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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="BBF4183AD25A530FAAA63812E11B8198" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Effossana tenuemarginata Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="BE2F32D3-AEAD-4D31-B730-36E4D44E9683" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>81</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>79</mods:start>
<mods:end>164</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e81961</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-81-79</mods:identifier>
<mods:identifier type="ZooBank">5C1FDC828F9F4869ADDD83FA96E507ED</mods:identifier>
<mods:identifier type="Pensoft-UUID">CA16EA0C5A33560FBC955981DFC9D502</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:BE2F32D3-AEAD-4D31-B730-36E4D44E9683" httpUri="http://treatment.plazi.org/id/BBF4183AD25A530FAAA63812E11B8198" lastPageNumber="79" pageId="0" pageNumber="79">
<subSubSection pageId="0" pageNumber="79" type="nomenclature">
<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="https://zoobank.org/BE2F32D3-AEAD-4D31-B730-36E4D44E9683" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Effossana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Effossana tenuemarginata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="tenuemarginata" status="sp. nov.">Effossana tenuemarginata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">Figs 41</figureCitation>
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Species of Effossana gen. nov., dorsal (left) and lateral (right) views. A, B E. gloma comb. nov., male, DZUP. C, D E. intrinseca sp. nov., holotype male, DZUP. E, F E. scapa comb. nov., holotype male, BMNH. G, H E. tenuemarginata sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure43" httpUri="https://binary.pensoft.net/fig/798196" pageId="0" pageNumber="79">, 43G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in lateral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41C</figureCitation>
), crown-face transition thin. Pygofer (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41G</figureCitation>
) without processes. Style, in lateral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41K</figureCitation>
), with blade short, dorsal margin forming conspicuous process directed dorsally, flattened laterally and abruptly tapered apically. Aedeagus (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41L, M</figureCitation>
) apodemal processes narrow and straight, almost as long as shaft, gradually tapered toward apex; shaft with pair of processes directed dorsally, anterior margin bearing a small spiniform process near mid-length.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 9.7 mm; paratypes, males (n = 5) 9.7-10.3 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A-B</figureCitation>
,
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Species of Effossana gen. nov., dorsal (left) and lateral (right) views. A, B E. gloma comb. nov., male, DZUP. C, D E. intrinseca sp. nov., holotype male, DZUP. E, F E. scapa comb. nov., holotype male, BMNH. G, H E. tenuemarginata sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure43" httpUri="https://binary.pensoft.net/fig/798196" pageId="0" pageNumber="79">43G-H</figureCitation>
) reddish-brown. Crown (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A</figureCitation>
) with pair of rounded black spots near posterior margin, behind ocelli. Face (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41B</figureCitation>
) without maculae. Pronotum (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A</figureCitation>
) with black punctures; larger black spots near anterior margin, behind eye; humeral angels with a small yellow macula; posterior margin with poorly defined dark brown transverse band; proepimeron (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41C</figureCitation>
), with transverse black stripe below pronotal carina. Mesonotum (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A</figureCitation>
) with three pairs of yellow spots: two pairs near anterior margin and one near scutoscutelar suture. Scutellum (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A</figureCitation>
) with pair of large yellow spots. Forewing (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41D</figureCitation>
) strongly marked by dark-brown mottling and yellow maculae; membrane translucent brown; veins dark brown. Metatibia (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in dorsal view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41A</figureCitation>
), transocular width 8.1 tenths of humeral width of pronotum. Head, in frontal view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41B</figureCitation>
), frons 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; clypeus 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; lateral margins parallel. Head, in lateral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41C</figureCitation>
), crown-face transition thin, with 5 transverse striae. Forewing (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41D</figureCitation>
) with appendix developed, as wide as first apical cell maximum width. Protibia AD row without differentiated setae. Metatibia PD, AD and AV rows with 23-24, 12 and 17-19 macrosetae respectively. Metatarsomere I elongated, about 4
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 8-10 setae; pecten with 4 platellae. Metatarsomere II pecten with 2 platellae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41E</figureCitation>
) 1.6
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, lateral margins parallel; posterior margin with deep rounded excavation. Valve (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41F</figureCitation>
) 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin narrowly excavated, in lateral view, weakly convex. Pygofer, in lateral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41G</figureCitation>
), 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; anteroventral margin rounded; posterodorsal and posteroventral margins straight and slightly convergent posterad; apex broadly rounded. Subgenital plate, in ventral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41H</figureCitation>
), elongated, 4.7
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; inner margin straight; outer margin slightly rounded; apex rounded. Connective (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41I</figureCitation>
) with arms moderately robust. Style, in lateral view (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41K</figureCitation>
), with blade short, widened towards apex; ventral margin straight and serrated on apical 2\3; dorsal margin forming conspicuous process directed dorsally, flattened laterally and abruptly tapered apically, forming acute small tip curved anterad; apex rounded. Aedeagus (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41L, M</figureCitation>
) dorsal apodeme developed, dorsal margin slightly excavated, lateral margins weakly projected dorsolaterally; apodemal process elongated and narrow, almost as long as shaft, slightly curved dorsally and gradually tapered toward apex; shaft very long, slightly flattened dorsoventrally, basal portion curved, posterior portion almost straight; apex truncated, with pair of processes directed dorsally, anterior margin bearing a small spiniform process near mid-length. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the crown-face transition thin.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2012-01-04" collectingDateMax="2012-01-17" collectingDateMin="2012-01-04" collectorName="W. Datilo, G. R. Mazao" country="Brazil" county="Brasil" latitude="-9.591389" location="Porto" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" specimenCount="1" stateProvince="Rondonia" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male:
<collectingCountry name="Brazil">Brazil</collectingCountry>
:
<normalizedToken originalValue="Rondônia">
<collectingRegion country="Brazil" name="Rondonia">Rondonia</collectingRegion>
</normalizedToken>
</emphasis>
: &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:C08C8C9E8816C260DB8E61D1B31D77DF" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Porto" stateProvince="Rondonia">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:CA3BF954B3BFC3D1087C0200F2488CA4" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Velho" stateProvince="Rondonia">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:E5D97E7CAD0BD4E743C7E2797972AF1D" country="Brazil" county="Brasil" latitude="-9.591389" longLatPrecision="21" longitude="-65.049164" municipality="Rondonia" name="Rio Madeira" stateProvince="Rondonia">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
\ Malaise
<collectingDate value="2012-01-04" valueMax="2012-01-17" valueMin="2012-01-04">04-17.I.2012</collectingDate>
\
<collectorName>W. Datilo</collectorName>
&amp;
<collectorName>
G.R.
<normalizedToken originalValue="Mazão”">Mazao&quot;</normalizedToken>
</collectorName>
(MZSP)
</materialsCitation>
.
<materialsCitation collectingDate="2011-06-18" collectingDateMax="2011-06-29" collectingDateMin="2011-06-18" country="Brazil" county="Brasil" latitude="-9.435555" location="Porto" longLatPrecision="21" longitude="-64.802505" municipality="Rondonia" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Paratypes">
<typeStatus>Paratypes</typeStatus>
</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
,
<normalizedToken originalValue="Rondônia">
<collectingMunicipality>Rondonia</collectingMunicipality>
</normalizedToken>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:AB21FB84201DC7FF4A3B90DBBE90BED2" country="Brazil" county="Brasil" latitude="-9.435555" longLatPrecision="21" longitude="-64.802505" municipality="Rondonia" name="Porto" stateProvince="Paratypes">Porto</location>
\
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:338ACE484CFF6ACC67C9A3C644B40A3B" country="Brazil" county="Brasil" latitude="-9.435555" longLatPrecision="21" longitude="-64.802505" municipality="Rondonia" name="Velho" stateProvince="Paratypes">Velho</location>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:3A350B5A0CF956DA26E86259625E459E" country="Brazil" county="Brasil" latitude="-9.435555" longLatPrecision="21" longitude="-64.802505" municipality="Rondonia" name="Rio Madeira" stateProvince="Paratypes">Rio Madeira</location>
\
<geoCoordinate degrees="09" direction="south" minutes="26" orientation="latitude" precision="15" seconds="08" value="-9.435555">09°26′08″S</geoCoordinate>
<geoCoordinate degrees="64" direction="west" minutes="48" orientation="longitude" precision="15" seconds="09" value="-64.802505">64°48′09″W</geoCoordinate>
\ Malaise
<collectingDate value="2011-06-18" valueMax="2011-06-29" valueMin="2011-06-18">18-29.VI.2011</collectingDate>
\ Albuquerque &amp; Ferreira&quot; (MZSP)
</materialsCitation>
;
<specimenCount type="male">1♂</specimenCount>
, same as preceding except
<geoCoordinate degrees="09" direction="south" minutes="26" orientation="latitude" precision="15" seconds="14" value="-9.4372225">09°26′14″S</geoCoordinate>
<geoCoordinate degrees="64" direction="west" minutes="49" orientation="longitude" precision="15" seconds="58" value="-64.83278">64°49′58″W</geoCoordinate>
(MZSP);
<specimenCount type="male">2♂</specimenCount>
, same as preceding except
<geoCoordinate degrees="09" direction="south" minutes="35" orientation="latitude" precision="15" seconds="29" value="-9.591389">09°35′29″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="02" orientation="longitude" precision="15" seconds="57" value="-65.049164">65°02′57″W</geoCoordinate>
(DZUP). -
<materialsCitation collectingDate="2011-05-11" collectingDateMax="2011-05-15" collectingDateMin="2011-05-11" country="Brazil" county="Brasil" elevation="175" latitude="-7.363055" location="Rio Liberdade" longLatPrecision="21" longitude="-71.868614" municipality="Ipixuna" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Amazonas">Amazonas</collectingRegion>
</emphasis>
:
<specimenCount type="male">1♂</specimenCount>
, &quot;
<collectingCounty>Brasil</collectingCounty>
: AM,
<collectingMunicipality>Ipixuna</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:51D4B0578CA03A51F305CD52DCAC9B04" country="Brazil" county="Brasil" latitude="-7.363055" longLatPrecision="21" longitude="-71.868614" municipality="Ipixuna" name="Rio Liberdade" stateProvince="Amazonas">Rio Liberdade</location>
,
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:766FE6E90C2210342F11E52A884BAD58" country="Brazil" county="Brasil" latitude="-7.363055" longLatPrecision="21" longitude="-71.868614" municipality="Ipixuna" name="Com." stateProvince="Amazonas">Com.</location>
\
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:47E181EFA8C8FF049385B665ECE90D79" country="Brazil" county="Brasil" latitude="-7.363055" longLatPrecision="21" longitude="-71.868614" municipality="Ipixuna" name="Sao Vicente" stateProvince="Amazonas">
<normalizedToken originalValue="São">Sao</normalizedToken>
Vicente
</location>
no
<location LSID="urn:lsid:plazi:treatment:BBF4183AD25A530FAAA63812E11B8198:8A98644A195296571A68F7A55FE42EBE" country="Brazil" county="Brasil" latitude="-7.363055" longLatPrecision="21" longitude="-71.868614" municipality="Ipixuna" name="Estirao da Preta" stateProvince="Amazonas">
<normalizedToken originalValue="Estirão">Estirao</normalizedToken>
da Preta
</location>
\
<geoCoordinate degrees="07" direction="south" minutes="21" orientation="latitude" precision="15" seconds="47" value="-7.363055">07°21′47″S</geoCoordinate>
<geoCoordinate degrees="071" direction="west" minutes="52" orientation="longitude" precision="15" seconds="07" value="-71.868614">071°52′07″W</geoCoordinate>
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.75" unit="m" value="175.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.75" unit="m" value="175.0">175m</elevation>
</quantity>
\
<collectingDate value="2011-05-11" valueMax="2011-05-15" valueMin="2011-05-11">11-15.v.2011</collectingDate>
Cavichioli,\
<normalizedToken originalValue="Gonçalves">Goncalves</normalizedToken>
, Rafael\ &amp; Takiya et al.&quot; (DZRJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Effossana" kingdom="Animalia" lsidName="Effossana tenuemarginata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="tenuemarginata">
<emphasis italics="true" pageId="0" pageNumber="79">Effossana tenuemarginata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
resembles
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. gatuna" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="gatuna">
<emphasis italics="true" pageId="0" pageNumber="79">E. gatuna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
and
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. scapa" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="scapa">
<emphasis italics="true" pageId="0" pageNumber="79">E. scapa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
by the shape of aedeagus with the apodemal processes straight. However,
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. tenuemarginata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="tenuemarginata">
<emphasis italics="true" pageId="0" pageNumber="79">E. tenuemarginata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be separated from
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. gatuna" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="gatuna">
<emphasis italics="true" pageId="0" pageNumber="79">E. gatuna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
and
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. scapa" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="scapa">
<emphasis italics="true" pageId="0" pageNumber="79">E. scapa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
by the style (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Effossana tenuemarginata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure41" httpUri="https://binary.pensoft.net/fig/798194" pageId="0" pageNumber="79">41K</figureCitation>
), with blade short and dorsal margin forming a conspicuous process directed dorsally, flattened laterally and abruptly tapered apically.
</paragraph>
</subSubSection>
</treatment>
</document>
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