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<document id="300F1FD8941C32B431DED89DF3851144" ID-CLB-Dataset="63555" ID-DOI="10.11646/zootaxa.5150.1.5" ID-GBIF-Dataset="0e9631c5-3211-4f61-9549-db14a51dbc30" ID-ISSN="1175-5326" ID-Zenodo-Dep="6610039" ID-ZooBank="2F6FD286-5E1E-40F7-B658-490EFD453BDF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654245021749" checkinUser="plazi" docAuthor="Golovatch, Sergei &amp; Vandenspiegel, Didier" docDate="2022" docId="F33487D0FFDDFFFCFF78FF56FE0518AE" docLanguage="en" docName="zootaxa.5150.1.5.pdf" docOrigin="Zootaxa 5150 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Costabolus baculus Golovatch &amp; Vandenspiegel 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="107" masterDocId="0F0DFFA8FFDBFFF6FFEFFFC1FFC61903" masterDocTitle="Carinobolus gen. nov. and Costabolus gen. nov., two new, remarkably crested monospecific genera of spirobolidan millipedes from WestAfrica, with the proposal of a new tribe, Amblybolini tribus nova (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="110" masterPageNumber="97" pageNumber="103" updateTime="1699338865200" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="76CCC28EC1EAE64EC69B92953BC7D188" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C2F90774D45DEB7D4A3D360C09C59C9">
<mods:title id="6FC2216DC3692B2FF72EAE60658573F8">Carinobolus gen. nov. and Costabolus gen. nov., two new, remarkably crested monospecific genera of spirobolidan millipedes from WestAfrica, with the proposal of a new tribe, Amblybolini tribus nova (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
</mods:titleInfo>
<mods:name id="331164997CA33B65534B4FBCF5B74950" type="personal">
<mods:role id="A03D2B2117A44501015DD4E4DE8FD66C">
<mods:roleTerm id="D77C622334A6033FF7B206FBF82CF117">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B048C67F0FF9EDD59CE6915E22C565B5">Golovatch, Sergei</mods:namePart>
<mods:affiliation id="813AD65B3BC9BF7763669BC7E3C1E140">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow, 119071 Russia. Biological Collection and Data Management Unit, Royal Museum for Central Africa, Tervuren, Belgium. didier. van. den. spiegel @ africamuseum. be; https: // orcid. org / 0000 - 0002 - 8696 - 9810</mods:affiliation>
</mods:name>
<mods:name id="0FD4A56B212F07C3114564BBD9B6342B" type="personal">
<mods:role id="DFBF026E20DB8F5BA235964FD2C9F9A9">
<mods:roleTerm id="C7410D6023F78874035694688D1FD63A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4E5F74E2E01E1354326F102AD4B4F68">Vandenspiegel, Didier</mods:namePart>
<mods:nameIdentifier id="D09D25DD08BF909531DA883242F6B61F" type="ORCID">0000-0002-8696-9810</mods:nameIdentifier>
<mods:nameIdentifier id="28DC5DDF7390619ED8CE82D54828BC09" type="email">didier.van.den.spiegel@africamuseum.be</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="6A76E99657BE9448A1E7A43AE3EE13DA">text</mods:typeOfResource>
<mods:relatedItem id="5830E8776393A6468210D426DD47EB77" type="host">
<mods:titleInfo id="155E1788164BB984FC31CC05978B2657">
<mods:title id="7CFB6ABCB72F592D5C3C2B3125C5FD9F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2D0BC0D6EA76D16D1C41FB937EAB12C4">
<mods:date id="343CD4C696B2E956FF97DD139F2610C1">2022</mods:date>
<mods:detail id="4E1B507D95E19197128DA39475A3AF22" type="pubDate">
<mods:number id="204A8EA63D7705E6673F306DC491497B">2022-06-03</mods:number>
</mods:detail>
<mods:detail id="F98429DE20E6F24C1A6EB5AEC6D54507" type="volume">
<mods:number id="2973F133928A68C0E57F5E8E45082D86">5150</mods:number>
</mods:detail>
<mods:detail id="26983B1C4EF7D6B545068DD7F6CFC996" type="issue">
<mods:number id="5DBDD9336F0AB58C057A80387FA86623">1</mods:number>
</mods:detail>
<mods:extent id="B4C4204D35476406D9FA25C112EC3FA5" unit="page">
<mods:start id="80C891D68B5772397B4172FBCEEF3F42">97</mods:start>
<mods:end id="EDDAB47B0CBAAC92AF243F5D2B78391E">110</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9CD593854C005DEF8351041A6BFF7859">journal article</mods:classification>
<mods:identifier id="8A122D81981393DCF8157372E25EAC92" type="CLB-Dataset">63555</mods:identifier>
<mods:identifier id="25F7D404E774AB785269EE15F2F03BF1" type="DOI">10.11646/zootaxa.5150.1.5</mods:identifier>
<mods:identifier id="E6A81FB6B76810964E885144374F9400" type="GBIF-Dataset">0e9631c5-3211-4f61-9549-db14a51dbc30</mods:identifier>
<mods:identifier id="805DA4E99D7BE41976D175B1D11952E6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6019066D01DC3199F4B6FC54FA7A09CA" type="Zenodo-Dep">6610039</mods:identifier>
<mods:identifier id="FACA5D66F96D77E24BFBB3AB2667E85C" type="ZooBank">2F6FD286-5E1E-40F7-B658-490EFD453BDF</mods:identifier>
</mods:mods>
<treatment id="F33487D0FFDDFFFCFF78FF56FE0518AE" ID-DOI="http://doi.org/10.5281/zenodo.6610031" ID-GBIF-Taxon="195730569" ID-Zenodo-Dep="6610031" LSID="urn:lsid:plazi:treatment:F33487D0FFDDFFFCFF78FF56FE0518AE" httpUri="http://treatment.plazi.org/id/F33487D0FFDDFFFCFF78FF56FE0518AE" lastPageId="10" lastPageNumber="107" pageId="6" pageNumber="103">
<subSubSection id="3387654DFFDDFFF0FF78FF56FE2E19B2" box="[151,488,151,178]" pageId="6" pageNumber="103" type="nomenclature">
<paragraph id="7B2236C6FFDDFFF0FF78FF56FE2E19B2" blockId="6.[151,488,151,213]" box="[151,488,151,178]" pageId="6" pageNumber="103">
<heading id="206A81AAFFDDFFF0FF78FF56FE2E19B2" bold="true" box="[151,488,151,178]" fontSize="11" level="1" pageId="6" pageNumber="103" reason="1">
<emphasis id="49E9EAD4FFDDFFF0FF78FF56FE2E19B2" bold="true" box="[151,488,151,178]" pageId="6" pageNumber="103">
<taxonomicName id="BC9D4D45FFDDFFF0FF78FF56FE4519B1" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[151,387,151,178]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFDDFFF0FF78FF56FE4519B1" bold="true" box="[151,387,151,178]" italics="true" pageId="6" pageNumber="103">Costabolus baculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52DA57AFFFDDFFF0FE65FF56FE2E19B2" box="[394,488,151,177]" pageId="6" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3387654DFFDDFFF0FF78FF7AFF3E19D6" box="[151,248,187,213]" pageId="6" pageNumber="103" type="description">
<paragraph id="7B2236C6FFDDFFF0FF78FF7AFF3E19D6" blockId="6.[151,488,151,213]" box="[151,248,187,213]" pageId="6" pageNumber="103">
<figureCitation id="E3A62A43FFDDFFF0FF78FF7AFF3E19D6" box="[151,248,187,213]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="6.[151,250,1942,1966]" captionStartId-1="7.[151,250,1776,1800]" captionStartId-2="8.[151,250,1776,1800]" captionStartId-3="9.[151,250,605,629]" captionTargetBox-0="[151,1436,669,1918]" captionTargetBox-1="[189,1398,181,1753]" captionTargetBox-2="[238,1349,181,1753]" captionTargetBox-3="[151,1436,181,582]" captionTargetId-0="figure-185@6.[151,1436,669,1918]" captionTargetId-1="figure-21@7.[189,1398,181,1753]" captionTargetId-2="figure-23@8.[238,1349,181,1753]" captionTargetId-3="figure-21@9.[151,1436,181,582]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 4. Costabolus baculus sp. nov., male holotype (VNHM 198). A &amp; B. Habitus, lateral and ventral views, respectively. Scale bars: 5 mm." captionText-1="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." captionText-2="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." captionText-3="FIGURE 7. Costabolus baculus sp. nov., female paratype (VMNH 154). A. Ventral view of both vulvae, B. Left vulva, lateral view. Abbreviations: av, anterior valve; cx, coxite of leg 1; cxv, coxite of vulva; o, operculum; pv, posterior valve. Scale bars: 500 µm (A), 200 µm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.6610051" figureDoi-1="http://doi.org/10.5281/zenodo.6610053" figureDoi-2="http://doi.org/10.5281/zenodo.6610060" figureDoi-3="http://doi.org/10.5281/zenodo.6610064" httpUri-0="https://zenodo.org/record/6610051/files/figure.png" httpUri-1="https://zenodo.org/record/6610053/files/figure.png" httpUri-2="https://zenodo.org/record/6610060/files/figure.png" httpUri-3="https://zenodo.org/record/6610064/files/figure.png" pageId="6" pageNumber="103">Figs 4–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3387654DFFDDFFF0FF78FEC2FED21B1A" pageId="6" pageNumber="103" type="materials_examined">
<paragraph id="7B2236C6FFDDFFF0FF78FEC2FBE91842" blockId="6.[151,1437,259,646]" pageId="6" pageNumber="103">
<emphasis id="49E9EAD4FFDDFFF0FF78FEC2FE46181E" bold="true" box="[151,384,259,285]" pageId="6" pageNumber="103">Material examined.</emphasis>
<materialsCitation id="CBF53C9BFFDDFFF0FE69FEC2FBE91842" ID-GBIF-Occurrence="3802884304" collectingDate="1984-08-12" collectionCode="VNHM" country="Nigeria" elevation="20" location="Calabar" pageId="6" pageNumber="103" specimenCode="VNHM 198" specimenCount="1" typeStatus="holotype">
<typeStatus id="A4268864FFDDFFF0FE69FEC2FE28181E" box="[390,494,259,285]" pageId="6" pageNumber="103" type="holotype">Holotype</typeStatus>
male (with exposed gonopods, body broken into two pieces) (
<specimenCode id="2B3B9EBDFFDDFFF0FB4CFEC2FAEB181E" box="[1187,1325,259,285]" collectionCode="VNHM" pageId="6" pageNumber="103">VNHM 198</specimenCode>
),
<collectingCountry id="038A7656FFDDFFF0FAAEFEC2FA5E181E" box="[1345,1432,259,285]" name="Nigeria" pageId="6" pageNumber="103">Nigeria</collectingCountry>
,
<quantity id="BC659B23FFDDFFF0FF78FEE6FF181842" box="[151,222,295,321]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="103" unit="km" value="27.0">27 km</quantity>
NE of
<location id="7E42601DFFDDFFF0FEDFFEE6FE4F1842" LSID="urn:lsid:plazi:treatment:F33487D0FFDDFFFCFF78FF56FE0518AE:7E42601DFFDDFFF0FEDFFEE6FE4F1842" box="[304,393,295,321]" country="Nigeria" name="Calabar" pageId="6" pageNumber="103">Calabar</location>
,
<quantity id="BC659B23FFDDFFF0FE7BFEE6FE0B1841" box="[404,461,295,322]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="103" unit="m" value="20.0">
<elevation id="F0B0D1F5FFDDFFF0FE7BFEE6FE0B1841" box="[404,461,295,322]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="103" unit="m" value="20.0">20 m</elevation>
</quantity>
inside native forest,
<date id="0F231006FFDDFFF0FD59FEE6FC8D1842" box="[694,843,295,321]" pageId="6" pageNumber="103" value="1984-08-12">
<collectingDate id="1F67E9EEFFDDFFF0FD59FEE6FC8D1842" box="[694,843,295,321]" pageId="6" pageNumber="103" value="1984-08-12">12.VIII.1984</collectingDate>
</date>
, collector unknown.
</materialsCitation>
</paragraph>
<paragraph id="7B2236C6FFDDFFF0FF28FE8DFD9618D2" blockId="6.[151,1437,259,646]" pageId="6" pageNumber="103">
<materialsCitation id="CBF53C9BFFDDFFF0FF28FE8DFC8E1866" ID-GBIF-Occurrence="3802884309" box="[199,840,331,358]" collectingDate="1984-08-12" collectionCode="VNHM" country="Nigeria" elevation="20" location="Calabar" pageId="6" pageNumber="103" specimenCode="VNHM 198" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="A4268864FFDDFFF0FF28FE8DFEFE1865" box="[199,312,332,358]" pageId="6" pageNumber="103" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="6D9BFD4FFFDDFFF0FEABFE8AFE5C1866" box="[324,410,331,357]" pageId="6" pageNumber="103" type="male">2 males</specimenCount>
(
<specimenCode id="2B3B9EBDFFDDFFF0FE45FE8AFDF01866" box="[426,566,331,357]" collectionCode="VNHM" pageId="6" pageNumber="103">VNHM 198</specimenCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="CBF53C9BFFDDFFF0FCBBFE8AFF2118AE" ID-GBIF-Occurrence="3802884302" collectingDate="1984-04-28" collectionCode="VMNH" country="Nigeria" location="Calabar" pageId="6" pageNumber="103" specimenCode="VMNH 154" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="6D9BFD4FFFDDFFF0FCBBFE8AFC731866" box="[852,949,331,357]" pageId="6" pageNumber="103" type="female">1 female</specimenCount>
(
<specimenCode id="2B3B9EBDFFDDFFF0FC2AFE8AFB971865" box="[965,1105,331,358]" collectionCode="VMNH" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" pageId="6" pageNumber="103">VMNH 154</specimenCode>
),
<collectingCountry id="038A7656FFDDFFF0FB89FE8AFB7B1866" box="[1126,1213,331,357]" name="Nigeria" pageId="6" pageNumber="103">Nigeria</collectingCountry>
,
<quantity id="BC659B23FFDDFFF0FB26FE8AFAD71866" box="[1225,1297,331,357]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="103" unit="km" value="27.0">27 km</quantity>
NE
<location id="7E42601DFFDDFFF0FAAAFE8AFA5A1866" LSID="urn:lsid:plazi:treatment:F33487D0FFDDFFFCFF78FF56FE0518AE:7E42601DFFDDFFF0FAAAFE8AFA5A1866" box="[1349,1436,331,357]" country="Nigeria" name="Calabar" pageId="6" pageNumber="103">Calabar</location>
(Ekang road), under logs on fairly dry soil in new canopy forest farmed to young maize,
<date id="0F231006FFDDFFF0FB6CFEAEFAC7188A" box="[1155,1281,367,393]" pageId="6" pageNumber="103" value="1984-04-28">
<collectingDate id="1F67E9EEFFDDFFF0FB6CFEAEFAC7188A" box="[1155,1281,367,393]" pageId="6" pageNumber="103" value="1984-04-28">28.IV.1984</collectingDate>
</date>
, collector unknown
</materialsCitation>
;
<materialsCitation id="CBF53C9BFFDDFFF0FF1DFE52FD8A18D2" ID-GBIF-Occurrence="3802884307" collectingDate="1984-08-12" collectionCode="VMHN" country="Nigeria" elevation="20" location="Calabar" pageId="6" pageNumber="103" specimenCode="VMHN 199" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="6D9BFD4FFFDDFFF0FF1DFE52FEF918AE" box="[242,319,403,429]" pageId="6" pageNumber="103" type="male">1 male</specimenCount>
,
<specimenCount id="6D9BFD4FFFDDFFF0FEA5FE52FE6D18AE" box="[330,427,403,429]" pageId="6" pageNumber="103" type="female">1 female</specimenCount>
(
<specimenCode id="2B3B9EBDFFDDFFF0FE56FE52FD8218AE" box="[441,580,403,429]" collectionCode="VMHN" pageId="6" pageNumber="103">VMHN 199</specimenCode>
),
<collectingCountry id="038A7656FFDDFFF0FDB7FE52FD6918AE" box="[600,687,403,429]" name="Nigeria" pageId="6" pageNumber="103">Nigeria</collectingCountry>
,
<quantity id="BC659B23FFDDFFF0FD56FE52FCC618AE" box="[697,768,403,429]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="103" unit="km" value="27.0">27 km</quantity>
N of
<location id="7E42601DFFDDFFF0FCD0FE52FC5E18AE" LSID="urn:lsid:plazi:treatment:F33487D0FFDDFFFCFF78FF56FE0518AE:7E42601DFFDDFFF0FCD0FE52FC5E18AE" box="[831,920,403,429]" country="Nigeria" name="Calabar" pageId="6" pageNumber="103">Calabar</location>
, between trees, native forest,
<quantity id="BC659B23FFDDFFF0FB0EFE52FADC18AD" box="[1249,1306,403,430]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="103" unit="m" value="20.0">
<elevation id="F0B0D1F5FFDDFFF0FB0EFE52FADC18AD" box="[1249,1306,403,430]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="103" unit="m" value="20.0">20 m</elevation>
</quantity>
off logging road,
<date id="0F231006FFDDFFF0FF38FE76FEAA18D2" box="[215,364,439,465]" pageId="6" pageNumber="103" value="1984-08-12">
<collectingDate id="1F67E9EEFFDDFFF0FF38FE76FEAA18D2" box="[215,364,439,465]" pageId="6" pageNumber="103" value="1984-08-12">12.VIII.1984</collectingDate>
</date>
, collector unknown
</materialsCitation>
.
</paragraph>
<paragraph id="7B2236C6FFDDFFF0FF28FE1AFED21B1A" blockId="6.[151,1437,259,646]" pageId="6" pageNumber="103">
<emphasis id="49E9EAD4FFDDFFF0FF28FE1AFECB18F6" bold="true" box="[199,269,475,501]" pageId="6" pageNumber="103">Name</emphasis>
.
<emphasis id="49E9EAD4FFDDFFF0FEF3FE1AFEB018F6" box="[284,374,475,501]" italics="true" pageId="6" pageNumber="103">Baculus</emphasis>
in Latin means “crutch”, referring to the long coxal processes on male legs 2 and 3; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="3387654DFFDDFFF0FF28FDE2FDC61B86" pageId="6" pageNumber="103" type="diagnosis">
<paragraph id="7B2236C6FFDDFFF0FF28FDE2FDC61B86" blockId="6.[151,1437,259,646]" pageId="6" pageNumber="103">
<emphasis id="49E9EAD4FFDDFFF0FF28FDE2FEFC1B3E" bold="true" box="[199,314,547,573]" pageId="6" pageNumber="103">Diagnosis</emphasis>
. Basically as in the genus. Body short, up to
<quantity id="BC659B23FFDDFFF0FCDDFDE2FC461B3D" box="[818,896,547,574]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="103" unit="mm" value="27.0">27 mm</quantity>
long, with 38 body rings plus telson, metazonae with simple longitudinal crests/ribs, epiproct slightly projecting caudally. Male with tarsal pads and prominent coxal projections on leg-pairs 2 and 3.
</paragraph>
</subSubSection>
<caption id="2FE2664EFFDDFFF0FF78F857FE901ED1" ID-DOI="http://doi.org/10.5281/zenodo.6610051" ID-Zenodo-Dep="6610051" httpUri="https://zenodo.org/record/6610051/files/figure.png" pageId="6" pageNumber="103" startId="6.[151,250,1942,1966]" targetBox="[151,1436,669,1918]" targetPageId="6">
<paragraph id="7B2236C6FFDDFFF0FF78F857FE901ED1" blockId="6.[151,1437,1942,2002]" pageId="6" pageNumber="103">
<emphasis id="49E9EAD4FFDDFFF0FF78F857FECB1EAD" bold="true" box="[151,269,1942,1966]" pageId="6" pageNumber="103">FIGURE 4</emphasis>
.
<taxonomicName id="BC9D4D45FFDDFFF0FEF6F857FE261EAD" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[281,480,1942,1966]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="103" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFDDFFF0FEF6F857FE261EAD" box="[281,480,1942,1966]" italics="true" pageId="6" pageNumber="103">Costabolus baculus</emphasis>
</taxonomicName>
<emphasis id="49E9EAD4FFDDFFF0FE09F857FDF01EAD" bold="true" box="[486,566,1942,1966]" pageId="6" pageNumber="103">
<taxonomicNameLabel id="52DA57AFFFDDFFF0FE09F857FDF01EAD" box="[486,566,1942,1966]" pageId="6" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (VNHM 198).
<emphasis id="49E9EAD4FFDDFFF0FC80F857FC741EAD" bold="true" box="[879,946,1942,1966]" pageId="6" pageNumber="103">A &amp; B</emphasis>
. Habitus, lateral and ventral views, respectively.
<emphasis id="49E9EAD4FFDDFFF0FF78F87BFEC81ED1" bold="true" box="[151,270,1978,2002]" pageId="6" pageNumber="103">Scale bars:</emphasis>
5 mm.
</paragraph>
</caption>
<caption id="2FE2664EFFDCFFF1FF78F931FDE21EE2" ID-DOI="http://doi.org/10.5281/zenodo.6610053" ID-Zenodo-Dep="6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="7" pageNumber="104" startId="7.[151,250,1776,1800]" targetBox="[189,1398,181,1753]" targetPageId="7">
<paragraph id="7B2236C6FFDCFFF1FF78F931FDE21EE2" blockId="7.[151,1437,1776,2017]" pageId="7" pageNumber="104">
<emphasis id="49E9EAD4FFDCFFF1FF78F931FEC81E0A" bold="true" box="[151,270,1776,1801]" pageId="7" pageNumber="104">FIGURE 5</emphasis>
.
<taxonomicName id="BC9D4D45FFDCFFF1FEF3F930FE231E0A" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[284,485,1777,1801]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="7" pageNumber="104" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFDCFFF1FEF3F930FE231E0A" box="[284,485,1777,1801]" italics="true" pageId="7" pageNumber="104">Costabolus baculus</emphasis>
</taxonomicName>
<emphasis id="49E9EAD4FFDCFFF1FE03F930FDF81E0A" bold="true" box="[492,574,1777,1801]" pageId="7" pageNumber="104">
<taxonomicNameLabel id="52DA57AFFFDCFFF1FE03F930FDF81E0A" box="[492,574,1777,1801]" pageId="7" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (VNHM 198).
<emphasis id="49E9EAD4FFDCFFF1FC91F931FC021E0A" bold="true" box="[894,964,1776,1801]" pageId="7" pageNumber="104">A &amp; B</emphasis>
. Head, lateral and frontal views, respectively.
<emphasis id="49E9EAD4FFDCFFF1FF78F8D5FF6F1E2F" bold="true" box="[151,169,1812,1836]" pageId="7" pageNumber="104">C</emphasis>
. Tip of antenna.
<emphasis id="49E9EAD4FFDCFFF1FEB3F8D4FEA81E2E" bold="true" box="[348,366,1813,1837]" pageId="7" pageNumber="104">D</emphasis>
. Left antenna, lateral view.
<emphasis id="49E9EAD4FFDCFFF1FD7DF8D4FD651E2E" bold="true" box="[658,675,1813,1837]" pageId="7" pageNumber="104">E</emphasis>
. Clypeus, frontolateral view.
<emphasis id="49E9EAD4FFDCFFF1FC38F8D4FBD91E2F" bold="true" box="[983,1055,1812,1837]" pageId="7" pageNumber="104">F &amp; G</emphasis>
. Mandible, ventral and mesal views, respectively.
<emphasis id="49E9EAD4FFDCFFF1FEF3F8F8FEF61E52" bold="true" box="[284,304,1849,1873]" pageId="7" pageNumber="104">H</emphasis>
. Gnathochilarium, ventral view.
<emphasis id="49E9EAD4FFDCFFF1FD94F8F8FD431E52" bold="true" box="[635,645,1849,1873]" pageId="7" pageNumber="104">I</emphasis>
. Right lateral margin of collum, lateral view.
<emphasis id="49E9EAD4FFDCFFF1FBA0F8F8FB9A1E52" bold="true" box="[1103,1116,1849,1873]" pageId="7" pageNumber="104">J</emphasis>
. A midbody metazona.
<emphasis id="49E9EAD4FFDCFFF1FAA5F8F8FA981E52" bold="true" box="[1354,1374,1849,1873]" pageId="7" pageNumber="104">K</emphasis>
. Ozopore.
<emphasis id="49E9EAD4FFDCFFF1FF3DF89CFF251E76" bold="true" box="[210,227,1885,1909]" pageId="7" pageNumber="104">L</emphasis>
. Limbus of a midbody ring.
<emphasis id="49E9EAD4FFDCFFF1FDE3F89CFDE21E76" bold="true" box="[524,548,1885,1909]" pageId="7" pageNumber="104">M</emphasis>
. Posterior part of body, ventral view.
<emphasis id="49E9EAD4FFDCFFF1FC46F89CFC351E77" bold="true" box="[937,1011,1884,1909]" pageId="7" pageNumber="104">N &amp; O</emphasis>
. Midbody legs, ventral and lateral views, respectively.
<emphasis id="49E9EAD4FFDCFFF1FEF2F840FEEA1E9A" bold="true" box="[285,300,1921,1945]" pageId="7" pageNumber="104">P</emphasis>
. Tarsus of a midbody leg, lateral view.
<emphasis id="49E9EAD4FFDCFFF1FD52F841FCBF1E9A" bold="true" box="[701,889,1920,1945]" pageId="7" pageNumber="104">Abbreviations: et</emphasis>
, external tooth;
<emphasis id="49E9EAD4FFDCFFF1FBCFF841FBE91E9B" bold="true" box="[1056,1071,1920,1944]" pageId="7" pageNumber="104">it</emphasis>
, internal teeth;
<emphasis id="49E9EAD4FFDCFFF1FB20F840FB181E9A" bold="true" box="[1231,1246,1921,1945]" pageId="7" pageNumber="104">lt</emphasis>
, lateral tooth;
<emphasis id="49E9EAD4FFDCFFF1FA9CF840FA501E9A" bold="true" box="[1395,1430,1921,1945]" pageId="7" pageNumber="104">mp</emphasis>
, molar plate
<emphasis id="49E9EAD4FFDCFFF1FEFEF864FEEF1EBE" bold="true" box="[273,297,1957,1981]" pageId="7" pageNumber="104">oz</emphasis>
, ozopore;
<emphasis id="49E9EAD4FFDCFFF1FE7AF864FE761EBE" bold="true" box="[405,432,1957,1981]" pageId="7" pageNumber="104">pa</emphasis>
, tarsal pad;
<emphasis id="49E9EAD4FFDCFFF1FDC2F864FD841EBE" bold="true" box="[557,578,1957,1981]" pageId="7" pageNumber="104">pl</emphasis>
, pectinate lamella.
<emphasis id="49E9EAD4FFDCFFF1FCE6F865FC471EBE" bold="true" box="[777,897,1956,1981]" pageId="7" pageNumber="104">Scale bars:</emphasis>
500 µm (
<emphasis id="49E9EAD4FFDCFFF1FC09F865FC3E1EBF" bold="true" box="[998,1016,1956,1980]" pageId="7" pageNumber="104">A</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FBEAF864FBD01EBE" bold="true" box="[1029,1046,1957,1981]" pageId="7" pageNumber="104">B</emphasis>
), 100 µm (
<emphasis id="49E9EAD4FFDCFFF1FB65F864FB5A1EBE" bold="true" box="[1162,1180,1957,1981]" pageId="7" pageNumber="104">D</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FB46F864FB7C1EBE" bold="true" box="[1193,1210,1957,1981]" pageId="7" pageNumber="104">E</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FB28F864FB101EBE" bold="true" box="[1223,1238,1957,1981]" pageId="7" pageNumber="104">F</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FB0BF865FB3E1EBF" bold="true" box="[1252,1272,1956,1980]" pageId="7" pageNumber="104">G</emphasis>
), 50 µm (
<emphasis id="49E9EAD4FFDCFFF1FAB0F865FAB71EBF" bold="true" box="[1375,1393,1956,1980]" pageId="7" pageNumber="104">C</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FA91F864FA4B1EBE" bold="true" box="[1406,1421,1957,1981]" pageId="7" pageNumber="104">P</emphasis>
), 200 µm (
<emphasis id="49E9EAD4FFDCFFF1FF1AF808FEC01EE2" bold="true" box="[245,262,1993,2017]" pageId="7" pageNumber="104">L</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FEFDF808FEE01EE2" bold="true" box="[274,294,1993,2017]" pageId="7" pageNumber="104">H</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FEDCF808FE861EE2" bold="true" box="[307,320,1993,2017]" pageId="7" pageNumber="104">J</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FEA3F808FEA61EE2" bold="true" box="[332,352,1993,2017]" pageId="7" pageNumber="104">K</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FE83F808FEB81EE2" bold="true" box="[364,382,1993,2017]" pageId="7" pageNumber="104">N</emphasis>
,
<emphasis id="49E9EAD4FFDCFFF1FE64F809FE591EE3" bold="true" box="[395,415,1992,2016]" pageId="7" pageNumber="104">O</emphasis>
), 10 µm (
<emphasis id="49E9EAD4FFDCFFF1FDEAF808FDD01EE2" bold="true" box="[517,534,1993,2017]" pageId="7" pageNumber="104">L</emphasis>
).
</paragraph>
</caption>
<caption id="2FE2664EFFD3FFFEFF78F931FCEC1EE2" ID-DOI="http://doi.org/10.5281/zenodo.6610060" ID-Zenodo-Dep="6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="8" pageNumber="105" startId="8.[151,250,1776,1800]" targetBox="[238,1349,181,1753]" targetPageId="8">
<paragraph id="7B2236C6FFD3FFFEFF78F931FCEC1EE2" blockId="8.[151,1437,1776,2017]" pageId="8" pageNumber="105">
<emphasis id="49E9EAD4FFD3FFFEFF78F931FECB1E0B" bold="true" box="[151,269,1776,1800]" pageId="8" pageNumber="105">FIGURE 6</emphasis>
.
<taxonomicName id="BC9D4D45FFD3FFFEFEF6F930FE261E0A" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[281,480,1777,1801]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="8" pageNumber="105" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFD3FFFEFEF6F930FE261E0A" box="[281,480,1777,1801]" italics="true" pageId="8" pageNumber="105">Costabolus baculus</emphasis>
</taxonomicName>
<emphasis id="49E9EAD4FFD3FFFEFE09F930FDF01E0A" bold="true" box="[486,566,1777,1801]" pageId="8" pageNumber="105">
<taxonomicNameLabel id="52DA57AFFFD3FFFEFE09F930FDF01E0A" box="[486,566,1777,1801]" pageId="8" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (VNHM 198).
<emphasis id="49E9EAD4FFD3FFFEFC81F931FC461E0B" bold="true" box="[878,896,1776,1800]" pageId="8" pageNumber="105">A</emphasis>
. Anterior part of body, ventral view.
<emphasis id="49E9EAD4FFD3FFFEFB1AF930FAC01E0A" bold="true" box="[1269,1286,1777,1801]" pageId="8" pageNumber="105">B</emphasis>
. Coxal projections on leg-pairs 2 and 3.
<emphasis id="49E9EAD4FFD3FFFEFE44F8D5FE7B1E2F" bold="true" box="[427,445,1812,1836]" pageId="8" pageNumber="105">C</emphasis>
. Leg-pair 2, posterior view.
<emphasis id="49E9EAD4FFD3FFFEFD0CF8D4FD331E2E" bold="true" box="[739,757,1813,1837]" pageId="8" pageNumber="105">D</emphasis>
. Leg-pair 3, posterior view.
<emphasis id="49E9EAD4FFD3FFFEFBF4F8D4FBF81E2E" bold="true" box="[1051,1086,1813,1837]" pageId="8" pageNumber="105">E-I</emphasis>
. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively.
<emphasis id="49E9EAD4FFD3FFFEFC8BF8F8FCB71E52" bold="true" box="[868,881,1849,1873]" pageId="8" pageNumber="105">J</emphasis>
. Right posterior gonopod, lateral view.
<emphasis id="49E9EAD4FFD3FFFEFAE8F8F8FAE71E52" bold="true" box="[1287,1313,1849,1873]" pageId="8" pageNumber="105">K.</emphasis>
Drawing of left posterior gonopod, caudal view.
<emphasis id="49E9EAD4FFD3FFFEFDFBF89DFD6B1E77" bold="true" box="[532,685,1884,1908]" pageId="8" pageNumber="105">Abbreviations</emphasis>
:
<emphasis id="49E9EAD4FFD3FFFEFD52F89DFD0C1E77" bold="true" box="[701,714,1884,1908]" pageId="8" pageNumber="105">1</emphasis>
, median membranous area of telopodite;
<emphasis id="49E9EAD4FFD3FFFEFB94F89DFB4E1E77" bold="true" box="[1147,1160,1884,1908]" pageId="8" pageNumber="105">2</emphasis>
, lateral sclerotized area of telopodite;
<emphasis id="49E9EAD4FFD3FFFEFEE4F841FEF61E9B" bold="true" box="[267,304,1920,1944]" pageId="8" pageNumber="105">cx1</emphasis>
, coxite of anterior gonopod;
<emphasis id="49E9EAD4FFD3FFFEFD8FF841FD421E9B" bold="true" box="[608,644,1920,1944]" pageId="8" pageNumber="105">cx2</emphasis>
, coxite of posterior gonopod;
<emphasis id="49E9EAD4FFD3FFFEFC50F840FC1C1E9A" bold="true" box="[959,986,1921,1945]" pageId="8" pageNumber="105">pg</emphasis>
, posterior gonopod;
<emphasis id="49E9EAD4FFD3FFFEFB5DF840FB081E9A" bold="true" box="[1202,1230,1921,1945]" pageId="8" pageNumber="105">hp</emphasis>
, basal projection of prefemur 2;
<emphasis id="49E9EAD4FFD3FFFEFEFBF864FEE01EBE" bold="true" box="[276,294,1957,1981]" pageId="8" pageNumber="105">st</emphasis>
, sternite;
<emphasis id="49E9EAD4FFD3FFFEFE65F864FE5D1EBE" bold="true" box="[394,411,1957,1981]" pageId="8" pageNumber="105">sl</emphasis>
, solenomere;
<emphasis id="49E9EAD4FFD3FFFEFDC6F865FDF91EBF" bold="true" box="[553,575,1956,1980]" pageId="8" pageNumber="105">t1</emphasis>
and
<emphasis id="49E9EAD4FFD3FFFEFD80F865FD421EBF" bold="true" box="[623,644,1956,1980]" pageId="8" pageNumber="105">t2</emphasis>
, telopoditomeres 1 and 2 of anterior gonopod;
<emphasis id="49E9EAD4FFD3FFFEFBB0F864FBA81EBE" bold="true" box="[1119,1134,1957,1981]" pageId="8" pageNumber="105">tl</emphasis>
, telopodite of posterior gonopod.
<emphasis id="49E9EAD4FFD3FFFEFF25F809FE871EE2" bold="true" box="[202,321,1992,2017]" pageId="8" pageNumber="105">Scale bars:</emphasis>
500 µm (
<emphasis id="49E9EAD4FFD3FFFEFE4BF809FE701EE3" bold="true" box="[420,438,1992,2016]" pageId="8" pageNumber="105">A</emphasis>
,
<emphasis id="49E9EAD4FFD3FFFEFE2CF809FE131EE3" bold="true" box="[451,469,1992,2016]" pageId="8" pageNumber="105">C</emphasis>
), 100 µm (
<emphasis id="49E9EAD4FFD3FFFEFDA7F808FD9F1EE2" bold="true" box="[584,601,1993,2017]" pageId="8" pageNumber="105">B</emphasis>
), 200 µm (
<emphasis id="49E9EAD4FFD3FFFEFD23F808FD181EE2" bold="true" box="[716,734,1993,2017]" pageId="8" pageNumber="105">D</emphasis>
,
<emphasis id="49E9EAD4FFD3FFFEFD04F808FCDA1EE2" bold="true" box="[747,796,1993,2017]" pageId="8" pageNumber="105">E–K</emphasis>
).
</paragraph>
</caption>
<caption id="2FE2664EFFD2FFFFFF78FD9CFE5F1BBD" ID-DOI="http://doi.org/10.5281/zenodo.6610064" ID-Zenodo-Dep="6610064" httpUri="https://zenodo.org/record/6610064/files/figure.png" pageId="9" pageNumber="106" startId="9.[151,250,605,629]" targetBox="[151,1436,181,582]" targetPageId="9">
<paragraph id="7B2236C6FFD2FFFFFF78FD9CFE5F1BBD" blockId="9.[151,1436,605,702]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF78FD9CFECB1B75" bold="true" box="[151,269,605,630]" pageId="9" pageNumber="106">FIGURE 7</emphasis>
.
<taxonomicName id="BC9D4D45FFD2FFFFFEF6FD9FFE271B75" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[281,481,606,630]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFD2FFFFFEF6FD9FFE271B75" box="[281,481,606,630]" italics="true" pageId="9" pageNumber="106">Costabolus baculus</emphasis>
</taxonomicName>
<emphasis id="49E9EAD4FFD2FFFFFE08FD9FFDFE1B75" bold="true" box="[487,568,606,630]" pageId="9" pageNumber="106">
<taxonomicNameLabel id="52DA57AFFFD2FFFFFE08FD9FFDFE1B75" box="[487,568,606,630]" pageId="9" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype (VMNH 154).
<emphasis id="49E9EAD4FFD2FFFFFC6AFD9CFC511B76" bold="true" box="[901,919,605,629]" pageId="9" pageNumber="106">A</emphasis>
. Ventral view of both vulvae,
<emphasis id="49E9EAD4FFD2FFFFFB25FD9FFB1D1B75" bold="true" box="[1226,1243,606,630]" pageId="9" pageNumber="106">B</emphasis>
. Left vulva, lateral view.
<emphasis id="49E9EAD4FFD2FFFFFF3CFD40FE551B99" bold="true" box="[211,403,641,666]" pageId="9" pageNumber="106">Abbreviations: av</emphasis>
, anterior valve;
<emphasis id="49E9EAD4FFD2FFFFFDD8FD43FD891B99" bold="true" box="[567,591,642,666]" pageId="9" pageNumber="106">cx</emphasis>
, coxite of leg 1;
<emphasis id="49E9EAD4FFD2FFFFFD18FD43FCDA1B99" bold="true" box="[759,796,642,666]" pageId="9" pageNumber="106">cxv</emphasis>
, coxite of vulva;
<emphasis id="49E9EAD4FFD2FFFFFC24FD43FC1E1B99" bold="true" box="[971,984,642,666]" pageId="9" pageNumber="106">o</emphasis>
, operculum;
<emphasis id="49E9EAD4FFD2FFFFFBB2FD43FBBE1B99" bold="true" box="[1117,1144,642,666]" pageId="9" pageNumber="106">pv</emphasis>
, posterior valve.
<emphasis id="49E9EAD4FFD2FFFFFAC9FD40FA5A1B99" bold="true" box="[1318,1436,641,666]" pageId="9" pageNumber="106">Scale bars:</emphasis>
500 µm (
<emphasis id="49E9EAD4FFD2FFFFFF1AFD64FEC11BBE" bold="true" box="[245,263,677,701]" pageId="9" pageNumber="106">A</emphasis>
), 200 µm (
<emphasis id="49E9EAD4FFD2FFFFFE96FD67FE4C1BBD" bold="true" box="[377,394,678,702]" pageId="9" pageNumber="106">B</emphasis>
).
</paragraph>
</caption>
<subSubSection id="3387654DFFD2FFFCFF28FD2DFE0518AE" lastPageId="10" lastPageNumber="107" pageId="9" pageNumber="106" type="description">
<paragraph id="7B2236C6FFD2FFFFFF28FD2DFD6F1A29" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FD2DFE9F1A05" bold="true" box="[199,345,748,774]" pageId="9" pageNumber="106">Description.</emphasis>
<emphasis id="49E9EAD4FFD2FFFFFEB0FD2CFDC71A05" box="[351,513,749,774]" italics="true" pageId="9" pageNumber="106">Measurements</emphasis>
: male
<typeStatus id="A4268864FFD2FFFFFDA3FD2DFD681A05" box="[588,686,748,774]" pageId="9" pageNumber="106" type="holotype">holotype</typeStatus>
with 39 body rings (38 + telson), ca
<quantity id="BC659B23FFD2FFFFFBAAFD2DFB551A05" box="[1093,1171,748,774]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="106" unit="mm" value="27.0">27 mm</quantity>
long,
<quantity id="BC659B23FFD2FFFFFB37FD2DFADE1A05" box="[1240,1304,748,774]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="106" unit="mm" value="3.0">3 mm</quantity>
wide.
<typeStatus id="A4268864FFD2FFFFFA8EFD2DFF131A29" pageId="9" pageNumber="106" type="paratype">Paratypes</typeStatus>
, both male and female, same as
<typeStatus id="A4268864FFD2FFFFFDD0FCD1FD631A29" box="[575,677,784,810]" pageId="9" pageNumber="106" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FCF5FB031A71" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FCF5FE841A4D" box="[199,322,820,846]" italics="true" pageId="9" pageNumber="106">Coloration</emphasis>
faded in alcohol, but a darker pattern still visible dorsally and laterally, yellowish ventrally. Head, ommatidia and anterior part of collum dark brown, antennae, legs and telson yellowish (
<figureCitation id="E3A62A43FFD2FFFFFB9CFC99FB7E1A71" box="[1139,1208,856,882]" captionStart="FIGURE 4" captionStartId="6.[151,250,1942,1966]" captionTargetBox="[151,1436,669,1918]" captionTargetId="figure-185@6.[151,1436,669,1918]" captionTargetPageId="6" captionText="FIGURE 4. Costabolus baculus sp. nov., male holotype (VNHM 198). A &amp; B. Habitus, lateral and ventral views, respectively. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6610051" httpUri="https://zenodo.org/record/6610051/files/figure.png" pageId="9" pageNumber="106">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FCBDFE201AB8" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FCBDFEC21A95" box="[199,260,892,918]" italics="true" pageId="9" pageNumber="106">Head</emphasis>
capsule with an axial suture, this being especially distinct on labrum; ca 22 ommatidia located in an irregular oval cluster (
<figureCitation id="E3A62A43FFD2FFFFFE90FC61FE1F1AB8" box="[383,473,928,955]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FC05FB991D6C" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FC05FEE61ADD" box="[199,288,964,990]" italics="true" pageId="9" pageNumber="106">Labrum</emphasis>
as usual, with three irregular teeth and a single row of 12 stout marginal setae. Clypeus with two setiferous foveolae on each side (
<figureCitation id="E3A62A43FFD2FFFFFE2DFC29FDDE1D00" box="[450,536,1000,1027]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5E</figureCitation>
). Incisura lateralis closed (
<figureCitation id="E3A62A43FFD2FFFFFCAEFC29FC7A1D01" box="[833,956,1000,1027]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5A, D</figureCitation>
). Antennal cavity/groove present, length of antennae ca
<quantity id="BC659B23FFD2FFFFFECBFBCDFEB81D25" box="[292,382,1036,1062]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="106" unit="mm" value="1.3">1.3 mm</quantity>
; antennae shorter than body diameter. Relative lengths of antennomeres: 3=4=5&lt;1&lt;6&lt;2 (
<figureCitation id="E3A62A43FFD2FFFFFA80FBCDFF7A1D48" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5D</figureCitation>
). Terminal antennomere with four large sensory cones clustered together inside a membranous area (
<figureCitation id="E3A62A43FFD2FFFFFAD9FBF1FA561D48" box="[1334,1424,1072,1099]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5C</figureCitation>
). Antennomeres 5 and 6 each with an apicolateral field of specialized sensilla (
<figureCitation id="E3A62A43FFD2FFFFFC17FB95FB941D6C" box="[1016,1106,1108,1135]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FBB9FCAE1DFC" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FBB9FE4E1D91" box="[199,392,1144,1170]" italics="true" pageId="9" pageNumber="106">Gnathochilarium</emphasis>
as usual, of spirobolidan conformation. Stipites separated in basal portion, each bearing three apical setae, but no setae on ventral surface. Lamellae linguales separated by anterior portion of mentum, each with two usual setae located behind one another; mentum without a large swollen area apically between both lamellae linguales, subdivided basally by a well-marked suture (
<figureCitation id="E3A62A43FFD2FFFFFCEEFB25FC9A1DFC" box="[769,860,1252,1279]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FAC9FB711C45" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FAC9FEFA1C21" box="[199,316,1288,1314]" italics="true" pageId="9" pageNumber="106">Mandible:</emphasis>
external tooth (
<emphasis id="49E9EAD4FFD2FFFFFE00FAC9FDC31C21" bold="true" box="[495,517,1288,1314]" pageId="9" pageNumber="106">et</emphasis>
) prominent, inner tooth (
<emphasis id="49E9EAD4FFD2FFFFFCCDFAC9FCF51C21" bold="true" box="[802,819,1288,1314]" pageId="9" pageNumber="106">it</emphasis>
) with three smaller inner teeth, a thin lateral tooth (
<emphasis id="49E9EAD4FFD2FFFFFA94FAC9FA4A1C21" bold="true" box="[1403,1420,1288,1314]" pageId="9" pageNumber="106">lt</emphasis>
), six rows of pectinate lamellae (
<emphasis id="49E9EAD4FFD2FFFFFE19FAEDFDCB1C45" bold="true" box="[502,525,1324,1350]" pageId="9" pageNumber="106">pl</emphasis>
), molar plate (
<emphasis id="49E9EAD4FFD2FFFFFD5BFAEDFD1D1C45" bold="true" box="[692,731,1324,1350]" pageId="9" pageNumber="106">mp</emphasis>
) with five transverse furrows (
<figureCitation id="E3A62A43FFD2FFFFFBDAFAEDFB6D1C45" box="[1077,1195,1324,1351]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5F, G</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FA91FC851C8C" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FA91FEDF1C69" box="[199,281,1360,1386]" italics="true" pageId="9" pageNumber="106">Collum</emphasis>
with a prominent groove along anterior margin, lateroventral margin broadly rounded, not extending ventrad as far as ventrolateral corner of body ring 2 (
<figureCitation id="E3A62A43FFD2FFFFFD08FAB5FCF31C8C" box="[743,821,1396,1423]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5I</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28FA59FB1A1F41" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28FA59FE891CB1" box="[199,335,1432,1458]" italics="true" pageId="9" pageNumber="106">Body rings:</emphasis>
Metazonae heavily crested (
<figureCitation id="E3A62A43FFD2FFFFFD70FA59FCD21CB0" box="[671,788,1432,1459]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5J–L</figureCitation>
); mesozonae with longitudinal impressions, these being especially distinct ventrally (
<figureCitation id="E3A62A43FFD2FFFFFE0BFA7DFD871CD4" box="[484,577,1468,1495]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5K</figureCitation>
), caudal fringe/limbus membranous, with very small and regularly spaced spinicles (
<figureCitation id="E3A62A43FFD2FFFFFEE6FA21FE991CF8" box="[265,351,1504,1531]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5L</figureCitation>
). Pre-anal ring with a small dorsal tip of epiproct overhanging the paraprocts (
<figureCitation id="E3A62A43FFD2FFFFFB21FA21FAEA1CF9" box="[1230,1324,1504,1530]" captionStart="FIGURE 4" captionStartId="6.[151,250,1942,1966]" captionTargetBox="[151,1436,669,1918]" captionTargetId="figure-185@6.[151,1436,669,1918]" captionTargetPageId="6" captionText="FIGURE 4. Costabolus baculus sp. nov., male holotype (VNHM 198). A &amp; B. Habitus, lateral and ventral views, respectively. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6610051" httpUri="https://zenodo.org/record/6610051/files/figure.png" pageId="9" pageNumber="106">Figs 4A</figureCitation>
,
<figureCitation id="E3A62A43FFD2FFFFFAD7FA21FAA41CF8" box="[1336,1378,1504,1531]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">5M</figureCitation>
); hypoproct rounded; paraprocts (anal valves) well-rounded and devoid of spines or setae (
<figureCitation id="E3A62A43FFD2FFFFFBBBF9C5FB751F1C" box="[1108,1203,1540,1567]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5M</figureCitation>
). Posterior margin of paraprocts in lateral view without distinct lips (
<figureCitation id="E3A62A43FFD2FFFFFD47F9E9FCC11F40" box="[680,775,1576,1603]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5M</figureCitation>
). Ozopores (
<emphasis id="49E9EAD4FFD2FFFFFC79F9E9FC761F41" bold="true" box="[918,944,1576,1602]" pageId="9" pageNumber="106">oz</emphasis>
) starting with body ring 6.
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28F98CFAE31FF5" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28F98CFEC01F65" box="[199,262,1613,1638]" italics="true" pageId="9" pageNumber="106">Legs:</emphasis>
Midbody legs ca
<quantity id="BC659B23FFD2FFFFFE21F98DFDE21F65" box="[462,548,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="106" unit="mm" value="1.4">1.4 mm</quantity>
long, with prefemur longer than coxa and as long as the other podomeres; with small tarsal pads (
<emphasis id="49E9EAD4FFD2FFFFFE8DF9B1FE461F89" bold="true" box="[354,384,1648,1674]" pageId="9" pageNumber="106">pa</emphasis>
) (
<figureCitation id="E3A62A43FFD2FFFFFE77F9B1FDC91F89" box="[408,527,1648,1675]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="9" pageNumber="106">Fig. 5O, P</figureCitation>
). Male tarsi 3 up to midbody legs with a small tarsal pad not protruding past base of claw, and with a pair of setiform apical spines (
<figureCitation id="E3A62A43FFD2FFFFFD2EF955FCDD1FAC" box="[705,795,1684,1711]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6D</figureCitation>
). Male leg-pairs 1, 2 and 3 modified, coxae 1 fused medially at base only, each prefemur 2 with a well-developed, unciform, basal projection (
<emphasis id="49E9EAD4FFD2FFFFFBBBF979FBB51FD1" bold="true" box="[1108,1139,1720,1746]" pageId="9" pageNumber="106">hp</emphasis>
) (
<figureCitation id="E3A62A43FFD2FFFFFB64F979FAC51FD1" box="[1163,1283,1720,1746]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6B, C</figureCitation>
), each coxa 3 with a long and spoon-shaped coxal process (
<emphasis id="49E9EAD4FFD2FFFFFD7BF91DFD761FF5" bold="true" box="[660,688,1756,1782]" pageId="9" pageNumber="106">cp</emphasis>
), this being almost as long as the leg itself (
<figureCitation id="E3A62A43FFD2FFFFFB72F91DFADE1FF5" box="[1181,1304,1756,1782]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6B, D</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFFFF28F8C1FEC11EA9" blockId="9.[151,1437,748,2035]" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28F8C1FE5D1E19" box="[199,411,1792,1818]" italics="true" pageId="9" pageNumber="106">Anterior gonopods</emphasis>
stout and relatively simple (
<figureCitation id="E3A62A43FFD2FFFFFD33F8C1FC8A1E18" box="[732,844,1792,1819]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6E–I</figureCitation>
); sternite (
<emphasis id="49E9EAD4FFD2FFFFFC2CF8C1FC111E19" bold="true" box="[963,983,1792,1818]" pageId="9" pageNumber="106">st</emphasis>
) produced into a wide, broadly rounded lobe, bearing clear traces of axial fusion (
<figureCitation id="E3A62A43FFD2FFFFFD87F8E5FD791E3C" box="[616,703,1828,1855]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6E</figureCitation>
); coxite (
<emphasis id="49E9EAD4FFD2FFFFFCC6F8E5FC971E3D" bold="true" box="[809,849,1828,1854]" pageId="9" pageNumber="106">cx1</emphasis>
) subtending much of anterior gonopods; telopodites basically 2-segmented, each consisting of a smaller, roundish, apically flattened, basal telopoditomere (
<emphasis id="49E9EAD4FFD2FFFFFAC2F889FA821E61" bold="true" box="[1325,1348,1864,1890]" pageId="9" pageNumber="106">t1</emphasis>
) with a very small, accessory, subtriangular segment at base (
<figureCitation id="E3A62A43FFD2FFFFFD01F8ADFC981E85" box="[750,862,1900,1926]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6H, I</figureCitation>
), and a much larger, lobe-shaped, distal telopoditomere (
<emphasis id="49E9EAD4FFD2FFFFFF30F851FF311EA9" bold="true" box="[223,247,1936,1962]" pageId="9" pageNumber="106">t2</emphasis>
).
</paragraph>
<paragraph id="7B2236C6FFD2FFFCFF28F874FC8219FA" blockId="9.[151,1437,748,2035]" lastBlockId="10.[151,1437,151,429]" lastPageId="10" lastPageNumber="107" pageId="9" pageNumber="106">
<emphasis id="49E9EAD4FFD2FFFFFF28F874FE631ECD" box="[199,421,1972,1998]" italics="true" pageId="9" pageNumber="106">Posterior gonopods</emphasis>
(
<figureCitation id="E3A62A43FFD2FFFFFE5BF875FDE01ECD" box="[436,550,1972,1998]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6J, K</figureCitation>
,
<emphasis id="49E9EAD4FFD2FFFFFDDEF875FD891ECD" bold="true" box="[561,591,1972,1998]" pageId="9" pageNumber="106">pg</emphasis>
in H)
<emphasis id="49E9EAD4FFD2FFFFFD79F874FD1D1ECD" box="[662,731,1973,1998]" italics="true" pageId="9" pageNumber="106">in situ</emphasis>
almost completely concealed inside anterior gonopods, clearly divided into a slender coxite (
<emphasis id="49E9EAD4FFD2FFFFFE1FF819FDDF1EF1" bold="true" box="[496,537,2008,2034]" pageId="9" pageNumber="106">cx2</emphasis>
) and a shorter telopodite (
<emphasis id="49E9EAD4FFD2FFFFFCA3F819FC9B1EF1" bold="true" box="[844,861,2008,2034]" pageId="9" pageNumber="106">tl</emphasis>
), both being slender (
<figureCitation id="E3A62A43FFD2FFFFFBB3F819FB6B1EF0" box="[1116,1197,2008,2035]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="9" pageNumber="106">Fig. 6J</figureCitation>
) and connected by a tiny triangular sternite. Telopodite consisting of two components: (1) a median membranous area, and (2) a lateral sclerotized area. Sperm channel running at mesal margin of coxite through telopodite’s membranous area, ending up subapically on a rudimentary solenomere (
<emphasis id="49E9EAD4FFD1FFFCFD77FF1EFD6D19FA" bold="true" box="[664,683,223,249]" pageId="10" pageNumber="107">sl</emphasis>
) (
<figureCitation id="E3A62A43FFD1FFFCFD2BFF1EFCF119FA" box="[708,823,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="10" pageNumber="107">Fig. 6J, K</figureCitation>
).
</paragraph>
<paragraph id="7B2236C6FFD1FFFCFF28FEC2FE0518AE" blockId="10.[151,1437,151,429]" pageId="10" pageNumber="107">
<emphasis id="49E9EAD4FFD1FFFCFF28FEC2FE4B181E" box="[199,397,259,285]" italics="true" pageId="10" pageNumber="107">
<typeStatus id="A4268864FFD1FFFCFF28FEC2FEFA181F" box="[199,316,259,284]" pageId="10" pageNumber="107" type="paratype">Paratypes</typeStatus>
. Males
</emphasis>
as
<typeStatus id="A4268864FFD1FFFCFE57FEC2FDD8181E" box="[440,542,259,285]" pageId="10" pageNumber="107" type="holotype">holotype</typeStatus>
.
<emphasis id="49E9EAD4FFD1FFFCFDC5FEC2FDB8181E" box="[554,638,259,285]" italics="true" pageId="10" pageNumber="107">Female</emphasis>
similar in size and appearance to males, but devoid of prefemoral or coxal projections on leg-pairs 2 and 3, respectively. Vulva simple, consisting of two simple, sclerotized plates, bivalve-like and subequal in size (
<figureCitation id="E3A62A43FFD1FFFCFE15FE8AFDFB1866" box="[506,573,331,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,605,629]" captionTargetBox="[151,1436,181,582]" captionTargetId="figure-21@9.[151,1436,181,582]" captionTargetPageId="9" captionText="FIGURE 7. Costabolus baculus sp. nov., female paratype (VMNH 154). A. Ventral view of both vulvae, B. Left vulva, lateral view. Abbreviations: av, anterior valve; cx, coxite of leg 1; cxv, coxite of vulva; o, operculum; pv, posterior valve. Scale bars: 500 µm (A), 200 µm (B)." figureDoi="http://doi.org/10.5281/zenodo.6610064" httpUri="https://zenodo.org/record/6610064/files/figure.png" pageId="10" pageNumber="107">Fig. 7</figureCitation>
). Both coxite (
<emphasis id="49E9EAD4FFD1FFFCFD0EFE8AFCCF1866" bold="true" box="[737,777,331,357]" pageId="10" pageNumber="107">cxv</emphasis>
) and operculum (
<emphasis id="49E9EAD4FFD1FFFCFC21FE8AFC1A1866" bold="true" box="[974,988,331,357]" pageId="10" pageNumber="107">o</emphasis>
) of vulva very small and inconspicuous. Both valves only basally with one row of setae directed towards the opening. Anterior valve (
<emphasis id="49E9EAD4FFD1FFFCFB4FFEAEFB7A188A" bold="true" box="[1184,1212,367,393]" pageId="10" pageNumber="107">av</emphasis>
) slightly larger than posterior one (
<emphasis id="49E9EAD4FFD1FFFCFED5FE52FE9E18AE" bold="true" box="[314,344,403,429]" pageId="10" pageNumber="107">pv</emphasis>
) (
<figureCitation id="E3A62A43FFD1FFFCFE9EFE52FE7318AE" box="[369,437,403,429]" captionStart="FIGURE 7" captionStartId="9.[151,250,605,629]" captionTargetBox="[151,1436,181,582]" captionTargetId="figure-21@9.[151,1436,181,582]" captionTargetPageId="9" captionText="FIGURE 7. Costabolus baculus sp. nov., female paratype (VMNH 154). A. Ventral view of both vulvae, B. Left vulva, lateral view. Abbreviations: av, anterior valve; cx, coxite of leg 1; cxv, coxite of vulva; o, operculum; pv, posterior valve. Scale bars: 500 µm (A), 200 µm (B)." figureDoi="http://doi.org/10.5281/zenodo.6610064" httpUri="https://zenodo.org/record/6610064/files/figure.png" pageId="10" pageNumber="107">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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