treatments-xml/data/FD/C6/B6/FDC6B62E04D64767836C78F21D6A91B0.xml
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<mods:title id="ED8CFC950B9DBDCE9A50AE77027502AD">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<paragraph id="898C32C1DBFCCCAB0D096C7AE9CA8C21" pageId="195" pageNumber="196">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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<subSubSection id="DF6F568056483E5661B726158E441920" pageId="195" pageNumber="196" type="nomenclature">
<paragraph id="45DEE7C2CD2A200FE80F252D79470890" pageId="195" pageNumber="196">
<taxonomicName id="B5E0BD4E0833C726755BE90A84B1C15A" ID-CoL="3RM7L" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="195" pageNumber="196" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus Sharp, 1882</taxonomicName>
Figs 162-163, 344-346, 491, 565
</paragraph>
</subSubSection>
<subSubSection id="89229BAC53D6793B6EAE50A47C0139B1" lastPageId="196" lastPageNumber="197" pageId="195" pageNumber="196" type="reference_group">
<paragraph id="C42349D531DF19CEEBFF79FDFDFA72E8" lastPageId="196" lastPageNumber="197" pageId="195" pageNumber="196">
<taxonomicName id="E5AAFC6BB6138DF525994407B9217E03" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="195" pageNumber="196" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
<bibRefCitation id="C7BF7E5176F6A8DF796064A5786104DB" author="Sharp, D" journalOrPublisher="The Scientific Transactions of the Royal Dublin Society" pageId="263" pageNumber="264" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
: 306, 307 (original description, faunistics, discussion); v. d.
<bibRefCitation id="414BC16F1CC8CDDD6AD07846E6CCD33E" author="Branden, C van den" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="255" pageNumber="256" pagination="1 - 118" title="Catalogue des coleopteres carnassiers aquatiques (Haliplidae, Amphizoidae, Pelobiidae et Dytiscidae)." volume="28" year="1885">Branden 1885</bibRefCitation>
: 21 (catalogue, faunistics);
<bibRefCitation id="3477617E8A5982F7DFFC06246D357A1D" author="Regimbart, M" journalOrPublisher="Memoires de la Societe entomologique de Belgique" pageId="262" pageNumber="263" pagination="1 - 244" title="Revision des Dytiscidae et Gyrinidae d'Afrique, Madagascar et iles voisines. En contribution a la faune entomologique du Congo." volume="4" year="1895">
<normalizedToken id="68FD9F5F77F4B0416103C88B80FDE43A" originalValue="Régimbart">Regimbart</normalizedToken>
1895
</bibRefCitation>
: 141 (description, faunistics);
<bibRefCitation id="528F5CB2520067D3C81B68E1E21DBFB3" author="Regimbart, M" journalOrPublisher="Bullettino della Societa Entomologica Italiana" pageId="262" pageNumber="263" pagination="201 - 226" title="Materiali per lo studio della fauna Eritrea raccolti nel 1901 - 03 dal Dr. A, Andreini tenente medico." volume="36" year="1905">
<normalizedToken id="CCA565D25304D7A6B59896111F847C7C" originalValue="Régimbart">Regimbart</normalizedToken>
1905
</bibRefCitation>
: 208 (faunistics);
<bibRefCitation id="F984E0133BF9ABD05B1CDB2604ADC9B8" author="Regimbart, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="262" pageNumber="263" pagination="235 - 278" title="Dytiscidae, Gyrinidae, Hydrophilidae. Voyage de M. Ch. Alluaud dans l'Afrique orientale." volume="75" year="1906">
<normalizedToken id="02BA27226BF81AF6B8A773D09E6B11B5" originalValue="Régimbart">Regimbart</normalizedToken>
1906
</bibRefCitation>
: 249 (faunistics);
<bibRefCitation id="C5D02DB2BCFB812BF2B3320F43112E63" author="Regimbart, M" journalOrPublisher="Almquist &amp; Wiksells Boktryckeri-A. B., Uppsala" pageId="262" pageNumber="263" title="Coleoptera. 1. Dytiscidae, Gyrinidae, Hydrophilidae. In Sjoestedt, Y Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und der umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906" year="1908">
<normalizedToken id="CFEB9064BED8432D7D8592171BAB375B" originalValue="Régimbart">Regimbart</normalizedToken>
1908
</bibRefCitation>
a: 5 (faunistics);
<bibRefCitation id="ADF8B88F7BA1ED208B834FD5162E8CE2" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
: 19 (catalogue, faunistics);
<bibRefCitation id="34974B3A41A719F61B53C39DEFA77245" author="Regimbart, M" journalOrPublisher="Voyage de M. Le Baron Maurice de Rothschild en Ethiopie et Afrique orientale Anglaise (1904 - 1905)" pageId="262" pageNumber="263" pagination="528 - 536" title="Dytiscides, gyrinides et hydrophilides." volume="2" year="1922">
<normalizedToken id="E7B43CED310F2BDEC64E8ACECBC8A7E8" originalValue="Régimbart">Regimbart</normalizedToken>
1922
</bibRefCitation>
: 532 (faunistics);
<bibRefCitation id="7E014E1CF5F9900B29624B85F7368FE0" pageId="195" pageNumber="196">Omer-Cooper 1931</bibRefCitation>
: 758 (description, discussion, biology, faunistics);
<bibRefCitation id="568182847C99FAC755C32DE31461881A" author="Gschwendtner, L" journalOrPublisher="Exploration du Parc National Albert. Mission G. F. de Witte (1933 - 1935)" pageId="256" pageNumber="257" pagination="3 - 22" title="Haliplidae und Dytiscidae." volume="13" year="1938 a">Gschwendtner 1938a</bibRefCitation>
: (faunistics, discussion);
<bibRefCitation id="761FC758995B5E8EBCD59EB70175D6B5" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
: 263, 264, 265, 279, 312 (description, faunistics, discussion);
<bibRefCitation id="8F00BC87326AE49F63D0A65064E3998E" author="Legros, C" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="259" pageNumber="260" pagination="265 - 280" title="XXI. Coleopteres Hydrocanthares (Mission M. Lamotte, 1942)." volume="40" year="1954">Legros 1954</bibRefCitation>
: 268 (discussion);
<bibRefCitation id="F35E5A238FB1A1E91B5C3E282DE83EC7" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 544, 548, 549, 550, 552 (description, faunistics, discus
<pageBreakToken id="E00CD03C740BFDD6854A98F0B18C5D6D" pageId="196" pageNumber="197" start="start">sion</pageBreakToken>
);
<bibRefCitation id="B89444642D4D9E082A55CAD744F43D8D" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
: 286, 287 (description, discussion);
<bibRefCitation id="C1E441EDF825FC07CCFC46FA520CD4C2" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
: 79, 94 (faunistics, discussion, biology);
<bibRefCitation id="2896F1D854C509ED6673E713237E3C35" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 244 (catalogue, faunistics);
<bibRefCitation id="5FC46F7A296F8AF97AA47255E504E50B" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 212 (catalogue, faunistics). [Comments: information on
<taxonomicName id="2575CED6CFCA051DB040D88195071051" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
outside Ethiopia and Eritrea is considered uncertain.]
</paragraph>
</subSubSection>
<subSubSection id="2ADFBA630BE1ED8A26762C60901D8484" pageId="196" pageNumber="197" type="type locality">
<paragraph id="889677A732EA5D6B8FF4698137BE4C1F" pageId="196" pageNumber="197">Type locality.</paragraph>
<paragraph id="D61558167DF0DBCAF4B18039AA8ED64D" pageId="196" pageNumber="197">Ethiopia: Abyssinia.</paragraph>
</subSubSection>
<subSubSection id="0E2CFE5AB7A27ACED04CAD7079DD50DE" pageId="196" pageNumber="197" type="type material">
<paragraph id="75BB05E52EB408DF9FF68DCC8E4C858D" pageId="196" pageNumber="197">Type material studied</paragraph>
<paragraph id="A0BE8AEA2DD3FDD8EF296E6138152184" pageId="196" pageNumber="197">
(3 exs.). Lectotype (by present desgination): male: &quot;Type 569 / Type / Abyssinia / Sharp Coll. 1905-313 /
<taxonomicName id="11ECE103A3774E213D736E9715A22D5C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
sp. n. Abyssinia&quot; (BMNH). - Paralecotytpes: &quot;569 / Co-type / Abyssinia/Sharp Coll. 1905-313 /
<taxonomicName id="36EB990507EF27D7973817ABD93EB9C4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
Shp Co-type&quot;(1 female ex. BMNH); &quot;Co-type / Abyss. Raffray / Abyssinia A. Raffray /
<taxonomicName id="A3656FCEDB637E9CF5110F6AE7E34750" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
Shp Co-type&quot;(1 male ex. BMNH).
</paragraph>
</subSubSection>
<subSubSection id="C8A48994D95101824A1FE98EC13493AD" pageId="196" pageNumber="197" type="materials_examined">
<paragraph id="8BE86134D644CED8E917CE825A1BEF63" pageId="196" pageNumber="197">Additional material studied</paragraph>
<paragraph id="7FD7349F2C64A28C54CCC120973D12DC" pageId="196" pageNumber="197">
(16 exs.). Ethiopia: &quot;Stream W of Zaguala 6000 ft. 27.10. 1926 JOC.&quot; (5 exs. AMGS); &quot;Abyss. Raffr. /
<taxonomicName id="9615B9E87A66DBE51A9E5FE8C3952C49" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
Shp det. M. Brancucci&quot; (3 exs. NHMB); &quot;Tigray Province ca. 20 km E Axum 2000 m 17.4. 2006 leg. Wewalka&quot; (4 exs. CGW; habitus in Fig. 491). - Eritrea: &quot;Adi-Ugri 5. 1901 Andreini /
<taxonomicName id="48DB0BE25C44D22168525D9CE0A970FA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
Sharp det. Rocchi 1995&quot; (2 exs. CSR); &quot;Asmara-Decamere 25.5. 1963 Linnavuori&quot; (1 ex. MZH); &quot;Ghinda Levander&quot; (6 exs. MZH).
</paragraph>
</subSubSection>
<subSubSection id="A6C2E4D6D7F38E1D84E1E36423056CF4" pageId="196" pageNumber="197" type="diagnosis">
<paragraph id="AAEF44098CE7C381FF3079A1CA9416FC" pageId="196" pageNumber="197">Diagnosis.</paragraph>
<paragraph id="7C337345667DE24114151082C721C41D" pageId="196" pageNumber="197">
<taxonomicName id="B3BAE8ACC60EB2ABB924CBF6E417FD1A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus grammicus" order="Coleoptera" pageId="196" pageNumber="197" phylum="Arthropoda" rank="species" species="grammicus">Laccophilus grammicus</taxonomicName>
resembles a number of species in the same species group and can therefore be difficult to identify solely using external features as appearance of body colour pattern. The species is, however, easily separated by examination of male genitalia; penis in apical half tapers gradually towards apex; in lateral aspect, penis very strongly curved.
</paragraph>
</subSubSection>
<subSubSection id="30BB06BCAF9D8CBAF06BB771D144FA9C" lastPageId="197" lastPageNumber="198" pageId="196" pageNumber="197" type="description">
<paragraph id="97FE7A5D745697B553D43A9A2E5893A4" pageId="196" pageNumber="197">Description.</paragraph>
<paragraph id="CB788E0628AEABBCF3EC8132E6FFA621" pageId="196" pageNumber="197">Body length 4.1-4.3 mm, width 2.2-2.4 mm. Dorsal, colour pattern of body distinct and quite uniform (Fig. 491).</paragraph>
<paragraph id="731D564CEAAB795ED6B7A4E21F3460ED" pageId="196" pageNumber="197">Head: Pale ferrugineous. Slightly mat, finely microsculptured. Reticulation double: number of fine meshes in one large mesh varies from 2-5. In part, double reticulation weakly developed and rather indistinct: fine and large meshes difficult to distinguish. At eyes and medially from eyes towards middle of head finely and sparsely punctured. Head extensively impunctate.</paragraph>
<paragraph id="2DCE777BDCBA02554A19DB5016D7D1C3" pageId="196" pageNumber="197">Pronotum: Pale ferrugineous; mediobasally with narrow blackish ferrugineous marking. Submat, finely microsculptured; reticulation double: large meshes contain 2-6 fine meshes. At margins with irregular, sparse row of fine punctures. At base, row of punctures in part absent.</paragraph>
<paragraph id="0ABDD5B1FF82185097DED9E9FEC9595D" pageId="196" pageNumber="197">Elytra: Pale ferrugineous, with distinct blackish to dark ferrugineous, longitudinal markings, which are in part reduced basally, slightly posterior to middle and apically (Fig. 491). Submat, finely microsculptured. Reticulation predominantly simple; large meshes discernible but vague and weakly developed. Three rather indistinct rows of sparse punctures are discernible.</paragraph>
<paragraph id="636AA998618557A918E400E28A59A60C" pageId="196" pageNumber="197">Ventral aspect: Pale ferrugineous to dark ferrugineous. Almost impunctate. Very finely microsculptured; in part fine reticulation indistinct or absent. Metacoxal plates in anterior half with shallow, transversely located, in part rather indistinct furrows. Abdomen with fine, curved striae. Apical ventrite with sharp knob on one side (Fig. 162). Prosternal process slender, pointed.</paragraph>
<paragraph id="63278EE7C19D8312944A8267FA09B9BB" pageId="197" pageNumber="198">
<pageBreakToken id="C93C61860144653817BAFB059CA2C06A" pageId="197" pageNumber="198" start="start">Legs</pageBreakToken>
: Pro- and mesotarsus quite long, slightly enlarged, provided with suckers.
</paragraph>
<paragraph id="B29EFE7703BB9D5BB5CD304003A3BC21" pageId="197" pageNumber="198">Male genitalia: Penis in apical half tapers rather evenly and gradually towards apex; in lateral aspect, penis very strongly curved (Figs 344-346).</paragraph>
<paragraph id="782F3D62D6F8FBCED775EA6567FB4B51" pageId="197" pageNumber="198">Female: Apical ventrite lacks sharp knob (Fig. 163). Pro- and mesotarsus slender.</paragraph>
</subSubSection>
<subSubSection id="EEA0B7E4437453800D881E8AFC1E0EAB" pageId="197" pageNumber="198" type="distribution">
<paragraph id="94DE33871FC6E5658F5ADDC6286E736D" pageId="197" pageNumber="198">Distribution.</paragraph>
<paragraph id="7E1AC3750514CADFD366BE9BEEF2E71D" pageId="197" pageNumber="198">Ethiopia, Eritrea (Fig. 565). Records outside Ethiopia and Eritrea are considered doubtful.</paragraph>
</subSubSection>
<subSubSection id="0458F6D941560414EF15B591F366B9C4" pageId="197" pageNumber="198" type="collecting circumstances">
<paragraph id="90A5A8FD24710F42AA91920EE4A0338E" pageId="197" pageNumber="198">Collecting circumstances.</paragraph>
<paragraph id="05D0480F05252DF875B094594444A2E9" pageId="197" pageNumber="198">
Collected at high altitudes (
<bibRefCitation id="9AC0A7A89ACABD52AAF0D1D4C9D16813" pageId="197" pageNumber="198">Omer-Cooper 1931</bibRefCitation>
,
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).
</paragraph>
</subSubSection>
</treatment>
</document>