FD33B3100D1E6E3C24">
<mods:date id="1E81042164A70A36813C642C068603F9">2020</mods:date>
<mods:detail id="2F4E0996E726D6C10DAE4FF5BFCC194D" type="pubDate">
<mods:number id="032A08F17F323C6E85F2083C75392C57">2020-01-15</mods:number>
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<mods:detail id="BB44E2F31555DC1E4C240170BA4F36A2" type="volume">
<mods:number id="2FAED30034D34E9EBC00B1EAE0C56F8C">68</mods:number>
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<mods:classification id="AC038C824AD0F9A5AB07950FDBEFFCBE">journal article</mods:classification>
<mods:identifier id="2DF610738F577C125F0D46A2F5E1FE3D" type="DOI">10.26107/RBZ-2020-0001</mods:identifier>
<mods:identifier id="7896D8776E36635A6EFC9D00899B97B2" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="33ABFE9105FA46A0DE10746B6C2E4638" type="Zenodo-Dep">4577165</mods:identifier>
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<treatment id="03EE1927C501140FFF7AFA0CFE851DFD" LSID="urn:lsid:plazi:treatment:03EE1927C501140FFF7AFA0CFE851DFD" httpUri="http://treatment.plazi.org/id/03EE1927C501140FFF7AFA0CFE851DFD" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="C35DFBBAC501140CFF7AFA0CFD541BCC" box="[160,706,1484,1509]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BF8A831C501140CFF7AFA0CFD541BCC" blockId="5.[160,706,1484,1541]" box="[160,706,1484,1509]" pageId="5" pageNumber="6">
<heading id="D0B01F5DC501140CFF7AFA0CFD541BCC" bold="true" box="[160,706,1484,1509]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="6" reason="2">
<emphasis id="B9337423C501140CFF7AFA0CFD541BCC" bold="true" box="[160,706,1484,1509]" pageId="5" pageNumber="6">
<taxonomicName id="4C47D3B2C501140CFF7AFA0CFDB81BCD" authority="Pall-Gergely" authorityName="Pall-Gergely in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" box="[160,558,1484,1509]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="ljudmilena" status="sp. nov.">
<emphasis id="B9337423C501140CFF7AFA0CFE031BCC" bold="true" box="[160,405,1484,1508]" italics="true" pageId="5" pageNumber="6">Sinoennea ljudmilena</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C501140CFDE6FA0DFD541BCC" box="[572,706,1484,1509]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC501140CFEAAFA2DFE64182C" box="[368,498,1516,1541]" pageId="5" pageNumber="6" type="description">
<paragraph id="8BF8A831C501140CFEAAFA2DFE64182C" blockId="5.[160,706,1484,1541]" box="[368,498,1516,1541]" pageId="5" pageNumber="6">
(
<figureCitation id="137CB4B4C501140CFEA2FA2DFE53182D" box="[376,453,1517,1541]" captionStart="Fig" captionStartId="5.[113,148,1215,1237]" captionTargetBox="[113,1451,189,1200]" captionTargetId="figure-12@5.[113,1451,189,1203]" captionTargetPageId="5" captionText="Fig. 3. A, Sinoennea ljudmilena Páll-Gergely, new species (holotype); B, C, paratypes. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577173" httpUri="https://zenodo.org/record/4577173/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
,
<figureCitation id="137CB4B4C501140CFE09FA2CFE7F182C" box="[467,489,1516,1540]" captionStart="Fig" captionStartId="10.[135,170,1193,1215]" captionTargetBox="[135,1473,190,1177]" captionTargetId="figure-112@10.[135,1473,190,1177]" captionTargetPageId="10" captionText="Fig. 6. Synoptic view of Sinoennea and Laoennea, new genus, species of Laos (all species shown) and northern Thailand. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species; B, Sinoennea infantilis Páll-Gergely &amp; Grego, new species; C–E, Sinoennea variabilis Páll-Gergely &amp; Grego, new species; F, Sinoennea sutchariti Páll-Gergely &amp; Hunyadi, new species; G, Sinoennea panhai Páll-Gergely &amp; Hunyadi, new species; H, Sinoennea euryomphala Inkhavilay &amp; Panha, 2016; I, Sinoennea ljudmilena Páll-Gergely, new species; J, Sinoennea lizae Maassen, 2008; K, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species; L, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577179" httpUri="https://zenodo.org/record/4577179/files/figure.png" pageId="5" pageNumber="6">6I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC501140CFFABF9EDFDBA188D" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BF8A831C501140CFFABF9EDFDBA188D" blockId="5.[113,753,1580,1701]" pageId="5" pageNumber="6">
<emphasis id="B9337423C501140CFFABF9EDFEB6186C" bold="true" box="[113,288,1580,1605]" pageId="5" pageNumber="6">Type material.</emphasis>
<materialsCitation id="3B2FA26CC501140CFEF0F9EDFDBA188D" collectionCode="H, D, MNHN-" collectorName="Province du Tran Ninh &amp; Saurin" country="Laos" pageId="5" pageNumber="6" specimenCount="19" typeStatus="holotype">
<typeStatus id="54FC1693C501140CFEF0F9EDFE19186D" box="[298,399,1581,1605]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
(1 shell,
<collectionCode id="ED5630F4C501140CFDDAF9EDFD81186C" box="[512,535,1581,1604]" pageId="5" pageNumber="6">H</collectionCode>
:
<quantity id="4CBF05D4C501140CFDFCF9EDFD1B186C" box="[550,653,1581,1605]" metricMagnitude="-3" metricUnit="m" metricValue="4.47" pageId="5" pageNumber="6" unit="mm" value="4.47">4.47 mm</quantity>
,
<collectionCode id="ED5630F4C501140CFD41F9EDFD24186C" box="[667,690,1581,1604]" pageId="5" pageNumber="6">D</collectionCode>
:
<quantity id="4CBF05D4C501140CFD1BF9ECFF09184C" metricMagnitude="-3" metricUnit="m" metricValue="2.56" pageId="5" pageNumber="6" unit="mm" value="2.56">2.56 mm</quantity>
;
<figureCitation id="137CB4B4C501140CFF75F98DFF60184D" box="[175,246,1613,1637]" captionStart="Fig" captionStartId="5.[113,148,1215,1237]" captionTargetBox="[113,1451,189,1200]" captionTargetId="figure-12@5.[113,1451,189,1203]" captionTargetPageId="5" captionText="Fig. 3. A, Sinoennea ljudmilena Páll-Gergely, new species (holotype); B, C, paratypes. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577173" httpUri="https://zenodo.org/record/4577173/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) (MNHN-IM-2012-27277),
<collectingCountry id="F350E8A1C501140CFD98F98DFDED184C" box="[578,635,1613,1636]" name="Laos" pageId="5" pageNumber="6">Laos</collectingCountry>
, Pah Hia,
<collectorName id="26B2CDE7C501140CFFA8F9ADFEFD18AD" box="[114,363,1645,1669]" pageId="5" pageNumber="6">Province du Tran Ninh</collectorName>
, coll.
<collectorName id="26B2CDE7C501140CFE71F9ADFE6218AD" box="[427,500,1645,1669]" pageId="5" pageNumber="6">Saurin</collectorName>
;
<specimenCount id="9D4163B8C501140CFE25F9ADFD1118AD" box="[511,647,1645,1669]" pageId="5" pageNumber="6" type="generic" typeStatus="paratypes">19 paratypes</specimenCount>
(
<collectionCode id="ED5630F4C501140CFD4FF9ADFD6718AD" box="[661,753,1645,1669]" pageId="5" pageNumber="6">MNHN-</collectionCode>
IM-2012-27278), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC501140CFFABF90CFD86190D" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BF8A831C501140CFFABF90CFD86190D" blockId="5.[113,752,1740,1829]" pageId="5" pageNumber="6">
<emphasis id="B9337423C501140CFFABF90CFF7C18CC" bold="true" box="[113,234,1740,1764]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
A large, glossy species with knots along the suture in the last three whorls, and an ovoid aperture with a very strong, curved parietal lamella.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC501140FFFABF88CFDC81F5D" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8BF8A831C501140FFFABF88CFE7B1F3D" blockId="5.[113,753,1868,2085]" lastBlockId="6.[136,776,189,277]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B9337423C501140CFFABF88CFE90194C" bold="true" box="[113,262,1868,1892]" pageId="5" pageNumber="6">Description.</emphasis>
Shell yellowish, ovoid. Some specimens nearly cylindrical with domed upper side. Whorls of several specimens grow irregularly resulting in a roof tile-like structure (
<figureCitation id="137CB4B4C501140CFF07F86DFEA419ED" box="[221,306,1965,1989]" captionStart="Fig" captionStartId="5.[113,148,1215,1237]" captionTargetBox="[113,1451,189,1200]" captionTargetId="figure-12@5.[113,1451,189,1203]" captionTargetPageId="5" captionText="Fig. 3. A, Sinoennea ljudmilena Páll-Gergely, new species (holotype); B, C, paratypes. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577173" httpUri="https://zenodo.org/record/4577173/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
). Penultimate or antepenultimate whorl is widest from standard apertural view. Entire shell consisting of 7–7.75 whorls. Protoconch consisting of 1.5–1.75 whorls, glossy, very finely pitted, sometimes with traces of inconspicuous radial wrinkles. Teleoconch with irregular sculpture. Initial whorls usually with full ribs extending from suture to suture, last ca. three whorls (cylindrical part of shell) glossy, with remains of ribs (knots) near upper suture, or very rarely with a few full ribs, which are probably indications of juvenile peristomes. Last half whorl again with full ribs. Individual ribs look like wrinkles or waves created by pushing the shell surface in upper (towards protoconch) direction. Aperture suboval to rectangular. Parietal callus rounded, reaches ca. half of preceding whorl, thin, “smeared” onto the preceding whorl. Apertural dentition five-fold, closes passage so effectively that it is hard to see inside aperture even by viewing obliquely. Parietal lamella very strong, elevated, outer portion strongly oblique, bent in direction of columella. Inner part of parietal lamella lower, abruptly bent in the other direction. Sinulus relatively small, strongly isolated (i.e., parietal lamella and upper palatal tooth nearly reaching each other), opens somewhat laterally. Three palatal denticles pointed, and of comparable size, situated along an oblique ridge. Outermost denticle sits on the edge of sinulus; columellar tooth deeply situated, elongated lamella-like, sharp. Peristome thickened, expanded, and slightly reflected. Basal swelling situated in some distance from peristome, very slight. Anterior to basal swelling with a point-like depression, which corresponds with the lower palatal denticle. Umbilicus narrow but open, teardrop-shaped due to the swollen inner part of the body whorl, whereby swelling corresponds with deeply situated columellar tooth.
</paragraph>
<paragraph id="8BF8A831C502140FFF52FEFDFDC81F5D" blockId="6.[136,774,317,373]" pageId="6" pageNumber="7">
<emphasis id="B9337423C502140FFF52FEFDFEAC1F7D" bold="true" box="[136,314,317,341]" pageId="6" pageNumber="7">Measurements.</emphasis>
H =
<quantity id="4CBF05D4C502140FFEA1FEFDFDB61F7D" box="[379,544,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="4.84" metricValueMax="5.31" metricValueMin="4.37" pageId="6" pageNumber="7" unit="mm" value="4.84" valueMax="5.31" valueMin="4.37">4.37–5.31 mm</quantity>
, D =
<quantity id="4CBF05D4C502140FFDBCFEFDFC901F7D" box="[614,774,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="2.585" metricValueMax="2.78" metricValueMin="2.39" pageId="6" pageNumber="7" unit="mm" value="2.585" valueMax="2.78" valueMin="2.39">2.39–2.78 mm</quantity>
(smallest and largest specimens measured).
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC502140FFF52FE5DFC9C1C9D" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BF8A831C502140FFF52FE5DFC9C1C9D" blockId="6.[136,778,413,693]" pageId="6" pageNumber="7">
<emphasis id="B9337423C502140FFF52FE5DFE141F9D" bold="true" box="[136,386,413,437]" pageId="6" pageNumber="7">Differential diagnosis.</emphasis>
There are no similar species in
<collectingCountry id="F350E8A1C502140FFD11FE5EFC971F9D" box="[715,769,414,437]" name="Laos" pageId="6" pageNumber="7">Laos</collectingCountry>
,
<collectingCountry id="F350E8A1C502140FFF52FE7DFF711FFD" box="[136,231,445,469]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
, or
<collectingCountry id="F350E8A1C502140FFED4FE7DFEE61FFD" box="[270,368,445,469]" name="Thailand" pageId="6" pageNumber="7">Thailand</collectingCountry>
. The Chinese
<emphasis id="B9337423C502140FFDDDFE7EFC911FFD" box="[519,775,446,469]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C47D3B2C502140FFDDDFE7EFDE01FFD" authorityName="Kobelt" authorityYear="1904" box="[519,630,446,469]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">Sinoennea</taxonomicName>
manyunensis
</emphasis>
B. Fan, M. Tian &amp; Y.-X. Chen, 2014, is similar in its large size, oval shell, and aperture formation. However, it has ribs extending from suture to suture, and has a long palatal ridge running into the aperture from the peristome.
<taxonomicName id="4C47D3B2C502140FFD4FFDFEFF7A1C5D" authority="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi, 2020" authorityName="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi" authorityYear="2020" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="otostoma" status="sp. nov.">
<emphasis id="B9337423C502140FFD4FFDFEFF7A1C5D" italics="true" pageId="6" pageNumber="7">Sinoennea otostoma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C502140FFF27FD9EFE111C5D" box="[253,391,605,629]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, has a similar aperture formation, and therefore, it is probably closely related. However, it is much smaller and glossier, and has a bean-shaped aperture.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC502140FFF52FD1DFEDC1D1D" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BF8A831C502140FFF52FD1DFEDC1D1D" blockId="6.[136,775,733,821]" pageId="6" pageNumber="7">
<emphasis id="B9337423C502140FFF52FD1DFE9C1CDD" bold="true" box="[136,266,733,757]" pageId="6" pageNumber="7">Etymology.</emphasis>
This species is named after its resemblance to the genus
<emphasis id="B9337423C502140FFF23FD3DFEE41D3D" box="[249,370,765,789]" italics="true" pageId="6" pageNumber="7">Ljudmilena</emphasis>
<bibRefCitation id="EFD6D5C0C502140FFEA1FD3DFDBB1D3D" author="Schileyko AA" box="[379,557,765,789]" pageId="6" pageNumber="7" refId="ref8597" refString="Schileyko AA (1984) Terrestrial molluscs of the suborder Pupillina of the USSR fauna (Gastropoda, Pulmonata, Geophila). In: Fauna SSSR. Molljuski. Volume 3. No. 3. Nauka, Leningrad, 399 pp." type="book" year="1984">Schileyko, 1984</bibRefCitation>
, of the
<taxonomicName id="4C47D3B2C502140FFD59FD3DFED11D1D" authority="B. B. Woodward, 1903" authorityName="B. B. Woodward" authorityYear="1903" class="Gastropoda" family="Enidae" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="family">
Enidae B.B.
<bibRefCitation id="EFD6D5C0C502140FFF52FCDDFED11D1D" author="Woodward BB" box="[136,327,797,821]" pageId="6" pageNumber="7" pagination="352 - 367" refId="ref8834" refString="Woodward BB (1903) List of British non-marine Mollusca. Journal of Conchology, 10 (12): 352 - 367." type="journal article" year="1903">Woodward, 1903</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC502140FFF52FC9DFE851DFD" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BF8A831C502140FFF52FC9DFE851DFD" blockId="6.[136,775,861,981]" pageId="6" pageNumber="7">
<emphasis id="B9337423C502140FFF52FC9DFF611D5D" bold="true" box="[136,247,861,885]" pageId="6" pageNumber="7">Remarks.</emphasis>
The
<typeStatus id="54FC1693C502140FFEF1FC9EFECF1D5E" box="[299,345,862,886]" pageId="6" pageNumber="7">type</typeStatus>
locality (Pah Hia) could not be found on maps. In an earlier article, we speculated that its geographic position could be at
<geoCoordinate id="EE73CEF6C502140FFEB5FC5CFE5A1D9D" box="[367,460,924,949]" degrees="18" direction="north" minutes="59" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="18.983334">18°59′N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C502140FFE00FC5CFDA81D9D" box="[474,574,924,949]" degrees="103" direction="east" minutes="16" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="103.26667">103°16′E</geoCoordinate>
(see
<bibRefCitation id="EFD6D5C0C502140FFDA7FC5DFE911DFD" author="Pall-Gergely B &amp; Muratov IV &amp; Asami T" pageId="6" pageNumber="7" pagination="1 - 26" refId="ref8367" refString="Pall-Gergely B, Muratov IV &amp; Asami T (2016) The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species. ZooKeys, 592: 1 - 26." type="journal article" year="2016">Páll-Gergely et al., 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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