treatments-xml/data/D8/16/CC/D816CC852981571EA5B1E555A3DEC960.xml

<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="D816CC852981571EA5B1E555A3DEC960" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) brunki Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="C5B5E518-02FF-4E24-A1C1-70F560699170" docUuidSource="ZooBank" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wrase, David W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Muilwijk, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Machard, Patrice</mods:namePart>
<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-06-13</mods:number>
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<mods:number>69</mods:number>
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<mods:number>1</mods:number>
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<mods:start>71</mods:start>
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<mods:url>http://dx.doi.org/10.3897/dez.69.83840</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-71</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:C5B5E518-02FF-4E24-A1C1-70F560699170" httpUri="http://treatment.plazi.org/id/D816CC852981571EA5B1E555A3DEC960" lastPageNumber="71" pageId="0" pageNumber="71">
<subSubSection pageId="0" pageNumber="71" type="nomenclature">
<paragraph pageId="0" pageNumber="71">
14. 
<taxonomicName LSID="http://zoobank.org/C5B5E518-02FF-4E24-A1C1-70F560699170" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) brunki</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14F</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">Yemen, 'Amran Governorate, Thula District, between Kaukaban and Shibam, approximate GPS coordinates: 15.503, 43.901.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀, 'YEMEN, ~ 40 km NW Sanaa, / Kaukaban/Shibam, small waste- / waterstream, ruralic vegetation, / sand, 2700-2930 m Handaufsam. / 
<geoCoordinate degrees="15" direction="north" minutes="30" orientation="latitude" precision="1" seconds="9.98" value="15.502772">15°30'9.98&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="43" direction="east" minutes="54" orientation="longitude" precision="1" seconds="2.66" value="43.90074">43°54'2.66&quot;E</geoCoordinate>
/ 02. Dez. 2009, legit. Ingo Brunk [p, w] // Coll. / I. Brunk / Dresden/Germ [h, pn]' (cBRU).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
<materialsCitation collectingDate="1938-02-03" collectorName="El Errein, B. M. Exp., S. W. Arabia., H. Scott, E. B. Britton., Reiche" country="YEMEN" county="Remains" determinerName="E. B. Britton" elevation="2835" location="Haz" municipality="El Errein" specimenCount="1" stateProvince="San'a">
<collectingCounty>Remains</collectingCounty>
of a specimen of uncertain sex in bad condition (only pronotum and elytra preserved), 'UNDER A STONE // 
<collectingCountry name="Yemen">YEMEN</collectingCountry>
/ 
<collectorName>
<collectingMunicipality>El Errein</collectingMunicipality>
</collectorName>
, nr. 
<location LSID="urn:lsid:plazi:treatment:D816CC852981571EA5B1E555A3DEC960:3E358768E5D3A423AA89691B0268ECCE" country="YEMEN" county="Remains" municipality="El Errein" name="Haz" stateProvince="San'a">Haz</location>
, / about 
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.5749504" unit="mi" value="16.0">16 miles</quantity>
N.W. / of 
<normalizedToken originalValue="San’a">
<collectingRegion country="Yemen" name="San'a'">San'a</collectingRegion>
</normalizedToken>
, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">9,300ft.</elevation>
</quantity>
[= 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">2835 m</elevation>
</quantity>
] / 
<collectingDate value="1938-02-03">3.ii.1938</collectingDate>
.// 
<collectorName>B.M. Exp.</collectorName>
to / 
<collectorName>S.W. Arabia.</collectorName>
/ 
<collectorName>H. Scott</collectorName>
&amp; / 
<collectorName>E.B. Britton.</collectorName>
/ B.M. 1938-246, // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
/ 
<collectorName>Reiche</collectorName>
/ 
<determinerName>E.B. Britton</determinerName>
det. / 1946' (NHMUK)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="71">
TME: 
<specimenCount type="generic">2 specimens</specimenCount>
. TGE: 
<specimenCount type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">Latinized patronym based on the surname of Ingo Brunk (Dresden, Germany), who collected the first well-preserved representative of the new species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
<taxonomicName authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus brunki</emphasis>
</taxonomicName>
sp. nov. readily differs from all other 
<taxonomicName authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-taxa in its smallest value in ratio PW/PL (Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
). It is most closely related to 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
; both species are the only two within &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group with darkest coloration of integument and sides of pronotum not sinuate from widest point to base (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
has also sides of pronotum not sinuate to base, but much lighter coloration of integument and higher value in ratio EL/PL). The new species is distinct from 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., in: (1) pronotum with anterior angles more rounded at tips (vs. pronotum with anterior angles more pointed at tips, Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3B, C</figureCitation>
); (2) pronotum nearly as long as wide, PW/PL: 1.02-1.05 (vs. pronotum appreciably wider than long, PW/PL: 1.07-1.11); (3) pronotum subcordate, less constricted anteriorly, PW/PA: 1.38-1.42, with greater value in ratio PA/PB: 0.80-0.88 (vs. pronotum subquadrate, more constricted anteriorly, PW/PA: 1.44-1.48, with smaller value in ratio PA/PB: 0.76-0.79); (4) elytra compared to pronotum slightly shorter, EL/PL: 2.40-2.44 (vs. elytra compared to pronotum slightly longer, EL/PL: 2.47-2.56). Females are different from all other representatives of the &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the presence of two, instead of one, dorsolateral ensiform setae on the apical gonocoxite (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL of holotype: 9.50 mm; BW: 3.30 mm), with elongate, moderately convex body (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3C</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly reddish-brown, with palpi and antennae slightly lighter. Integument slightly to moderately shiny, head and pronotum shinier than elytra. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Microreticulation of pronotum and elytra as in 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. Ventral surface largely with well-impressed isodiametric or slight transverse sculpticells, epipleura, mesosternum and middle coxa with sculpticells less apparent. Head (holotype) nearly smooth, impunctate, wrinkles present only on clypeus and frons, including frontal foveae and area surrounding them. Pronotum surface largely smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled, as well as each side along lateral margin with a few shallow punctures not reaching anterior half. Elytral intervals without apparent punctation. Abdominal ventrites nearly smooth, impunctate, 3-5 with a few fine wrinkles at sides. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Head</emphasis>
(holotype). About two-thirds as wide as pronotum (PW/HW of holotype: 1.36). Eyes long, slightly convex. Labrum subrectangular, as long as clypeus, with anterior margin concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows shallow, subfoveolate. Paraorbital sulci straight, backward barely reaching posterior margin of eye, not reaching level of posterior supraorbital pore. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum almost as wide as long (PW/PL= 1.02-1.05), with widest point at second quarter (holotype) or medial fifth (specimen from &quot;El Errein, nr. Haz&quot;). Anterior and posterior transverse impressions slightly distinct medially, indistinct laterally. Sides not sinuate, smoothly convex anteriorly and medially, straight (specimen from &quot;El Errein, nr. Haz&quot;) or convex posteriorly (holotype); anterior bead present laterally, lacking in medial 1/8 to 1/10; basal bead present laterally, reduced to absent in medial half; posterior angles rounded (holotype) or obtuse (specimen from &quot;El Errein, nr. Haz&quot;). Metepisternum as long as wide, MA/MI= 1.00. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times as long as elytra (EL/EW= 1.58-1.59), two and a half times as long as pronotum (EL/PL= 2.40-2.44), and a one and half as wide as pronotum (EW/PW= 1.48), with widest point at medial third. Parascutellar striole scarcely distinct, very shallow; striae more impressed than striole, barely punctate; parascutellar striole not joining stria 1; striae 1-5 and sometimes 6 reaching basal bead, 7 ending little before. Interval 3 with three discal setiferous punctures, medial one larger than others, in midst of interval 3 (holotype) or near stria 2 (specimen from &quot;El Errein, nr. Haz&quot;), anterior and posterior punctures smaller, adjoining stria 3 (posterior puncture lacking on right elytron of holotype). Umbilicate setiferous series with 16 or 17 punctures of each elytron. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Left mesofemur with 5, right one with 4 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
The holotype was collected by sifting the waste-water vegetation and soil at a humid place where waste water falls down (Ingo Brunk, pers. comm.). The correct altitude of the type locality is ca. 2700 m a. s. l. The locality is situated approximately half-way on a very steep walking path starting from Kawkaban (town on the top of the Mount Kawkaban) downwards to Schibam Kawkaban (town on the bottom of the mountain). Together with the holotype, numerous specimens of 
<taxonomicName authorityName="Belousov" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus (Arabotrechus) rougemontiellus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="rougemontiellus" subGenus="Arabotrechus">Trechus (Arabotrechus) rougemontiellus</taxonomicName>
Belousov, 2017 and few 
<taxonomicName class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Bembidion</emphasis>
</taxonomicName>
spp. were collected.
</paragraph>
<caption doi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Figure 13.</emphasis>
Spermathecal complex and gonocoxites, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken originalValue="Mařan">Maran</normalizedToken>
, 1940, topotype female specimen; 
<emphasis bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., topotype female paratype; 
<emphasis bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., topotype female paratype; 
<emphasis bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., topotype female paratype; 
<emphasis bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., female paratype, col de Buglan 1500 m, border area of 
<normalizedToken originalValue="Bingöl">Bingoel</normalizedToken>
and 
<normalizedToken originalValue="Muş">Mus</normalizedToken>
provinces, Turkey; 
<emphasis bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: 
<emphasis bold="true" pageId="0" pageNumber="71">ag</emphasis>
- apical gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="71">at</emphasis>
- atrium; 
<emphasis bold="true" pageId="0" pageNumber="71">bc</emphasis>
- bursa copulatrix; 
<emphasis bold="true" pageId="0" pageNumber="71">bg</emphasis>
- basal gonocoxite; 
<emphasis bold="true" pageId="0" pageNumber="71">co</emphasis>
- common oviduct; 
<emphasis bold="true" pageId="0" pageNumber="71">rc</emphasis>
- receptaculum; 
<emphasis bold="true" pageId="0" pageNumber="71">sg</emphasis>
- spermathecal gland; 
<emphasis bold="true" pageId="0" pageNumber="71">smc</emphasis>
- seminal canal; 
<emphasis bold="true" pageId="0" pageNumber="71">spc</emphasis>
- spermathecal canal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
Yemen (
<normalizedToken originalValue="Sana’a">Sana'a</normalizedToken>
Governorate), see Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>