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<document id="C147E07D504979DDC6E49EFC287F59C4" ID-CLB-Dataset="63555" ID-DOI="10.11646/zootaxa.5150.1.5" ID-GBIF-Dataset="0e9631c5-3211-4f61-9549-db14a51dbc30" ID-ISSN="1175-5326" ID-Zenodo-Dep="6610039" ID-ZooBank="2F6FD286-5E1E-40F7-B658-490EFD453BDF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654245021749" checkinUser="plazi" docAuthor="Golovatch, Sergei &amp; Vandenspiegel, Didier" docDate="2022" docId="F33487D0FFD0FFFDFF78FEAEFEF81A82" docLanguage="en" docName="zootaxa.5150.1.5.pdf" docOrigin="Zootaxa 5150 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Costabolus Golovatch &amp; Vandenspiegel 2022, gen. nov." docType="treatment" docVersion="7" lastPageNumber="108" masterDocId="0F0DFFA8FFDBFFF6FFEFFFC1FFC61903" masterDocTitle="Carinobolus gen. nov. and Costabolus gen. nov., two new, remarkably crested monospecific genera of spirobolidan millipedes from WestAfrica, with the proposal of a new tribe, Amblybolini tribus nova (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="110" masterPageNumber="97" pageNumber="108" updateTime="1699338865200" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="09C910EED099FA165FDC899AEBD57EF1">Carinobolus gen. nov. and Costabolus gen. nov., two new, remarkably crested monospecific genera of spirobolidan millipedes from WestAfrica, with the proposal of a new tribe, Amblybolini tribus nova (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="7CE6D6196DAAE483872077202867BA87">Golovatch, Sergei</mods:namePart>
<mods:affiliation id="8F4E1CC865C97EE56E4FC73767A8AAC5">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow, 119071 Russia. Biological Collection and Data Management Unit, Royal Museum for Central Africa, Tervuren, Belgium. didier. van. den. spiegel @ africamuseum. be; https: // orcid. org / 0000 - 0002 - 8696 - 9810</mods:affiliation>
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<mods:namePart id="9F670BE81551DF59FA9CBA705117D591">Vandenspiegel, Didier</mods:namePart>
<mods:nameIdentifier id="FF5BFB0AAEDCE98082DF28F3B3C95084" type="ORCID">0000-0002-8696-9810</mods:nameIdentifier>
<mods:nameIdentifier id="896C18EE2D1F39663F25AAF23F06918B" type="email">didier.van.den.spiegel@africamuseum.be</mods:nameIdentifier>
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<mods:title id="B3A137BE110F9BC533C642040EA5C7D0">Zootaxa</mods:title>
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<mods:date id="09AF11B8D8095543EC32E42F798CE69A">2022</mods:date>
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<mods:number id="11C72F4B84ECED8D80E33B3F26181DEB">2022-06-03</mods:number>
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<mods:detail id="565B48F0E78E643FD8B6CF25933197A3" type="volume">
<mods:number id="94863B3587B35366CF962CC4F4A7DD6C">5150</mods:number>
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<mods:classification id="9DF2FBD18C5E7757403359BA51A8892D">journal article</mods:classification>
<mods:identifier id="6B62227DEA4782D30A2D38A76EC7C9A2" type="CLB-Dataset">63555</mods:identifier>
<mods:identifier id="C6031EF86EBD924128D4E149AF751529" type="DOI">10.11646/zootaxa.5150.1.5</mods:identifier>
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<treatment id="F33487D0FFD0FFFDFF78FEAEFEF81A82" ID-DOI="http://doi.org/10.5281/zenodo.6611459" ID-GBIF-Taxon="195730568" ID-Zenodo-Dep="6611459" LSID="urn:lsid:plazi:treatment:F33487D0FFD0FFFDFF78FEAEFEF81A82" httpUri="http://treatment.plazi.org/id/F33487D0FFD0FFFDFF78FEAEFEF81A82" lastPageNumber="108" pageId="11" pageNumber="108">
<subSubSection id="3387654DFFD0FFFDFF78FEAEFE54188A" box="[151,402,367,394]" pageId="11" pageNumber="108" type="nomenclature">
<paragraph id="7B2236C6FFD0FFFDFF78FEAEFE54188A" blockId="11.[151,402,367,394]" box="[151,402,367,394]" pageId="11" pageNumber="108">
<heading id="206A81AAFFD0FFFDFF78FEAEFE54188A" bold="true" box="[151,402,367,394]" fontSize="11" level="1" pageId="11" pageNumber="108" reason="1">
<emphasis id="49E9EAD4FFD0FFFDFF78FEAEFE54188A" bold="true" box="[151,402,367,394]" pageId="11" pageNumber="108">
<taxonomicName id="BC9D4D45FFD0FFFDFF78FEAEFEDB1889" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[151,285,367,394]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="108" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="49E9EAD4FFD0FFFDFF78FEAEFEDB1889" bold="true" box="[151,285,367,394]" italics="true" pageId="11" pageNumber="108">Costabolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52DA57AFFFD0FFFDFECBFEAEFE54188A" box="[292,402,367,393]" pageId="11" pageNumber="108" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3387654DFFD0FFFDFF78FE76FE1B18F6" pageId="11" pageNumber="108" type="type_taxon">
<paragraph id="7B2236C6FFD0FFFDFF78FE76FBEF18D2" blockId="11.[151,1065,439,502]" box="[151,1065,439,466]" pageId="11" pageNumber="108">
<materialsCitation id="CBF53C9BFFD0FFFDFF78FE76FBEF18D2" ID-GBIF-Occurrence="3802884305" box="[151,1065,439,466]" country="Nigeria" location="No" pageId="11" pageNumber="108" specimenCount="1" typeStatus="holotype">
<emphasis id="49E9EAD4FFD0FFFDFF78FE76FEF218D2" bold="true" box="[151,308,439,465]" pageId="11" pageNumber="108">
<typeStatus id="A4268864FFD0FFFDFF78FE76FF1418D2" box="[151,210,439,465]" pageId="11" pageNumber="108">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="BC9D4D45FFD0FFFDFED3FE76FDDE18D2" authority="Golovatch &amp; Vandenspiegel, 2022" authorityName="Golovatch &amp; Vandenspiegel" authorityYear="2022" box="[316,536,439,465]" class="Diplopoda" family="Pachybolidae" genus="Costabolus" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="108" phylum="Arthropoda" rank="species" species="baculus" status="sp. nov.">
<emphasis id="49E9EAD4FFD0FFFDFED3FE76FDDE18D2" box="[316,536,439,465]" italics="true" pageId="11" pageNumber="108">Costabolus baculus</emphasis>
</taxonomicName>
<emphasis id="49E9EAD4FFD0FFFDFDF0FE76FDBE18D2" bold="true" box="[543,632,439,465]" pageId="11" pageNumber="108">
<taxonomicNameLabel id="52DA57AFFFD0FFFDFDF0FE76FDBE18D2" box="[543,632,439,465]" pageId="11" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from
<collectingCountry id="038A7656FFD0FFFDFD2BFE76FCDD18D2" box="[708,795,439,465]" name="Nigeria" pageId="11" pageNumber="108">Nigeria</collectingCountry>
, by present designation.
</materialsCitation>
</paragraph>
<paragraph id="7B2236C6FFD0FFFDFF28FE1DFE1B18F6" blockId="11.[151,1065,439,502]" box="[199,477,475,502]" pageId="11" pageNumber="108">
<location id="7E42601DFFD0FFFDFF28FE1DFF2F18F5" LSID="urn:lsid:plazi:treatment:F33487D0FFD0FFFDFF78FEAEFEF81A82:7E42601DFFD0FFFDFF28FE1DFF2F18F5" box="[199,233,476,502]" country="Nigeria" name="No" pageId="11" pageNumber="108">No</location>
other species known.
</paragraph>
</subSubSection>
<subSubSection id="3387654DFFD0FFFDFF78FDE2FEF81A82" pageId="11" pageNumber="108" type="diagnosis">
<paragraph id="7B2236C6FFD0FFFDFF78FDE2FF1D1B62" blockId="11.[151,1437,547,897]" pageId="11" pageNumber="108">
<emphasis id="49E9EAD4FFD0FFFDFF78FDE2FECC1B3E" bold="true" box="[151,266,547,573]" pageId="11" pageNumber="108">Diagnosis</emphasis>
. A tribe of the family
<taxonomicName id="BC9D4D45FFD0FFFDFDEBFDE2FD5D1B3E" authorityName="Cook, 1897 sensu Hoffman" authorityYear="1980" box="[516,667,547,573]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="108" phylum="Arthropoda" rank="family">Pachybolidae</taxonomicName>
characterized by the following putative apomorphies or synapomorphies:
</paragraph>
<paragraph id="7B2236C6FFD0FFFDFF28FDADFA9A1B86" blockId="11.[151,1437,547,897]" box="[199,1372,619,646]" pageId="11" pageNumber="108">
- incisura lateralis closed (
<figureCitation id="E3A62A43FFD0FFFDFE02FDAAFD8F1B86" box="[493,585,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1563]" captionTargetId="figure-21@3.[151,1436,181,1563]" captionTargetPageId="3" captionText="FIGURE 2. Carinobolus complex sp. nov., male paratype (RMCA). A &amp; B. Head, frontal and lateral views, respectively. C. Tip of antenna, sublateral view. D. Gnathochilarium, ventral view. E, F &amp; G. Right mandible, submesal, mesal and subfrontal views, respectively. H. Anterior part of body, lateral view. I. Midbody rings with ozopores (oz), lateral view. J. A midbody metazona showing bimodal crests. K &amp; L. Posterior part of body, lateral and ventral views, respectively. M. Cross-section of midbody ring, frontal view. N. Details of a midbody ring, lateral view. O. Limbus of a midbody ring, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate; oz, ozopore; pl, pectinate lamella. Scale bars: 500 µm (A, I, K, L, M), 200 µm (B, D, H, J), 100 µm (C, E, F, G, N), 10 µm (O)." figureDoi="http://doi.org/10.5281/zenodo.6610043" httpUri="https://zenodo.org/record/6610043/files/figure.png" pageId="11" pageNumber="108">Figs 2B</figureCitation>
,
<figureCitation id="E3A62A43FFD0FFFDFDBAFDAAFDBC1B85" box="[597,634,619,646]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="11" pageNumber="108">5D</figureCitation>
), a synapomorphy shared with
<taxonomicName id="BC9D4D45FFD0FFFDFC37FDAAFB981B86" box="[984,1118,619,645]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="110" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">Pachybolini</taxonomicName>
(
<bibRefCitation id="1F0C4B37FFD0FFFDFB83FDAAFA971B86" author="Wesener, T. &amp; Enghoff, H. &amp; Wagele, J. - W." box="[1132,1361,619,645]" pageId="11" pageNumber="108" pagination="37 - 53" refId="ref7178" refString="Wesener, T., Enghoff, H. &amp; Wagele, J. - W. (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53. https: // doi. org / 10.1071 / IS 07008" type="journal article" year="2008">
Wesener
<emphasis id="49E9EAD4FFD0FFFDFB3AFDADFACB1B86" box="[1237,1293,619,645]" italics="true" pageId="11" pageNumber="108">et al.</emphasis>
2008
</bibRefCitation>
);
</paragraph>
<paragraph id="7B2236C6FFD0FFFDFF28FD51FF261BCE" blockId="11.[151,1437,547,897]" pageId="11" pageNumber="108">
- setae on lamellae linguales apical (
<figureCitation id="E3A62A43FFD0FFFDFD87FD4EFD0E1BAA" box="[616,712,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1587,1611]" captionTargetBox="[151,1436,181,1563]" captionTargetId="figure-21@3.[151,1436,181,1563]" captionTargetPageId="3" captionText="FIGURE 2. Carinobolus complex sp. nov., male paratype (RMCA). A &amp; B. Head, frontal and lateral views, respectively. C. Tip of antenna, sublateral view. D. Gnathochilarium, ventral view. E, F &amp; G. Right mandible, submesal, mesal and subfrontal views, respectively. H. Anterior part of body, lateral view. I. Midbody rings with ozopores (oz), lateral view. J. A midbody metazona showing bimodal crests. K &amp; L. Posterior part of body, lateral and ventral views, respectively. M. Cross-section of midbody ring, frontal view. N. Details of a midbody ring, lateral view. O. Limbus of a midbody ring, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate; oz, ozopore; pl, pectinate lamella. Scale bars: 500 µm (A, I, K, L, M), 200 µm (B, D, H, J), 100 µm (C, E, F, G, N), 10 µm (O)." figureDoi="http://doi.org/10.5281/zenodo.6610043" httpUri="https://zenodo.org/record/6610043/files/figure.png" pageId="11" pageNumber="108">Figs 2D</figureCitation>
,
<figureCitation id="E3A62A43FFD0FFFDFD3AFD4EFD3C1BA9" box="[725,762,655,682]" captionStart="FIGURE 5" captionStartId="7.[151,250,1776,1800]" captionTargetBox="[189,1398,181,1753]" captionTargetId="figure-21@7.[189,1398,181,1753]" captionTargetPageId="7" captionText="FIGURE 5. Costabolus baculus sp. nov., male paratype (VNHM 198). A &amp; B. Head, lateral and frontal views, respectively. C. Tip of antenna. D. Left antenna, lateral view. E. Clypeus, frontolateral view. F &amp; G. Mandible, ventral and mesal views, respectively. H. Gnathochilarium, ventral view. I. Right lateral margin of collum, lateral view. J. A midbody metazona. K. Ozopore. L. Limbus of a midbody ring. M. Posterior part of body, ventral view. N &amp; O. Midbody legs, ventral and lateral views, respectively. P. Tarsus of a midbody leg, lateral view. Abbreviations: et, external tooth; it, internal teeth; lt, lateral tooth; mp, molar plate oz, ozopore; pa, tarsal pad; pl, pectinate lamella. Scale bars: 500 µm (A, B), 100 µm (D, E, F, G), 50 µm (C, P), 200 µm (L, H, J, K, N, O), 10 µm (L)." figureDoi="http://doi.org/10.5281/zenodo.6610053" httpUri="https://zenodo.org/record/6610053/files/figure.png" pageId="11" pageNumber="108">5H</figureCitation>
), a synapomorphy shared with
<taxonomicName id="BC9D4D45FFD0FFFDFB8DFD4EFB2E1BAA" box="[1122,1256,655,681]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="110" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">Pachybolini</taxonomicName>
(
<bibRefCitation id="1F0C4B37FFD0FFFDFB17FD4EFF121BCE" author="Wesener, T. &amp; Enghoff, H. &amp; Wagele, J. - W." pageId="11" pageNumber="108" pagination="37 - 53" refId="ref7178" refString="Wesener, T., Enghoff, H. &amp; Wagele, J. - W. (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53. https: // doi. org / 10.1071 / IS 07008" type="journal article" year="2008">
Wesener
<emphasis id="49E9EAD4FFD0FFFDFA8DFD51FA5B1BAA" box="[1378,1437,655,681]" italics="true" pageId="11" pageNumber="108">et al.</emphasis>
2008
</bibRefCitation>
);
</paragraph>
<paragraph id="7B2236C6FFD0FFFDFF28FD19FCB91A16" blockId="11.[151,1437,547,897]" pageId="11" pageNumber="108">
- vulva subreniform and bivalve (
<figureCitation id="E3A62A43FFD0FFFDFDD1FD16FD7E1BF1" box="[574,696,727,754]" captionStart="FIGURE 7" captionStartId="9.[151,250,605,629]" captionTargetBox="[151,1436,181,582]" captionTargetId="figure-21@9.[151,1436,181,582]" captionTargetPageId="9" captionText="FIGURE 7. Costabolus baculus sp. nov., female paratype (VMNH 154). A. Ventral view of both vulvae, B. Left vulva, lateral view. Abbreviations: av, anterior valve; cx, coxite of leg 1; cxv, coxite of vulva; o, operculum; pv, posterior valve. Scale bars: 500 µm (A), 200 µm (B)." figureDoi="http://doi.org/10.5281/zenodo.6610064" httpUri="https://zenodo.org/record/6610064/files/figure.png" pageId="11" pageNumber="108">Fig. 7A, B</figureCitation>
), a synapomorphy shared with
<taxonomicName id="BC9D4D45FFD0FFFDFBF7FD16FB581BF2" box="[1048,1182,727,753]" class="Diplopoda" family="Pachybolidae" kingdom="Animalia" order="Spirobolida" pageId="11" pageNumber="110" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">Pachybolini</taxonomicName>
(
<bibRefCitation id="1F0C4B37FFD0FFFDFB43FD16FA561BF2" author="Wesener, T. &amp; Enghoff, H. &amp; Wagele, J. - W." box="[1196,1424,727,753]" pageId="11" pageNumber="108" pagination="37 - 53" refId="ref7178" refString="Wesener, T., Enghoff, H. &amp; Wagele, J. - W. (2008) Pachybolini - a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53. https: // doi. org / 10.1071 / IS 07008" type="journal article" year="2008">
Wesener
<emphasis id="49E9EAD4FFD0FFFDFAFAFD19FA8B1BF2" box="[1301,1357,727,753]" italics="true" pageId="11" pageNumber="108">et al.</emphasis>
2008
</bibRefCitation>
), but devoid of a central ridge, this being an apparent plesiomorphy.
</paragraph>
<paragraph id="7B2236C6FFD0FFFDFF28FCE1FEF81A82" blockId="11.[151,1437,547,897]" pageId="11" pageNumber="108">
- both posterior gonopods mostly connected by a tiny sternite, each distinctly biarticulate near midlength (
<figureCitation id="E3A62A43FFD0FFFDFA85FCDEFF121A5E" captionStart="FIGURE 3" captionStartId="4.[151,250,1799,1823]" captionTargetBox="[177,1411,181,1775]" captionTargetId="figure-23@4.[177,1411,181,1775]" captionTargetPageId="4" captionText="FIGURE 3. Carinobolus complex sp. nov., male paratype (RMCA). A. Midbody leg, lateral view. B. Apical parts of a postgonopodal leg with a tarsal pad, subventral view. C. First three legs, ventrocaudal view. D &amp; E. Anterior gonopods, posterior and anterior views, respectively. F. Left anterior gonopod, lateral view. G. Right posterior gonopod, lateral view. H. Posterior gonopods, caudal view. I. Drawing of right posterior gonopod, caudal view. Abbreviations: ap, antero-apical process; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; hp, basal projection of prefemur 2; pa, tarsal pad; sl, solenomere; st, sternite; t1 and t2, telopoditomeres 1 and 2; tl, telopodite. Scale bars: 200 µm (A, E, F, G), 50 µm (B), 500 µm (C, D), 0.5 mm (H, I)." figureDoi="http://doi.org/10.5281/zenodo.6610049" httpUri="https://zenodo.org/record/6610049/files/figure.png" pageId="11" pageNumber="108">Figs 3G–I</figureCitation>
,
<figureCitation id="E3A62A43FFD0FFFDFF0FFC82FED91A5D" box="[224,287,835,862]" captionStart="FIGURE 6" captionStartId="8.[151,250,1776,1800]" captionTargetBox="[238,1349,181,1753]" captionTargetId="figure-23@8.[238,1349,181,1753]" captionTargetPageId="8" captionText="FIGURE 6. Costabolus baculus sp. nov., male paratype (VNHM 198). A. Anterior part of body, ventral view. B. Coxal projections on leg-pairs 2 and 3. C. Leg-pair 2, posterior view. D. Leg-pair 3, posterior view. E-I. Everted gonopod block, anterior, ventrolateral, ventral, posterior and posterolateral views, respectively. J. Right posterior gonopod, lateral view. K. Drawing of left posterior gonopod, caudal view. Abbreviations: 1, median membranous area of telopodite; 2, lateral sclerotized area of telopodite; cx1, coxite of anterior gonopod; cx2, coxite of posterior gonopod; pg, posterior gonopod; hp, basal projection of prefemur 2; st, sternite; sl, solenomere; t1 and t2, telopoditomeres 1 and 2 of anterior gonopod; tl, telopodite of posterior gonopod. Scale bars: 500 µm (A, C), 100 µm (B), 200 µm (D, E–K)." figureDoi="http://doi.org/10.5281/zenodo.6610060" httpUri="https://zenodo.org/record/6610060/files/figure.png" pageId="11" pageNumber="108">6J–K</figureCitation>
), telopodite clearly twisted based on a subtransverse course of the seminal groove dividing the coxite and telopodite.
</paragraph>
</subSubSection>
</treatment>
</document>
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